List of Clubionidae species

Short description: none

Main article: Biology:Sac spider

This page lists all described species of the spider family Clubionidae accepted by the World Spider Catalog as of January 2021[update]:[1]

## Contents

* 1 A
* 1.1 Arabellata
* 2 C
* 2.1 Carteronius
* 2.2 † Chiapasona
* 2.3 Clubiona
* 2.4 Clubionina
* 3 D
* 3.1 † Desultor
* 4 E
* 4.1 Elaver
* 4.2 † Eobumbatrix
* 4.3 † Eodoter
* 4.4 † Eostentatrix
* 4.5 † Eoversatrix
* 5 I
* 5.1 Invexillata
* 6 M
* 6.1 † Machilla
* 6.2 Malamatidia
* 6.3 † Massula
* 6.4 Matidia
* 7 N
* 7.1 Nusatidia
* 8 P
* 8.1 Porrhoclubiona
* 8.2 Pristidia
* 8.3 † Prosocer
* 8.4 Pteroneta
* 9 S
* 9.1 Scopalio
* 9.2 Simalio
* 10 T
* 10.1 Tixcocoba
* 11 References

## A

### Arabellata

Arabellata Baert, Versteirt & Jocqué, 2010

* A. nimispalpata Baert, Versteirt & Jocqué, 2010 (type) — New Guinea
* A. terebrata Baert, Versteirt & Jocqué, 2010 — New Guinea

## C

### Carteronius

Carteronius Simon, 1897

* C. argenticomus (Keyserling, 1877) — Madagascar
* C. fuscus Simon, 1896 — Mauritius
* C. helluo Simon, 1896 (type) — Sierra Leone
* C. vittiger Simon, 1896 — Madagascar

### † Chiapasona

† Chiapasona Petrunkevitch, 1963[2]

