Argiles et Grès à Reptiles Formation
Stratigraphic range: Campanian-Early Maastrichtian, 74–70 Ma

PreЄ

Type| Geological formation
Underlies| Calcaire de Rognac
Thickness| Over 40 m (130 ft)
Lithology
Primary| Clay, shale
Other| Sandstone
Location
Coordinates|  [ ⚑ ] : 43°30′N 5°30′E﻿ / ﻿43.5°N 5.5°E﻿ / 43.5; 5.5
Paleocoordinates|  [ ⚑ ] 36°06′N 4°00′E﻿ / ﻿36.1°N 4.0°E﻿ / 36.1; 4.0
Region| Provence-Alpes-Cote d'Azur, Bouches-du-Rhône
Country|  France
Extent| Aix-en-Provence Basin

Argiles et Grès à Reptiles Formation (France)

Argiles et Grès à Reptiles Formation (meaning Reptile Clay and Sandstone) also known as the Argiles Rutilantes Formation[1] is an early Maastrichtian[2] France geologic formation in the département of Var preserving the remains of several types of dinosaurs and other extinct organisms.[3]

## Contents

* 1 Fossil content
* 1.1 Dinosaurs
* 1.2 Crocodylians
* 1.3 Turtles
* 1.4 Pterosaurs
* 1.5 Fish
* 1.6 Fossil eggs
* 1.7 Flora
* 2 See also
* 3 References
* 3.1 Bibliography
* 4 Further reading

## Fossil content

An abelisaurid similar to Arcovenator is known from Département des Bouches-du-Rhone, while enantiornithean and indeterminate avialan remains are known from Département du Var.[2]

### Dinosaurs

Dinosaurs of the Grès à Reptiles

Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images
Abelisauridae | Indeterminate  | Bouches du Rhône  | Lower  |  | A large abelisaurid that was similar to Arcovenator.  |
Arcovenator[4] | A. escotae | Var  | Lower  |  | A large abelisaurid.  |
Atsinganosaurus[5] | G. meridionalis | Bouches du Rhône  | Lower  |  | A titanosaur Lirainosaurinae.  |
Castignovolucris[6] | C. sebei | Castigno valley, Villespassans, Hérault[6] | Lower (i.e. continental red clays)[6] |  | An enantiornithine bird.  |
Garrigatitan[7] | A. velauciensis | Bouches du Rhône  | Lower  |  | A titanosaur Lirainosaurinae.  |
Gargantuavis[8] | G. philoinos[8] | Var[8] |  |  |  |
Hypselosaurus[9] | H. priscus[9] |

* Bouches-du-Rhône[10]
* Var[8]

|  |  | Actually indeterminate sauropod remains.[9] |

Matheronodon[9] | M. provincialis[9] |

* Bouches-du-Rhône [10]

|  |  | A rhabdodontid ornithopod |

Theropoda | Indeterminate  | Var[8] |  |  | Previously referred to "Megalosaurus" pannoniensis |
Pyroraptor[10] | P. olympius[10] | Bouches-du-Rhône[10] |  | "Fragmentary postcrania and teeth"[11] |  |

Rhabdodon[9] | R. priscus[9] |

* Bouches-du-Rhône[10]
* Var[8]

|  |  |  |

Rhodanosaurus[10] | R. lugdunensis[10] | Bouches-du-Rhône[10] |  |  |  |
Variraptor[9] | V. mechinorum[9] |

* Var[8]
* Bouches-du-Rhône[10]

|  |  |  |

Mammals of the Grès à Reptiles

Genus | Species | Location | Stratigraphic position | Abundance | Notes | Images
Mistralestes | M. arcensis | La Cairanne  |  |  |  |
Valentinella | V. vitrollense | Vitrolles-La Plaine  |  |  |  |

### Crocodylians

* Acynodon iberoccitanus[12]
* Allodaposuchus sp.[13]
* Alligatoridae indet.[14]
* Crocodylidae indet.[14]

### Turtles

* Foxemys mechinorum[15]
* cf. Polysternon sp.[16]
* cf. Solemys sp.[16]

### Pterosaurs

* Mistralazhdarcho maggii[17]

### Fish

* Hybodontiformes indet.[16]
* Lepisosteidae indet.[16]

### Fossil eggs

* Megaloolithus aureliensis[18]
* Megaloolithus petralta[18]
* Megaloolithus sp.[18]

### Flora

* Peckichara cancellata[16]
* P. pectinata[16]
* Platychara caudata[16]
* Saportanella maslovi[16]
* ?Amblyochara begudiana[16]

## See also

* List of dinosaur-bearing stratigraphic units
* List of fossiliferous stratigraphic units in France

