Branta

The black geese of the genus Branta are waterfowl belonging to the true geese and swans subfamily Anserinae. They occur in the northern coastal regions of the Palearctic and all over North America, migrating to more southerly coasts in winter, and as resident birds in the Hawaiian Islands. Alone in the Southern Hemisphere, a self-sustaining feral population derived from introduced Canada geese is also found in New Zealand.

The black geese derive their vernacular name for the prominent areas of black coloration found in all species. They can be distinguished from all other true geese by their legs and feet, which are black or very dark grey. Furthermore, they have black bills and large areas of black on the head and neck, with white (ochre in one species) markings that can be used to tell apart most species.[note 1] As with most geese, their undertail and uppertail coverts are white. They are also on average smaller than other geese, though some very large taxa are known, which rival the swan goose and the black-necked swan in size.

The Eurasian species of black geese have a more coastal distribution compared to the grey geese (genus Anser) which share the same general area of occurrence, not being found far inland even in winter (except for occasional stray birds or individuals escaped from captivity). This does not hold true for the American and Pacific species, in whose ranges grey geese are, for the most part, absent.

## Contents

* 1 Taxonomy
* 1.1 Species list
* 1.2 Early fossil record
* 2 Footnotes
* 3 References
* 4 Further reading

## Taxonomy

The genus Branta was introduced by the Austrian naturalist Giovanni Antonio Scopoli in 1769.[1] The name is a Latinised form of Old Norse Brandgás meaning burnt as in "burnt (black) goose".[2] The type species is the brant goose (Branta bernicla).[3]

Ottenburghs and colleagues published a study in 2016 that established the phylogenetic relationships between the species.[4]

Branta |  |  |

Brant (Branta bernicla) 60 px

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Red-breasted goose (Branta ruficollis) 60 px

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Nene (Branta sandvicensis) 60 px

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Canada goose (Branta canadensis) 60 px

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Barnacle goose (Branta leucopsis) 60 px

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Cackling goose (Branta hutchinsii) 60 px

### Species list

The genus contains six living species.[5]

Genus Branta  – Scopoli,, 1769 – six species  Common name  | Scientific name and subspecies  | Range  | Size and ecology  | IUCN status and estimated population

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Brant (U.S.) or brent goose (U.K.)

| Branta bernicla
(Linnaeus, 1758)

Three subspecies

* B. b. nigricans (Lawrence, 1846)
* B. b. bernicla (Linnaeus, 1758)
* B. b. hrota (Müller, OF, 1776)

| {{{range}}}
| Size:

Habitat:

Diet:  |  LC

Red-breasted goose

| Branta ruficollis
(Pallas, 1769) | Breeds in Arctic Siberia, mainly on the Taymyr Peninsula, with smaller populations in the Gydan and Yamal Peninsulas; wintering on northwestern shores of the Black Sea in Bulgaria, Romania and Ukraine, as well as in Azerbaijan
| Size:

Habitat:

Diet:  |  VU

Nene, nēnē, or Hawaiian goose

| Branta sandvicensis
(Vigors, 1834) | Hawaiian Islands  | Size:

Habitat:

Diet:  |  NT

Canada goose

| Branta canadensis
(Linnaeus, 1758)

Seven subspecies

* Atlantic Canada goose (B. c. canadensis) (Linnaeus, 1758)
* Interior Canada goose (B. c. interior) (Todd, 1938)
* Giant Canada goose (B. c. maxima) (Delacour, 1951)
* Moffitt's Canada goose (B. c. moffitti) (Aldrich, 1946)
* Vancouver Canada goose (B. c. fulva) (Delacour, 1951)
* Dusky Canada goose (B. c. occidentalis) (Baird, 1858)
* Lesser Canada goose (B. c. parvipes) (Cassin, 1852)

| Temperate regions of North America, introduced populations in Western and northern Europe, Japan, New Zealand, Chile, Argentina and the Falkland Islands
| Size:

Habitat:

Diet:  |  LC

Barnacle goose

| Branta leucopsis
(Bechstein, 1803) | Breeds in Arctic Russia, eastern Greenland and northern Europe; wintering in northern and northwestern Europe.
| Size:

Habitat:

Diet:  |  LC

Cackling goose

| Branta hutchinsii
(Richardson, 1832)

Four or five subspecies

* Richardson's cackling goose (B. h. hutchinsii)
* Aleutian cackling goose (B. h. leucopareia)
* Small cackling goose (B. h. minima)
* Taverner's cackling goose (B. h. taverneri)
* †Bering cackling goose (B. h. asiatica)[disputed]

| North America, northern Canada and Alaska  | Size:

Habitat:

Diet:  |  LC

Two species have been described from subfossil remains found in the Hawaiian Islands, where they became extinct in prehistoric times:

* †Nēnē-nui or wood-walking goose, Branta hylobadistes (prehistoric)

Similar but hitherto undescribed remains are also known from Kauaʻi and Oʻahu.[6]

* †Giant Hawaii goose, Branta rhuax (prehistoric), formerly Geochen rhuax

The relationships of the enigmatic Geochen rhuax, formerly known only from parts of a single bird's skeleton damaged due to apparently dying in a lava flow, were long unresolved. After reexamination of the subfossil material and comparisons with other subfossil bones from the island of Hawaiʻi assigned to the genus Branta, it was redescribed as Branta rhuax in 2013.[7] While a presumed relation between B. rhuax and the shelducks, proposed by Lester Short in 1970,[8] has thus been refuted, bones of a shelduck-like bird have been found more recently on Kaua‘i.[9]

Similarly, two bones found on Oʻahu indicate the erstwhile presence of a gigantic waterfowl on this island. Its relationships relative to this genus and the moa-nalos, enormous goose-like dabbling ducks, are completely undeterminable at present.[6]

### Early fossil record

Several fossil species of Branta have been described. Since the true geese are hardly distinguishable by anatomical features, the allocation of these to this genus is somewhat uncertain.

* Branta woolfendeni Bickart 1990 (Late Miocene of Arizona, USA)[10]
* Branta thessaliensis Boev & Koufos, 2006 (Late Miocene of Perivolaki, Greece)
* Branta dickeyi Miller 1924 (Late Pliocene – Late Pleistocene of W USA)[8][11][12][13]
* Branta esmeralda Burt 1929 (Early Pliocene of Nevada, USA)[12][13][14]
* Branta howardae Miller 1930 (Early Pliocene of California, USA)[13][14][15][16]
* Branta propinqua Shufeldt 1892 (Middle Pleistocene of Fossil Lake, Oregon, USA)[8][13]
* Branta hypsibata (Cope) 1878 (Middle Pleistocene of Fossil Lake, Oregon, USA)[8][12][13]

The former "Branta" minuscula is now placed with the prehistoric American shelducks, Anabernicula.[13] On the other hand, a goose fossil from the Early-Middle Pleistocene of El Salvador is highly similar to Anser and given its age and biogeography it is likely to belong to that genus or Branta.[17]

## Footnotes

1. ↑ The nēnē, which is aberrant in many respects, has no white on the head or neck and fairly little black, being quite similar to the swan goose in the color pattern of these areas, and was thus formerly assigned to the monotypic genus Nesochen. The swan goose, a grey goose, also has a black bill, but its reddish-orange legs indicate its actual relationships.