* † C. defuncta Petrunkevitch, 1963

### Clubiona

Clubiona Latreille, 1804

* C. abbajensis Strand, 1906 — Ethiopia, Somalia, Central, East Africa
* C. a. kibonotensis Lessert, 1921 — East Africa
* C. abboti L. Koch, 1866 — USA, Canada
* C. a. abbotoides Chamberlin & Ivie, 1946 — USA
* C. aberrans Dankittipakul, 2012 — Thailand
* C. abnormis Dankittipakul, 2008 — Thailand, Laos
* C. acanthocnemis Simon, 1906 — India
* C. achilles Hogg, 1896 — Australia (Central)
* C. acies Nicolet, 1849 — Chile
* C. aciformis Zhang & Hu, 1991 — China
* C. aculeata Zhang, Zhu & Song, 2007 — China
* C. adjacens Gertsch & Davis, 1936 — USA
* C. aducta Simon, 1932 — Portugal, Spain, Italy
* C. africana Lessert, 1921 — East Africa
* C. akagiensis Hayashi, 1985 — Japan
* C. alexeevi Mikhailov, 1990 — Russia (Caucasus)
* C. aliceae Chickering, 1937 — Panama
* C. allotorta Dankittipakul & Singtripop, 2008 — Thailand
* C. alluaudi Simon, 1898 — Mauritius
* C. alpicola Kulczyński, 1882 — Europe to Central Asia
* C. alticola Dankittipakul & Singtripop, 2008 — Thailand
* C. altissimoides Liu, Yan, Griswold & Ubick, 2007 — China
* C. altissimus Hu, 2001 — China
* C. alveolata L. Koch, 1873 — Samoa, Funafuti, Marquesas Is., Hawaii
* C. amurensis Mikhailov, 1990 — Russia (Far East), Japan
* C. analis Thorell, 1895 — India, Bangladesh, Myanmar
* C. andreinii Caporiacco, 1936 — Italy
* C. angulata Dondale & Redner, 1976 — Canada
* C. annuligera Lessert, 1929 — Congo, Mozambique
* C. anwarae Biswas & Raychaudhuri, 1996 — Bangladesh
* C. apiata Urquhart, 1893 — Australia (Tasmania)
* C. apiculata Dankittipakul & Singtripop, 2014 — Malaysia (Borneo)
* C. applanata Liu, Yan, Griswold & Ubick, 2007 — China
* C. aspidiphora Simon, 1910 — South Africa
* C. asrevida Ono, 1992 — Taiwan
* C. auberginosa Zhang, Yin, Bao & Kim, 1997 — China
* C. australiaca Kolosváry, 1934 — Australia (New South Wales)
* C. bachmaensis Ono, 2009 — Vietnam
* C. bagerhatensis Biswas & Raychaudhuri, 1996 — Bangladesh
* C. baimaensis Song & Zhu, 1991 — China
* C. baishishan Zhang, Zhu & Song, 2003 — China
* C. bakurovi Mikhailov, 1990 — Russia (Far East), China, Korea
* C. bandoi Hayashi, 1995 — Japan
* C. basarukini Mikhailov, 1990 — Russia (South Siberia, Far East), Mongolia, Japan
* C. bashkirica Mikhailov, 1992 — Russia (Urals, West Siberia)
* C. batikanoides Barrion, Barrion-Dupo & Heong, 2013 — China (Hainan)
* C. bengalensis Biswas, 1984 — India
* C. bevisi Lessert, 1923 — South Africa
* C. biaculeata Simon, 1897 — South Africa
* C. bicornis Yu & Li, 2019 — China
* C. bicuspidata Wu & Zhang, 2014 — China
* C. biembolata Deeleman-Reinhold, 2001 — Malaysia (Borneo), Indonesia (Sumatra)
* C. bifissurata Kritscher, 1966 — New Caledonia
* C. biforamina Liu, Peng & Yan, 2016 — China
* C. bifurcata Zhang, Yu & Zhong, 2018 — China
* C. bilobata Dhali, Roy, Saha & Raychaudhuri, 2016 — India
* C. bipinnata Yu, Zhang & Chen, 2017 — China
* C. bishopi Edwards, 1958 — USA, Canada
* C. blesti Forster, 1979 — New Zealand
* C. bomiensis Zhang & Zhu, 2009 — China
* C. boxaensis Biswas & Biswas, 1992 — India
* C. brachyptera Zhu & Chen, 2012 — China (Hainan)
* C. brevipes Blackwall, 1841 — Europe, Caucasus, Japan?
* C. brevispina Huang & Chen, 2012 — Taiwan
* C. bryantae Gertsch, 1941 — USA, Canada
* C. bucera Yang, Ma & Zhang, 2011 — China
* C. bukaea (Barrion & Litsinger, 1995) — Philippines
* C. cada Forster, 1979 — New Zealand
* C. caerulescens L. Koch, 1867 — Europe, Russia (Europe to Far East), Kazakhstan, China, Japan
* C. californica Fox, 1938 — USA
* C. calycina Wu & Zhang, 2014 — China
* C. cambridgei L. Koch, 1873 — New Zealand
* C. campylacantha Dankittipakul, 2008 — Thailand
* C. canaca Berland, 1930 — New Caledonia
* C. canadensis Emerton, 1890 — USA, Canada
* C. canberrana Dondale, 1966 — Australia (New South Wales)
* C. candefacta Nicolet, 1849 — Chile
* C. caohai Zhang & Yu, 2020 — China
* C. capensis Simon, 1897 — South Africa
* C. catawba Gertsch, 1941 — USA
* C. caucasica Mikhailov & Otto, 2017 — Caucasus (Georgia, Armenia, Azerbaijan, Russia), Turkey
* C. chabarovi Mikhailov, 1991 — Russia (Far East)
* C. chakrabartei Majumder & Tikader, 1991 — India
* C. charitonovi Mikhailov, 1990 — Russia (South Siberia, Far East)
* C. charleneae Barrion & Litsinger, 1995 — Philippines
* C. chathamensis Simon, 1905 — New Zealand (Chatham Is.)
* C. cheni Yu & Li, 2019 — China
* C. chevalieri Berland, 1936 — Cape Verde Is.
* C. chikunii Hayashi, 1986 — Japan
* C. chippewa Gertsch, 1941 — USA, Canada
* C. circulata Zhang & Yin, 1998 — China
* C. cirrosa Ono, 1989 — Japan (Ryukyu Is.)
* C. citricolor Lawrence, 1952 — South Africa
* C. clima Forster, 1979 — New Zealand
* C. cochlearis Yu & Li, 2019 — China
* C. cochleata Wang, Wu & Zhang, 2015 — China
* C. complicata Banks, 1898 — Mexico
* C. comta C. L. Koch, 1839 — Europe, North Africa, Turkey, Caucasus
* C. concinna (Thorell, 1887) — Myanmar
* C. congentilis Kulczyński, 1913 — Central Europe to Central Asia
* C. conica Dankittipakul & Singtripop, 2014 — Malaysia (Borneo)
* C. consensa Forster, 1979 — New Zealand
* C. contaminata O. Pickard-Cambridge, 1872 — Israel
* C. contrita Forster, 1979 — New Zealand
* C. convoluta Forster, 1979 — New Zealand
* C. cordata Zhang & Zhu, 2009 — China
