Paleontological sites of Lebanon

A plate with Pseudostacus sp. (lobster, left), Diplomystus birdii (fish, right), and a partial Dercetis triqueter, Cretaceous Hakel, Lebanon

The paleontological sites of Lebanon contain deposits of some of the best-preserved fossils in the world, and include some species found nowhere else. The most famous of these is the Lebanese lagerstätten of the Late Cretaceous age, which contain a well-preserved variety of different fossils. Some fossils date back to the Jurassic period, and younger fossils of mammals from a different site belong to the Miocene through the Pleistocene. The fossils found in Lebanon are displayed in natural history, geology and paleontology museums around the world.

## Contents

* 1 History
* 2 Locations and fossils
* 3 Fossils found in Lebanon
* 4 See also
* 5 References
* 6 External links

## History

The earliest known account of Lebanese fossils is attributed to Herodotus.[1][2] The existence of fossil fishes in the Lebanon is referred to by Jean de Joinville. In his Histoire de Saint Louis he wrote that during the sojourn of King Louis IX of France at Sidon in 1253, just before his return home from the Seventh Crusade, a stone was brought him,

“  | which was the most marvellous in the world, for when a layer of it was lifted, there was found between the two pieces the form of a fish. The fish was of stone, but lacked nothing in form, eyes, bones, colour, or anything necessary to a living fish.[3] | ”

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European scientists became interested in Lebanese fossils in the 19th century.[1]

## Locations and fossils

Rhinobatos whitfieldi, a species from the Cretaceous period found in Lebanon

There are three major fossil locations in Lebanon: Sahel Alma, Hajula and Hakel. Hajula and Hakel are each about twelve miles north-east of the coastal town of Jebeil, the ancient Byblos. Hajula is situated six miles south of Hakel; and between the two villages there are two westward-projecting spurs of Mount Lebanon and an intervening valley. Both villages are approximately 2500–3000 feet above sea level. In both places there is clear evidence of faulting by which the fish-bearing strata have been lowered into the midst of older strata. Those at Hakel have been lowered to the level of the hippurite limestone of Lebanon, being above the trigonia sandstone. The Hajula beds are thought to be an extension of those at Hakel. The study of fish fossils appears to show that the horizon of the beds at Hajula is somewhat higher than that of the beds at Hakel.[4]

The fishes found at Sahel Alma mostly belong to the same genera as those at Hakel and Hajula, but of around sixty species, probably not one is found elsewhere. Twenty-one species found at Hajula also occur at Hakel. These data make it quite certain that the beds at Sahel Alma are on a different level from those at Hakel and Hajula; while those at the latter places are on the same, or nearly the same, horizon. Opinions differ as to which are older, the fish-beds at Sahel Alma or those at Hakel, but the view of modern authorities is that those at Hakel are more ancient. This opinion appears to be supported by the character of the fishes in each. From a study of the fishes taken at Hajula, the beds containing them may belong to a slightly more recent time than that of the beds at Hakel.[5]

The landscape of Lebanon has been subject to volcanic eruptions, tectonic plate movement, and the rising and dropping of sea levels. The Late Cretaceous was characterized by very high sea levels. During that time fossil-bearing limestone was formed.[6]

One of the recent discoveries was made in 1989. M. Malez and A. Forsten described Hipparion from the Bekaa Valley.[7]

Another find was a snake with two legs. This provided a valuable example of evolution at work, illustrating how ancient lizards became modern snakes. [8]

Octopuses are some of the rarest marine fossils, since most species have no hard structures apart from a beak. An octopus's corpse cannot usually survive long enough to become fossilized, which is why the discovery of well-preserved octopuses in Lebanon in 2009 was remarkable. Dirk Fuchs of the Freie University Berlin said about these 95-million-year-old fossil octopuses found in Lebanon:

“  | This provides important evolutionary information. The more primitive relatives of octopuses had fleshy fins along their bodies. The new fossils are so well preserved that they show, like living octopus, that they didn't have these structures.[9] | ”