## References

1. ↑ Tabuce et al., 2013
2. ↑ 2.0 2.1 Weishampel et al., 2004, pp.588-593
3. ↑ Argiles et Grès à Reptiles Formation at Fossilworks.org
4. ↑ Tortosa, Thierry; Buffetaut, Eric; Vialle, Nicolas; Dutour, Yves; Turini, Eric; Cheylan, Gilles (2014-01-01). "A new abelisaurid dinosaur from the Late Cretaceous of southern France: Palaeobiogeographical implications" (in en). Annales de Paléontologie 100 (1): 63–86. doi:10.1016/j.annpal.2013.10.003. ISSN 0753-3969. Bibcode: 2014AnPal.100...63T. https://www.researchgate.net/publication/259295271.
5. ↑ Díez Díaz, Verónica; Garcia, Géraldine; Pereda Suberbiola, Xabier; Jentgen-Ceschino, Benjamin; Stein, Koen; Godefroit, Pascal; Valentin, Xavier (2020-12-21). "A new titanosaur (Dinosauria: Sauropoda) from the Upper Cretaceous of Velaux-La-Bastide Neuve (southern France)" (in en). Historical Biology 33 (11): 2998–3017. doi:10.1080/08912963.2020.1841184. ISSN 0891-2963. https://www.tandfonline.com/doi/full/10.1080/08912963.2020.1841184.
6. ↑ 6.0 6.1 6.2 Buffetaut, E.; Angst, D.; Tong, H. (2023). "A new enantiornithine bird from Upper Cretaceous non-marine deposits at Villespassans (Hérault, southern France)". Annales de Paléontologie 109 (1): 102585. doi:10.1016/j.annpal.2022.102585. Bibcode: 2023AnPal.10902585B.
7. ↑ B. VilaExpression error: Unrecognized word "et". (2012). "The diversity of sauropod dinosaurs and their first taxonomic succession from the latest Cretaceous of southwestern Europe: Clues to demise and extinction". Palaeogeography, Palaeoclimatology, Palaeoecology 350-352 (15): 19–38. doi:10.1016/j.palaeo.2012.06.008. Bibcode: 2012PPP...350...19V.
8. ↑ 8.0 8.1 8.2 8.3 8.4 8.5 8.6 "14.8 Département du Var, France; 2. Grès à Reptiles Formation," in Weishampel et al., 2004, p.589
9. ↑ 9.0 9.1 9.2 9.3 9.4 9.5 9.6 9.7 9.8 "14.8 Département du Var, France; 2. Grès à Reptiles Formation" and "14.38 Département des Bouches-du-Rhone, France; 1. Grès à Reptiles Formation," in Weishampel et al., 2004, pp.589-590
10. ↑ 10.0 10.1 10.2 10.3 10.4 10.5 10.6 10.7 10.8 10.9 "14.38 Département des Bouches-du-Rhone, France; 1. Grès à Reptiles Formation," in Weishampel et al., 2004, p.590
11. ↑ "Table 10.1," in Weishampel et al., 2004, p.199
12. ↑ Martin, 2007
13. ↑ Blanco & Brochu, 2016
14. ↑ 14.0 14.1 Le Loeuff & Buffetaut, 1998
15. ↑ Tong et al., 1998
16. ↑ 16.0 16.1 16.2 16.3 16.4 16.5 16.6 16.7 16.8 Martin et al., 2016
17. ↑ Vullo et al., 2018
18. ↑ 18.0 18.1 18.2 Viany-Liaud et al., 1994

### Bibliography

* Vullo, R.; G. Garcia; P. Godefroit; A. Cincotta, and X. Valentin. 2018. Mistralazhdarcho maggii, gen. et sp. nov., A New Azhdarchid Pterosaur from the Upper Cretaceous of Southeastern France. Journal of Vertebrate Paleontology 38\. 1-16.
* Blanco, A., and C. A. Brochu. 2016. Intra- and interspecific variability in allodaposuchid crocodylomorphs and the status of western European taxa. Historical Biology 29\. 395-408.
* Martin, J. E.; M. Delfino; G. Garcia; P. Godefroit; S. Berton, and X. Valentin. 2016. New specimens of Allodaposuchus precedens from France: intraspecific variability and the diversity of European Late Cretaceous eusuchians. Zoological Journal of the Linnean Society 176\. 607-631.
* Cheylan, Rodolphe; Thierry Tortosa; Monique Vianey-Liaud; Géraldine Garcia; Renaud Lebrun; Pascal Godefroit; Yves Dutour; Sévérine Berton, and Xavier Valentin. 2013. [ New eutherian mammals from the Late Cretaceous of Aix-en-Provence Basin, south-eastern France]. Zoological Journal of the Linnean Society 169\. 653–672. doi:10.1111/zoj.12074 ISSN 0024-4082
* Martin, J. E. 2007. New material of the Late Cretaceous globidontan Acynodon iberoccitanus (Crocodylia) from southern France. Journal of Vertebrate Paleontology 27\. 362-372.
* Weishampel, David B.; Peter Dodson, and Halszka Osmólska (eds.). 2004. The Dinosauria, 2nd edition, 1–880. Berkeley: University of California Press. Accessed 2019-02-21. ISBN:0-520-24209-2
* Le Loeuff, J., and É. Buffetaut. 1998. A new dromaeosaurid theropod from the Upper Cretaceous of southern France.. Oryctos 1\. 105-112.
* Tong, H.; E. S. Gaffney, and É. Buffetaut. 1998. Foxemys, a new side-necked turtle (Bothremydidae: Pelomedusoides) from the Late Cretaceous of France. American Museum Novitates 3251\. 1-19.
* Vianey-Liaud, M.; P. Mallan; O. Buscail, and C. Montgelard. 1994. Review of French dinosaur eggshells: morphology, structure, mineral, and organic composition. In K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies, 151–183. Cambridge University Press, Cambridge.

## Further reading

* R. Cousin, G. Breton, R. Fournier and J.-P. Watte. 1994. Dinosaur egglaying and nesting in France. K. Carpenter, K.F. Hirsch & J.R. Horner (eds.) Dinosaur Eggs and Babies. Cambridge University Press, Cambridge 56-74
* J. Le Loeuff, E. Buffetaut, P. Méchin and A. Méchin-Salessy. 1992. The first record of dromaeosaurid dinosaurs (Saurischia, Theropoda) in the Maastrichtian of southern Europe: palaeobiogeographical implications. Bulletin de la Société géologique de la France 163(3):337-343
* R. Dughi and F. Sirugue. 1958. Sur les oeufs de dinosaures du bassin fluvio-lacustre de Basse-Provence. Comptes Rendus du Congrès National des Sociétés Savantes, Section des Sciences, Paris 83:183-20

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