* C. coreana Paik, 1990 — Russia (Far East), China, Korea
* C. corrugata Bösenberg & Strand, 1906 — Russia (Far East), China, Taiwan, Korea, Japan, Thailand
* C. corticalis (Walckenaer, 1802) — Europe, Turkey, Caucasus
* C. c. concolor Kulczyński, 1897 — Hungary
* C. c. nigra Simon, 1878 — France
* C. cultrata Dankittipakul & Singtripop, 2014 — Indonesia (Borneo)
* C. cycladata Simon, 1909 — Australia (Western Australia)
* C. cylindrata Liu, Yan, Griswold & Ubick, 2007 — China
* C. cylindriformis Dankittipakul & Singtripop, 2014 — Malaysia (Borneo)
* C. dactylina Liu, Peng & Yan, 2016 — China
* C. dakong Zhang & Yu, 2020 — China
* C. damirkovaci Deeleman-Reinhold, 2001 — Malaysia
* C. debilis Nicolet, 1849 — Chile
* C. deletrix O. Pickard-Cambridge, 1885 — India, China, Taiwan, Japan
* C. delicata Forster, 1979 — New Zealand
* C. denticulata Dhali, Roy, Saha & Raychaudhuri, 2016 — India
* C. desecheonis Petrunkevitch, 1930 — Puerto Rico
* C. deterrima Strand, 1904 — Norway
* C. dichotoma Wang, Chen & Z. S. Zhang, 2018 — China
* C. didentata Zhang & Yin, 1998 — China
* C. digitata Dankittipakul, 2012 — Thailand
* C. dikita Barrion & Litsinger, 1995 — Philippines
* C. diversa O. Pickard-Cambridge, 1862 — Europe, Caucasus, Russia (Europe to Far East), Kazakhstan, Pakistan, Korea, Japan
* C. drassodes O. Pickard-Cambridge, 1874 — India, Bangladesh, China
* C. dubia O. Pickard-Cambridge, 1870 — St. Helena
* C. dunini Mikhailov, 2003 — Russia (Far East)
* C. duoconcava Zhang & Hu, 1991 — China
* C. durbana Roewer, 1951 — South Africa
* C. dyasia Gertsch, 1941 — USA
* C. dysderiformis (Guérin, 1838) — New Guinea
* C. elaphines Urquhart, 1893 — Australia (Tasmania)
* C. ericius Chrysanthus, 1967 — New Guinea
* C. eskovi Mikhailov, 1995 — Russia (Far East)
* C. estes Edwards, 1958 — USA
* C. esuriens Thorell, 1897 — Myanmar
* C. evoronensis Mikhailov, 1995 — Russia (Far East)
* C. excavata (Rainbow, 1920) — Australia (Lord Howe Is.)
* C. excisa O. Pickard-Cambridge, 1898 — Mexico
* C. ezoensis Hayashi, 1987 — Russia (Far East), Japan
* C. facilis O. Pickard-Cambridge, 1911 — Britain
* C. falcata Tang, Song & Zhu, 2005 — China, Mongolia
* C. falciforma Liu, Peng & Yan, 2016 — China
* C. fanjingshan Wang, Chen & Z. S. Zhang, 2018 — China
* C. femorocalcarata Huang & Chen, 2012 — Taiwan
* C. filicata O. Pickard-Cambridge, 1874 — Pakistan, India, Bangladesh, Myanmar, Thailand, Laos, China
* C. filifera Dankittipakul, 2008 — Thailand
* C. filoramula Zhang & Yin, 1998 — China
* C. flavocincta Nicolet, 1849 — Chile
* C. forcipa Yang, Song & Zhu, 2003 — China
* C. frisia Wunderlich & Schuett, 1995 — Europe to Central Asia
* C. frutetorum L. Koch, 1867 — Europe to Central Asia
* C. furcata Emerton, 1919 — North America, Russia (Middle to East Siberia)
* C. fusoidea Zhang, 1992 — China
* C. fuzhouensis Gong, 1985 — China
* C. gallagheri Barrion & Litsinger, 1995 — Indonesia (Java)
* C. germanica Thorell, 1871 — Europe, Caucasus, Russia (Europe to Far East), Central Asia
* C. gertschi Edwards, 1958 — USA
* C. gilva O. Pickard-Cambridge, 1872 — Israel
* C. giulianetti Rainbow, 1898 — New Guinea
* C. glatiosa Saito, 1934 — Japan
* C. globosa Wang, Chen & Z. S. Zhang, 2018 — China
* C. godfreyi Lessert, 1921 — East Africa
* C. golovatchi Mikhailov, 1990 — Russia (Europe), Caucasus
* C. gongi Zhang, Yin, Bao & Kim, 1997 — China
* C. gongshan He, Liu & Zhang, 2016 — China
* C. grucollaris Yu, Zhang & Chen, 2017 — China
* C. guianensis Caporiacco, 1947 — Guyana
* C. haeinsensis Paik, 1990 — Russia (Far East), China, Korea, Japan
* C. haplotarsa Simon, 1909 — São Tomé and Príncipe
* C. hatamensis (Thorell, 1881) — New Guinea
* C. haupti Tang, Song & Zhu, 2005 — China
* C. hedini Schenkel, 1936 — China
* C. helenae Mikhailov, 2003 — Russia (Far East)
* C. helva Simon, 1897 — South Africa
* C. heteroducta Zhang & Yin, 1998 — China
* C. heterosaca Yin, Yan, Gong & Kim, 1996 — China
* C. hexadentata Dhali, Roy, Saha & Raychaudhuri, 2016 — India
* C. hilaris Simon, 1878 — Mountains of Spain, France, Italy, Austria, Switzerland, North Macedonia and Romania
* C. hindu Deeleman-Reinhold, 2001 — Indonesia (Bali)
* C. hitchinsi Saaristo, 2002 — Seychelles, French Polynesia (Tuamotu)
* C. hoffmanni Schenkel, 1937 — Madagascar
* C. hooda Dong & Zhang, 2016 — China
* C. huaban Xin, Zhang, Li, Zheng & Yu, 2020 — China
* C. hugispaa Barrion & Litsinger, 1995 — Philippines
* C. hugisva Barrion & Litsinger, 1995 — Philippines
* C. huiming Wang, F. Zhang & Z. S. Zhang, 2018 — China
* C. hummeli Schenkel, 1936 — China
* C. hundeshageni Strand, 1907 — Indonesia (Moluccas)
* C. huttoni Forster, 1979 — New Zealand
* C. hwanghakensis Paik, 1990 — Korea
* C. hyrcanica Mikhailov, 1990 — Azerbaijan, Iran
* C. hysgina Simon, 1889 — India
* C. hystrix Berland, 1938 — Indonesia (Lesser Sunda Is.), Vanuatu
* C. iharai Ono, 1995 — Japan
* C. ikedai Ono, 1992 — Japan
* C. inaensis Hayashi, 1989 — Japan
* C. inquilina Deeleman-Reinhold, 2001 — Malaysia (Borneo)
* C. insulana Ono, 1989 — Taiwan, Japan (Ryukyu Is.)
* C. interjecta L. Koch, 1879 — Russia (West Siberia to Far East), Mongolia, China
* C. irinae Mikhailov, 1991 — Russia (Far East), China, Korea
* C. jaegeri Ono, 2011 — Palau Is.