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The scientific name of the new find is Keuppia levante. Some of them have been preserved so well that their ink is still visible.[9]

## Fossils found in Lebanon

* Enantiophoenix
* Palaeoctopus
* Keuppia
* Styletoctopus
* Eupodophis
* Palibacus
* Ichthyotringa

## See also

* Byblos Fossil Museum
* Dany Azar on Wikispecies
* Mim Museum

## References

1. ↑ 1.0 1.1 Fabio M. Dalla Vecchia1, Paolo Arduini and Alexander W. A. Kellner (2001). "The first pterosaur from the Cenomanian (Late Cretaceous) Lagerstatten of Lebanon". http://doc.rero.ch/lm.php?url=1000,43,39,20091210003025-ZZ/PAL_E2341.pdf.
2. ↑ Nature, Volume 36 By Nature Publishing Group, UM-MEDSEARCH Gateway. June 9, 1887. p. 230. https://books.google.com/books?id=on8CAAAAIAAJ&pg=PA132&lpg=PA132&dq=The+earliest+account+of+the+fossils+in+Lebanon+is+attributed+to+Herodotus&source=bl&ots=vDGDoe1LIO&sig=eNpc0Grnq6HI9qfGxlBjmJLmJvo&hl=en&ei=gcFeTM3-B46WsgPJ-6CqCw&sa=X&oi=book_result&ct=result&resnum=4&ved=0CCIQ6AEwAw#v=onepage&q&f=false.
3. ↑ Geological Society of London (1896). The Quarterly journal of the Geological Society of London, Volume 52. Harvard University. p. 230. https://books.google.com/books?ei=5OhdTM7lGZHCsAOnvfipCw&ct=result&output=text&id=-b4KAAAAYAAJ&dq=lebanon+fossils+hakel&q=hakel#v=snippet&q=sidon&f=false.
4. ↑ Joel Asaph Allen, American Museum of Natural History (1903). Bulletin of the American Museum of Natural History, Volume 19. Order of the Trustees. pp. 395–396. https://books.google.com/books?id=1S9KAAAAYAAJ&pg=PA396-IA3&dq=lebanon+fossils+hakel+elevation&hl=en&ei=o_NdTJ_NAov0tgP_voXJDQ&sa=X&oi=book_result&ct=result&resnum=3&ved=0CDsQ6AEwAg#v=onepage&q=hakel&f=false.
5. ↑ American Society of Naturalists (1903). The American naturalist, Volume 37. p. 687. https://books.google.com/books?id=9mTyAAAAMAAJ&pg=PA687&dq=hakel+lebanon+fossils&hl=en&ei=RAteTJPKGY68sQPog_mpCw&sa=X&oi=book_result&ct=result&resnum=6&ved=0CE4Q6AEwBQ#v=onepage&q=hakel lebanon fossils&f=false.
6. ↑ Chris Walley. "Geology of Lebanon". American University of Beirut. http://ddc.aub.edu.lb/projects/geology/geology-of-lebanon/.
7. ↑ "M Malez and A Forsten 1989". The Paleobiology Database. 1989. http://paleodb.geology.wisc.edu/cgi-bin/bridge.pl?action=displayReference&reference_no=9049.
8. ↑ Jonathan Amos (10 April 2008). "Ancient serpent shows its leg". BBC. http://news.bbc.co.uk/2/hi/science/nature/7339508.stm.
9. ↑ 9.0 9.1 "Cretaceous Octopus With Ink And Suckers -- The World's Least Likely Fossils?". www.sciencedaily.com. https://www.sciencedaily.com/releases/2009/03/090317111902.htm.

## External links

* "Fossils from the amber of Lebanon". American University in Beirut. 1998. http://ddc.aub.edu.lb/projects/amber/index.html. Retrieved 8 August 2010.
* Walley, C.D. (1998). "1.3 Fossils of Lebanon". The Geology of Lebanon - a summary. American University of Beirut. http://ddc.aub.edu.lb/projects/geology/geology-of-lebanon/. Retrieved 8 August 2010.
* Expo Hakel, a fossil museum in Hakel

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