* C. janae Edwards, 1958 — USA
* C. japonica L. Koch, 1878 — Russia (Sakhalin, Kurile Is.), China, Korea, Taiwan, Japan
* C. japonicola Bösenberg & Strand, 1906 — Russia (Far East) to Philippines, Indonesia
* C. jiandan Yu & Li, 2019 — China
* C. jiulongensis Zhang, Yin & Kim, 1996 — China
* C. johnsoni Gertsch, 1941 — USA, Canada
* C. jucunda (Karsch, 1879) — Russia (Far East), China, Korea, Taiwan, Japan
* C. juvenis Simon, 1878 — Europe, Iran, Uzbekistan
* C. kagani Gertsch, 1941 — USA
* C. kai Jäger & Dankittipakul, 2010 — China, Laos
* C. kaltenbachi Kritscher, 1966 — New Caledonia
* C. kapataganensis Barrion & Litsinger, 1995 — Philippines
* C. kasanensis Paik, 1990 — Korea, Japan
* C. kastoni Gertsch, 1941 — USA, Canada
* C. kasurensis Mukhtar & Mushtaq, 2005 — Pakistan
* C. katioryza Barrion & Litsinger, 1995 — Philippines
* C. kayashimai Ono, 1994 — Taiwan
* C. kiboschensis Lessert, 1921 — East Africa
* C. kigabensis Strand, 1915 — East Africa
* C. kimyongkii Paik, 1990 — Russia (Far East), China, Korea
* C. kiowa Gertsch, 1941 — North America
* C. komissarovi Mikhailov, 1992 — Russia (Far East), Korea
* C. kowong Chrysanthus, 1967 — New Guinea
* C. krisisensis Barrion & Litsinger, 1995 — Philippines, Indonesia (Borneo)
* C. kropfi Zhang, Zhu & Song, 2003 — China
* C. kuanshanensis Ono, 1994 — Taiwan
* C. kularensis Marusik & Koponen, 2002 — Russia (north-eastern Siberia, Far East)
* C. kulczynskii Lessert, 1905 — North America, Europe, Kazakhstan, Russia (Europe to Far East), Japan
* C. kumadaorum Ono, 1992 — Japan
* C. kunashirensis Mikhailov, 1990 — Russia (Sakhalin, Kurile Is.), Japan
* C. kurenshikovi Mikhailov, 1995 — Russia (Far East)
* C. kurilensis Bösenberg & Strand, 1906 — Russia (Far East), China, Taiwan, Korea, Japan
* C. kurosawai Ono, 1986 — Taiwan, Korea, Japan
* C. kuu Jäger & Dankittipakul, 2010 — Laos
* C. lala Jäger & Dankittipakul, 2010 — Laos
* C. lamellaris Zhang, Yu & Zhong, 2018 — China
* C. lamina Zhang, Zhu & Song, 2007 — China
* C. langei Mikhailov, 1991 — Russia (Far East)
* C. latericia Kulczyński, 1926 — Russia (Middle Siberia to Far East), USA (Alaska)
* C. laticeps O. Pickard-Cambridge, 1885 — China (Yarkand)
* C. latitans Pavesi, 1883 — Ethiopia, Somalia, Kenya
* C. laudabilis Simon, 1909 — Australia (Western Australia)
* C. lawrencei Roewer, 1951 — South Africa
* C. lena Bösenberg & Strand, 1906 — China, Korea, Japan
* C. leonilae Barrion & Litsinger, 1995 — Philippines
* C. leptosa Zhang, Yin, Bao & Kim, 1997 — China
* C. limpida Simon, 1897 — South Africa
* C. linea Xie, Yin, Yan & Kim, 1996 — China
* C. linzhiensis Hu, 2001 — China
* C. lirata Yang, Song & Zhu, 2003 — China
* C. littoralis Banks, 1895 — USA, Canada
* C. logunovi Mikhailov, 1990 — Russia (Far East)
* C. longipes Nicolet, 1849 — Chile
* C. luapalana Giltay, 1935 — Congo
* C. lucida He, Liu & Zhang, 2016 — China
* C. ludhianaensis Tikader, 1976 — India, Bangladesh
* C. lutescens Westring, 1851 — Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Kazakhstan, Korea, Japan. Introduced to North America
* C. lyriformis Song & Zhu, 1991 — China
* C. maculata Roewer, 1951 — Australia (Queensland)
* C. mahensis Simon, 1893 — Seychelles
* C. maipai Jäger & Dankittipakul, 2010 — Thailand
* C. mandschurica Schenkel, 1953 — Russia (Far East), China, Korea, Japan
* C. manshanensis Zhu & An, 1988 — China
* C. maracandica Kroneberg, 1875 — Uzbekistan
* C. maritima L. Koch, 1867 — USA, Canada, Caribbean
* C. marmorata L. Koch, 1866 — France to Ukraine and Turkey
* C. marna Roddy, 1966 — USA
* C. marusiki Mikhailov, 1990 — Russia (Far East)
* C. maya Hayashi & Yoshida, 1991 — Japan
* C. maysangarta Barrion & Litsinger, 1995 — Philippines
* C. mayumiae Ono, 1993 — Russia (Far East), Korea, Japan
* C. mazandaranica Mikhailov, 2003 — Azerbaijan, Iran
* C. medog Zhang, Zhu & Song, 2007 — China
* C. melanosticta Thorell, 1890 — Thailand, Indonesia (Sumatra, Krakatau), New Guinea
* C. melanothele Thorell, 1895 — Myanmar, Thailand, Laos, Indonesia (Sumatra)
* C. meraukensis Chrysanthus, 1967 — Malaysia, New Guinea
* C. microsapporensis Mikhailov, 1990 — Russia (Far East), Korea
* C. mikhailovi Deeleman-Reinhold, 2001 — Indonesia (Java)
* C. milingae Barrion-Dupo, Barrion & Heong, 2013 — China (Hainan)
* C. mimula Chamberlin, 1928 — USA, Canada
* C. minima (Ono, 2010) — Japan
* C. minuscula Nicolet, 1849 — Chile
* C. minuta Nicolet, 1849 — Chile
* C. mixta Emerton, 1890 — USA, Canada
* C. modesta L. Koch, 1873 — Australia (Queensland)
* C. moesta Banks, 1896 — USA, Canada, China
* C. moralis Song & Zhu, 1991 — China, Taiwan
* C. mordica O. Pickard-Cambridge, 1898 — Mexico
* C. mujibari Biswas & Raychaudhuri, 1996 — Bangladesh
* C. multidentata Liu, Peng & Yan, 2016 — China
* C. munda Thorell, 1887 — Myanmar
* C. munis Simon, 1909 — Australia (Western Australia)
* C. mutata Gertsch, 1941 — USA, Canada
* C. mutilata Bösenberg & Strand, 1906 — Japan
* C. mykolai Mikhailov, 2003 — Ukraine
* C. nataliae Trilikauskas, 2007 — Russia (Far East)
* C. natalica Simon, 1897 — South Africa
* C. neglecta O. Pickard-Cambridge, 1862 — Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asia, China, Korea
* C. neglectoides Bösenberg & Strand, 1906 — China, Korea, Japan
* C. nemorum Ledoux, 2004 — Réunion
* C. nenilini Mikhailov, 1995 — Russia (South Siberia)
* C. neocaledonica Berland, 1924 — New Caledonia
* C. newnani Ivie & Barrows, 1935 — USA
* C. nicholsi Gertsch, 1941 — USA
* C. nicobarensis Tikader, 1977 — India (Nicobar Is.)
* C. nigromaculosa Blackwall, 1877 — Seychelles, Réunion
* C. nilgherina Simon, 1906 — India
* C. ningpoensis Schenkel, 1944 — China
* C. nollothensis Simon, 1910 — South Africa
* C. norvegica Strand, 1900 — North America, Europe, Russia (Europe to West Siberia)
* C. notabilis L. Koch, 1873 — Australia (Queensland)
* C. obesa Hentz, 1847 — USA, Canada
* C. oceanica Ono, 2011 — Japan
* C. octoginta Dankittipakul, 2008 — Thailand
* C. odelli Edwards, 1958 — USA
* C. odesanensis Paik, 1990 — Russia (Far East), China, Korea
* C. ogatai Ono, 1995 — Japan
* C. oligerae Mikhailov, 1995 — Russia (Far East)
* C. opeongo Edwards, 1958 — Canada
* C. orientalis Mikhailov, 1995 — North Korea
* C. oteroana Gertsch, 1941 — USA
* C. ovalis Zhang, 1991 — China
* C. pacifica Banks, 1896 — USA, Canada
* C. paenuliformis (Strand, 1916) — Ghana
* C. pahilistapyasea Barrion & Litsinger, 1995 — Thailand, Indonesia (Borneo), Philippines
* C. paiki Mikhailov, 1991 — Russia (Far East)
* C. pala Deeleman-Reinhold, 2001 — Indonesia (Moluccas)
* C. pallidula (Clerck, 1757) (type) — Europe, Caucasus, Russia (Europe to Far East), Central Asia. Introduced to North America
* C. pantherina Chrysanthus, 1967 — New Guinea
* C. papillata Schenkel, 1936 — Russia (Far East), China, Korea
* C. papuana Chrysanthus, 1967 — New Guinea
* C. paralena Mikhailov, 1995 — North Korea
* C. parallela Hu & Li, 1987 — China
* C. paranghinlalakirta Barrion & Litsinger, 1995 — Philippines
* C. parangunikarta Barrion & Litsinger, 1995 — Philippines
* C. parconcinna Deeleman-Reinhold, 2001 — Thailand, Indonesia (Borneo)
* C. parva Seo, 2018 — Korea
* C. parvula Saito, 1933 — Japan
* C. peculiaris L. Koch, 1873 — New Zealand
* C. phansa Strand, 1911 — Indonesia (Aru Is.)
* C. phragmitis C. L. Koch, 1843 — Morocco, Algeria, Europe, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China, Korea
* C. phragmitoides Schenkel, 1963 — China
* C. pianmaensis Wang, Wu & Zhang, 2015 — China
* C. picturata Deeleman-Reinhold, 2001 — Indonesia (Bali)
* C. pikei Gertsch, 1941 — USA, Canada
* C. pila Dhali, Roy, Saha & Raychaudhuri, 2016 — India
* C. plumbi Gertsch, 1941 — USA
* C. pogonias Simon, 1906 — India
* C. pollicaris Wu, Zheng & Zhang, 2015 — China
* C. pomoa Gertsch, 1941 — USA
* C. pongolensis Lawrence, 1952 — South Africa
* C. pototanensis Barrion & Litsinger, 1995 — Philippines
* C. praematura Emerton, 1909 — North America, Russia (Far East)
* C. procera Chrysanthus, 1967 — New Guinea
* C. procteri Gertsch, 1941 — USA
* C. producta Forster, 1979 — New Zealand
* C. propinqua L. Koch, 1879 — Russia (Middle Siberia to Far East), North Korea, China
* C. proszynskii Mikhailov, 1995 — North Korea
* C. pruvotae Berland, 1930 — New Caledonia
* C. pseudocordata Dhali, Roy, Saha & Raychaudhuri, 2016 — India
* C. pseudogermanica Schenkel, 1936 — Russia (Far East), China, Korea, Japan
* C. pseudomaxillata Hogg, 1915 — New Guinea
* C. pseudoneglecta Wunderlich, 1994 — Morocco, Algeria, Europe, Caucasus, Iran
* C. pseudopteroneta Raven & Stumkat, 2002 — Australia (Queensland)
* C. pseudosaxatilis Mikhailov, 1992 — Russia (Central Asia, South Siberia), Kazakhstan
* C. pseudosimilis Mikhailov, 1990 — Algeria, Portugal, Greece (Crete), Caucasus
* C. pterogona Yang, Song & Zhu, 2003 — China
* C. puera Nicolet, 1849 — Chile
* C. pupillaris Lawrence, 1938 — South Africa
* C. pupula Thorell, 1897 — Myanmar
* C. pygmaea Banks, 1892 — USA, Canada
* C. pyrifera Schenkel, 1936 — China
* C. qianhuayuani Barrion, Barrion-Dupo & Heong, 2013 — China (Hainan)
* C. qini Tang, Song & Zhu, 2005 — China
* C. qiyunensis Xu, Yang & Song, 2003 — China
* C. quebecana Dondale & Redner, 1976 — USA, Canada
* C. rainbowi Roewer, 1951 — Australia (Lord Howe Is.)
* C. rama Dankittipakul & Singtripop, 2008 — India, Thailand, China
* C. ramoiensis (Thorell, 1881) — New Guinea
* C. rava Simon, 1886 — Senegal
* C. reclusa O. Pickard-Cambridge, 1863 — Europe, Turkey, Russia (Europe to South Siberia), Kazakhstan
* C. revillioidi Lessert, 1936 — South Africa, Mozambique
* C. rhododendri Barrows, 1945 — USA
* C. rileyi Gertsch, 1941 — USA
* C. riparia L. Koch, 1866 — Russia (Urals to Far East), Mongolia, China, Japan, North America
* C. risbeci Berland, 1930 — New Caledonia
* C. rivalis Pavesi, 1883 — Ethiopia
* C. robusta L. Koch, 1873 — Australia
* C. roeweri Caporiacco, 1940 — Ethiopia
* C. rosserae Locket, 1953 — Britain, France, Netherlands, Poland, Slovakia, Hungary, Romania
* C. rostrata Paik, 1985 — Russia (Far East), China, Korea, Japan
* C. rothschildi Berland, 1922 — Ethiopia
* C. rumpiana Lawrence, 1952 — South Africa
* C. rybini Mikhailov, 1992 — Kazakhstan, Kyrgyzstan
* C. ryukyuensis Ono, 1989 — Japan (Ryukyu Is.)
* C. saltitans Emerton, 1919 — USA, Canada
* C. saltuum Kulczyński, 1898 — Austria
* C. samoensis Berland, 1929 — Samoa, French Polynesia (Society Is., Austral Is.: Rapa)
* C. sapporensis Hayashi, 1986 — Russia (Far East), Korea, Japan
* C. saurica Mikhailov, 1992 — Kazakhstan
* C. savesi Berland, 1930 — New Caledonia
* C. saxatilis L. Koch, 1867 — France to Poland and south-eastern Europe
* C. scandens Deeleman-Reinhold, 2001 — Malaysia (Borneo)
* C. scatula Forster, 1979 — New Zealand
* C. scenica Nicolet, 1849 — Chile
* C. semicircularis Tang, Song & Zhu, 2005 — China
* C. sertungensis Hayashi, 1996 — Indonesia (Krakatau)
* C. shillongensis Majumder & Tikader, 1991 — India
* C. sichotanica Mikhailov, 2003 — Russia (Far East)
* C. sigillata Lawrence, 1952 — South Africa
* C. silvestris Deeleman-Reinhold, 2001 — Malaysia (Borneo)
* C. similis L. Koch, 1867 — Europe, Turkey, Caucasus
* C. sjostedti Lessert, 1921 — East Africa
* C. s. spinigera Lessert, 1921 — East Africa
* C. sopaikensis Paik, 1990 — Russia (Far East), Korea
* C. sparassella Strand, 1909 — South Africa
* C. spiralis Emerton, 1909 — USA, Canada
* C. stagnatilis Kulczyński, 1897 — Europe, Caucasus, Russia (Europe to South Siberia), Central Asia
* C. stiligera Deeleman-Reinhold, 2001 — Indonesia (Sumatra)
* C. straminea O. Pickard-Cambridge, 1872 — Israel
* C. subapplanata Wang, Chen & Z. S. Zhang, 2018 — China
* C. subasrevida Yu & Li, 2019 — China
* C. subborealis Mikhailov, 1992 — Russia (South Siberia, Far East), Mongolia
* C. subcylindrica Wang, Chen & Z. S. Zhang, 2018 — China
* C. subkuu Yu & Li, 2019 — China
* C. submaculata (Thorell, 1891) — India (Nicobar Is.)
* C. submoralis Wu, Zheng & Zhang, 2015 — China
* C. subnotabilis Strand, 1907 — Australia
* C. subparallela Zhang, Zhu & Song, 2007 — China
* C. subquebecana Yu & Li, 2019 — China
* C. subrama Yu & Li, 2019 — China
* C. subrostrata Zhang & Hu, 1991 — China
* C. subsultans Thorell, 1875 — Europe, Russia (Europe to South Siberia), Japan
* C. subtilis L. Koch, 1867 — Europe, Russia (Europe to Far East), Kyrgyzstan, Korea
* C. subtrivialis Strand, 1906 — Ethiopia, East Africa
* C. subyaginumai Yu & Li, 2019 — China
* C. subyangmingensis Gan & Wang, 2020 — China
* C. suthepica Dankittipakul, 2008 — Thailand
* C. tabupumensis Petrunkevitch, 1914 — Myanmar
* C. taiwanica Ono, 1994 — China, Taiwan
* C. tangi Liu, Peng & Yan, 2016 — China
* C. tanikawai Ono, 1989 — China, Taiwan, Japan (Ryukyu Is.)
* C. tateyamensis Hayashi, 1989 — Japan
* C. tenera (Thorell, 1890) — Indonesia (Sumatra, Java)
* C. tengchong Zhang, Zhu & Song, 2007 — China
* C. ternatensis (Thorell, 1881) — Indonesia (Moluccas)
* C. terrestris Westring, 1851 — Europe (without Russia), Turkey
* C. theoblicki Yu & Li, 2019 — China
* C. thorelli Roewer, 1951 — Indonesia (Sumatra)
* C. tiane Yu & Li, 2019 — China
* C. tiantongensis Zhang, Yin & Kim, 1996 — China
* C. tikaderi Majumder & Tikader, 1991 — India
* C. tongdaoensis Zhang, Yin, Bao & Kim, 1997 — China, Korea
* C. tongi Yu & Li, 2019 — China
* C. topakea Barrion & Litsinger, 1995 — Philippines
* C. torta Forster, 1979 — New Zealand
* C. tortuosa Zhang & Yin, 1998 — China
* C. transbaicalica Mikhailov, 1992 — Russia (South Siberia)
* C. tridentata Dhali, Roy, Saha & Raychaudhuri, 2016 — India
* C. trivialis C. L. Koch, 1843 — North America, Europe, Russia (Europe to Far East), China, Japan
* C. tsurusakii Hayashi, 1987 — Russia (Kurile Is.), Japan
* C. uenoi Ono, 1986 — Japan
* C. umbilensis Lessert, 1923 — South Africa
* C. unanoa Barrion & Litsinger, 1995 — Philippines
* C. unikarta Barrion & Litsinger, 1995 — Philippines
* C. upoluensis Marples, 1964 — Samoa
* C. vachoni Lawrence, 1952 — South Africa
* C. vacuna L. Koch, 1873 — New Guinea, Australia (Queensland)
* C. valens Simon, 1897 — South Africa
* C. venatoria Rainbow & Pulleine, 1920 — Australia (Lord Howe Is.)
* C. venusae Barrion & Litsinger, 1995 — Philippines
* C. venusta Paik, 1985 — China, Korea
* C. victoriaensis Barrion & Litsinger, 1995 — Philippines
* C. vigil Karsch, 1879 — Russia (Kurile Is.), Korea, Japan, China
* C. vigillella Strand, 1918 — Japan
* C. violaceovittata Schenkel, 1936 — China
* C. vukomi Jäger & Dankittipakul, 2010 — Thailand, Laos
* C. wolongica Zhu & An, 1999 — China
* C. wulingensis Yu & Chen, 2017 — China
* C. xinwenhui Barrion, Barrion-Dupo & Heong, 2013 — China (Hainan)
* C. yaginumai Hayashi, 1989 — Taiwan, Japan
* C. yangmingensis Hayashi & Yoshida, 1993 — Taiwan
* C. yanzhii Zhang & Yu, 2020 — China
* C. yaoi Yu & Li, 2019 — China
* C. yaroslavi Mikhailov, 2003 — Russia (Far East)
* C. yasudai Ono, 1991 — Japan
* C. yoshidai Hayashi, 1989 — Japan
* C. yueya Yu & Li, 2019 — China
* C. yurii Mikhailov, 2011 — Mongolia
* C. zacharovi Mikhailov, 1991 — Russia (Far East), Korea
* C. zandstrai Barrion & Litsinger, 1995 — Philippines
* C. zhanggureni Yu & Li, 2019 — China
* C. zhangmuensis Hu & Li, 1987 — China
* C. zhangyongjingi Li & Blick, 2019 — China
* C. zhengi Yu & Li, 2019 — China
* C. zhui Xu, Yang & Song, 2003 — China
* C. zilla Dönitz & Strand, 1906 — Japan
* C. zimmermanni Marples, 1964 — Samoa
* C. zyuzini Mikhailov, 1995 — Russia (Far East)
* † C. arcana Scudder, 1890
* † C. attenuata Koch and Berendt, 1854
* † C. curvispinosa Petrunkevitch, 1922
* † C. eversa Scudder, 1890
* † C. florissanti Petrunkevitch, 1922
* † C. lanata Koch and Berendt, 1854
* † C. microphthalma Koch and Berendt, 1854
* † C. pubescens Koch and Berendt, 1854
* † C. sericea Koch and Berendt, 1854
* † C. tomentosa Koch and Berendt, 1854

### Clubionina

Clubionina Berland, 1947

* C. pallida Berland, 1947 (type) — St. Paul Is.

## D

### † Desultor

† Desultor Petrunkevitch, 1942[2]

* † D. depressus Petrunkevitch, 1942

## E

### Elaver

Elaver O. Pickard-Cambridge, 1898

* E. achuca (Roddy, 1966) — USA
* E. albicans (Franganillo, 1930) — Cuba, Jamaica
* E. arawakan Saturnino & Bonaldo, 2015 — Haiti
* E. balboae (Chickering, 1937) — Panama to Brazil, Cuba
* E. barroana (Chickering, 1937) — Panama
* E. beni Saturnino & Bonaldo, 2015 — Peru, Brazil, Bolivia
* E. brevipes (Keyserling, 1891) — Brazil, Argentina
* E. calcarata (Kraus, 1955) — Mexico, El Salvador, Costa Rica
* E. candelaria Saturnino & Bonaldo, 2015 — Mexico
* E. carlota (Bryant, 1940) — Cuba
* E. chisosa (Roddy, 1966) — USA
* E. crinophora (Franganillo, 1934) — Cuba
* E. crocota (O. Pickard-Cambridge, 1896) — Mexico
* E. darwichi Saturnino & Bonaldo, 2015 — Panama, Ecuador
* E. depuncta O. Pickard-Cambridge, 1898 — Mexico
* E. elaver (Bryant, 1940) — Cuba
* E. excepta (L. Koch, 1866) — USA, Canada, Caribbean
* E. grandivulva (Mello-Leitão, 1930) — Brazil, Bolivia
* E. helenae Saturnino & Bonaldo, 2015 — Mexico
* E. hortoni (Chickering, 1937) — Panama
* E. implicata (Gertsch, 1941) — Hispaniola
* E. juana (Bryant, 1940) — Cuba, Bahama Is.
* E. juruti Saturnino & Bonaldo, 2015 — Brazil
* E. kawitpaaia (Barrion & Litsinger, 1995) — Philippines
* E. kohlsi (Gertsch & Jellison, 1939) — USA
* E. linguata (F. O. Pickard-Cambridge, 1900) — Guatemala
* E. lizae Saturnino & Bonaldo, 2015 — Costa Rica
* E. lutescens (Schmidt, 1971) — Panama to Brazil
* E. madera (Roddy, 1966) — USA
* E. mirabilis (O. Pickard-Cambridge, 1896) — Mexico, Belize, Nicaragua
* E. mulaiki (Gertsch, 1935) — USA
* E. multinotata (Chickering, 1937) — Costa Rica, Panama, Colombia, Venezuela, Peru, Brazil
* E. orvillei (Chickering, 1937) — Panama
* E. placida O. Pickard-Cambridge, 1898 — Mexico
* E. portoricensis (Petrunkevitch, 1930) — Puerto Rico, Virgin Is.
* E. quadrata (Kraus, 1955) — El Salvador
* E. richardi (Gertsch, 1941) — Honduras
* E. sericea O. Pickard-Cambridge, 1898 — Mexico
* E. shinguito Saturnino & Bonaldo, 2015 — Colombia, Peru, Brazil
* E. sigillata (Petrunkevitch, 1925) — Panama, Colombia, Peru, Brazil
* E. simplex (O. Pickard-Cambridge, 1896) — Guatemala
* E. tenera (Franganillo, 1935) — Cuba
* E. tenuis (Franganillo, 1935) — Cuba
* E. texana (Gertsch, 1933) — USA, Mexico
* E. tigrina O. Pickard-Cambridge, 1898 (type) — Mexico, Costa Rica
* E. tourinhoae Saturnino & Bonaldo, 2015 — Colombia, Brazil
* E. tricuspis (F. O. Pickard-Cambridge, 1900) — Guatemala, Panama
* E. tristani (Banks, 1909) — Costa Rica
* E. tumivulva (Banks, 1909) — Costa Rica
* E. turongdaliriana (Barrion & Litsinger, 1995) — Philippines
* E. valvula (F. O. Pickard-Cambridge, 1900) — Panama
* E. vieirae Saturnino & Bonaldo, 2015 — Brazil, Peru
* E. wheeleri (Roewer, 1933) — USA, Mexico
* † E. nutua Wunderlich, 1988

### † Eobumbatrix

† Eobumbatrix Petrunkevitch, 1922[2]

* † E. latebrosa Scudder, 1890

### † Eodoter

† Eodoter Petrunkevitch, 1958[2]

* † E. eopala Wunderlich, 2004
* † E. longimammillae Wunderlich, 2012
* † E. magnificus Petrunkevitch, 1958
* † E. scutatus Wunderlich, 2011
* † E. tibialis Wunderlich, 2011

### † Eostentatrix

† Eostentatrix Petrunkevitch, 1922[2]

* † E. cockerelli Petrunkevitch, 1922
* † E. ostentata Scudder, 1890

### † Eoversatrix

† Eoversatrix Petrunkevitch, 1922[2]

* † E. eversa Scudder, 1890

## I

### Invexillata

Invexillata Versteirt, Baert & Jocqué, 2010

* I. caerulea Versteirt, Baert & Jocqué, 2010 — New Guinea
* I. maculata Versteirt, Baert & Jocqué, 2010 (type) — New Guinea
* I. viridiflava Versteirt, Baert & Jocqué, 2010 — New Guinea

## M

### † Machilla

† Machilla Petrunkevitch, 1958[2]

* † M. setosa Petrunkevitch, 1958

### Malamatidia

Malamatidia Deeleman-Reinhold, 2001

* M. bohorokensis Deeleman-Reinhold, 2001 (type) — Indonesia (Sumatra, Borneo)
* M. christae Jäger & Dankittipakul, 2010 — Laos
* M. thorelli Deeleman-Reinhold, 2001 — Indonesia (Sulawesi)
* M. vethi Deeleman-Reinhold, 2001 — Malaysia, Indonesia (Borneo)
* M. zu Jäger & Dankittipakul, 2010 — Laos

### † Massula

† Massula Petrunkevitch, 1942[2]

* † M. klebsi Petrunkevitch, 1942

### Matidia

Matidia Thorell, 1878

* M. bipartita Deeleman-Reinhold, 2001 — Indonesia (Moluccas)
* M. calcarata Thorell, 1878 — Indonesia (Ambon)
* M. chlora Chrysanthus, 1967 — New Guinea
* M. flagellifera Simon, 1897 — Sri Lanka
* M. incurvata Reimoser, 1934 — India
* M. mas Deeleman-Reinhold, 2001 — Thailand
* M. missai Versteirt, Baert & Jocqué, 2010 — New Guinea
* M. muju Chrysanthus, 1967 — New Guinea
* M. paranga (Barrion & Litsinger, 1995) — Philippines
* M. simia Deeleman-Reinhold, 2001 — Indonesia (Sulawesi)
* M. simplex Simon, 1897 — Sri Lanka
* M. spatulata Chen & Huang, 2006 — Taiwan
* M. strobbei Versteirt, 2010 — New Guinea
* M. trinotata Thorell, 1890 — Malaysia
* M. virens Thorell, 1878 (type) — Indonesia (Moluccas, Sulawesi)
* M. viridissima Strand, 1911 — Indonesia (Aru Is.)

## N

### Nusatidia

Nusatidia Deeleman-Reinhold, 2001

* N. aeria (Simon, 1897) — Borneo
* N. bimaculata (Simon, 1897) — Sri Lanka
* N. borneensis Deeleman-Reinhold, 2001 — Indonesia (Sumatra, Borneo)
* N. camouflata Deeleman-Reinhold, 2001 — Thailand
* N. javana (Simon, 1897) (type) — Indonesia (Java, Krakatau)
* N. luzonica (Simon, 1897) — Philippines
* N. manipisea (Barrion & Litsinger, 1995) — Philippines
* N. melanobursa Deeleman-Reinhold, 2001 — Indonesia (Sumatra)
* N. pandalira Barrion-Dupo, Barrion & Heong, 2013 — China
* N. rama Deeleman-Reinhold, 2001 — Indonesia (Sumatra)
* N. snazelli Deeleman-Reinhold, 2001 — Indonesia (Java, Sumatra)
* N. vietnamensis Logunov & Jäger, 2015 — Vietnam

## P

### Porrhoclubiona

Porrhoclubiona Lohmander, 1944

* P. bosmansi Marusik & Omelko, 2018 — Tajikistan
* P. decora (Blackwall, 1859) — Madeira, Azores
* P. diniensis (Simon, 1878) — Algeria, Morocco, Tunisia, Portugal, Spain, France, Italy
* P. genevensis (L. Koch, 1866) (type) — Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asia
* P. laudata (O. Pickard-Cambridge, 1885) — China
* P. leucaspis (Simon, 1932) — Europe, Algeria, Morocco, Tunisia
* P. minor (Wunderlich, 1987) — Canary Is.
* P. moradmandi Marusik & Omelko, 2018 — Iran
* P. pseudominor (Wunderlich, 1987) — Canary Is.
* P. pteronetoides (Deeleman-Reinhold, 2001) — Thailand
* P. vegeta (Simon, 1918) — Canary Is., North Africa, Southern Europe, Caucasus, Iran
* P. viridula (Ono, 1989) — China, Taiwan, Thailand, Japan (Ryukyu Is.), Indonesia (Lesser Sunda Is.)
* P. wunderlichi (Mikhailov, 1992) — Mongolia

### Pristidia

Pristidia Deeleman-Reinhold, 2001

* P. cervicornuta Yu, Zhang & Chen, 2017 — China (Hainan)
* P. longistila Deeleman-Reinhold, 2001 — Malaysia (Borneo)
* P. prima Deeleman-Reinhold, 2001 (type) — Thailand, Malaysia (Peninsula), Indonesia (Sumatra, Java)
* P. ramosa Yu, Sun & Zhang, 2012 — China, Taiwan
* P. secunda Deeleman-Reinhold, 2001 — Indonesia (Sumatra)
* P. viridissima Deeleman-Reinhold, 2001 — Thailand, Malaysia, Indonesia (Sumatra, Borneo, Java)

### † Prosocer

† Prosocer Petrunkevitch, 1963[2]

* † P. mollis Petrunkevitch, 1963

### Pteroneta

Pteroneta Deeleman-Reinhold, 2001

* P. baiteta Versteirt, Deeleman-Reinhold & Baert, 2008 — New Guinea
* P. brevichela Versteirt, Deeleman-Reinhold & Baert, 2008 — New Guinea
* P. longichela Versteirt, Deeleman-Reinhold & Baert, 2008 — New Guinea
* P. madangiensis Versteirt, Deeleman-Reinhold & Baert, 2008 — New Guinea
* P. saltans Deeleman-Reinhold, 2001 (type) — Malaysia, Indonesia (Sulawesi, Lesser Sunda Is.), Borneo
* P. spinosa Raven & Stumkat, 2002 — Australia (Queensland)
* P. tertia Deeleman-Reinhold, 2001 — Singapore, Indonesia (Borneo, Sulawesi)
* P. ultramarina (Ono, 1989) — Japan (Ryukyu Is.)

## S

### Scopalio

Scopalio Deeleman-Reinhold, 2001

* S. verrens Deeleman-Reinhold, 2001 (type) — Borneo

### Simalio

Simalio Simon, 1897

* S. aurobindoi Patel & Reddy, 1991 — India
* S. biswasi Majumder & Tikader, 1991 — India
* S. castaneiceps Simon, 1906 — India
* S. lucorum Simon, 1906 — Sri Lanka
* S. percomis Simon, 1906 — India
* S. petilus Simon, 1897 (type) — Philippines
* S. phaeocephalus Simon, 1906 — Sri Lanka
* S. rubidus Simon, 1897 — Trinidad

## T

### Tixcocoba

Tixcocoba Gertsch, 1977

* T. maya Gertsch, 1977 (type) — Mexico

## References

1. ↑ "Family: Clubionidae Wagner,1887". Natural History Museum Bern. 2021. doi:10.24436/2. http://www.wsc.nmbe.ch/family/17. Retrieved 2021-02-08.
2. ↑ 2.0 2.1 2.2 2.3 2.4 2.5 2.6 2.7 2.8 Behrensmeyer, A. K.; Turner, A. (2013). "Taxonomic occurrences of Suidae recorded in the Paleobiology Database". Fossilworks. http://www.fossilworks.org/cgi-bin/bridge.pl?action=home.

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