## References

1. ↑ Scopoli, Giovanni Antonio (1769) (in Latin). Annus Historico-Naturalis. Part 1. Lipsiae [Leipzig]: Sumtib. C.G. Hilscheri. p. 67. https://www.biodiversitylibrary.org/page/31117656.
2. ↑ Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 77. ISBN 978-1-4081-2501-4. https://archive.org/details/Helm_Dictionary_of_Scientific_Bird_Names_by_James_A._Jobling.
3. ↑ Mayr, Ernst; Cottrell, G. William, eds (1979). Check-List of Birds of the World. 1 (2nd ed.). Cambridge, Massachusetts: Museum of Comparative Zoology. p. 440. https://www.biodiversitylibrary.org/page/16109080.
4. ↑ Ottenburghs, J.; Megens, H.-J.; Kraus, R.H.S.; Madsen, O.; van Hooft, P.; van Wieren, S.E.; Crooijmans, R.P.M.A.; Ydenberg, R.C. et al. (2016). "A tree of geese: A phylogenomic perspective on the evolutionary history of True Geese". Molecular Phylogenetics and Evolution 101: 303–313. doi:10.1016/j.ympev.2016.05.021. PMID 27233434. Bibcode: 2016MolPE.101..303O. http://nbn-resolving.de/urn:nbn:de:bsz:352-0-379362.
5. ↑ Gill, Frank; Donsker, David; Rasmussen, Pamela, eds (2020). "Screamers, ducks, geese & swans". IOC World Bird List Version 10.2. International Ornithologists' Union. http://www.worldbirdnames.org/bow/waterfowl/.
6. ↑ 6.0 6.1 Olson, Storrs L.; James, Helen F. (1991). "Descriptions of thirty-two new species of birds from the Hawaiian Islands: Part I. Non-Passeriformes". Ornithological Monographs 45 (45): 48. doi:10.2307/40166794. https://repository.si.edu/handle/10088/1745.
7. ↑ Olson, Storrs L. (2013). "Hawaii's first fossil bird: history, geological age, and taxonomic status of the extinct goose Geochen rhuax Wetmore (Aves: Anatidae).". Proceedings of the Biological Society of Washington (Washington: Biological Society of Washington) 126 (2): 161–168. doi:10.2988/0006-324x-126.2.161. http://www.bioone.org/doi/abs/10.2988/0006-324X-126.2.161. Retrieved 13 July 2013.
8. ↑ 8.0 8.1 8.2 8.3 Short, Lester L. (1970). "A new anseriform genus and species from the Nebraska Pliocene". Auk 87 (3): 537–543. doi:10.2307/4083796. http://sora.unm.edu/sites/default/files/journals/auk/v087n03/p0537-p0543.pdf.
9. ↑ Burney, D.A.; James, H.F.; Burney, L.P.; Olson, S.L.; Kikuchi, W.; Wagner, W.L.; Burney, W.; McCloskey, D. et al. (2001). "Fossil evidence for a diverse biota from Kaua'i and its transformation since human arrival". Ecological Monographs 71 (4): 615–641. doi:10.1890/0012-9615(2001)071[0615:FEFADB2.0.CO;2]. Bibcode: 2001EcoM...71..615B. https://repository.si.edu/handle/10088/109.
10. ↑ Bickart, K.J. (1990). "Part I: The birds of the late Miocene–early Pliocene Big Sandy Formation, Mohave County, Arizona". Ornithological Monographs 44 (44): 1–72. doi:10.2307/40166673. https://www.jstor.org/stable/40166673.
11. ↑ Miller, Loye (1924). "Branta dickeyi from the McKittrick Pleistocene". The Condor 26 (5): 178–180. doi:10.2307/1363171. http://sora.unm.edu/sites/default/files/journals/condor/v026n05/p0178-p0180.pdf.
12. ↑ 12.0 12.1 12.2 Miller, Loye (1944). "Some Pliocene Birds from Oregon and Idaho". The Condor 46 (1): 25–32. doi:10.2307/1364248. http://sora.unm.edu/sites/default/files/journals/condor/v046n01/p0025-p0032.pdf.
13. ↑ 13.0 13.1 13.2 13.3 13.4 13.5 Short, Lester L. (1969). "A new genus and species of gooselike swan from the Pliocene of Nebraska". American Museum Novitates (2369).
14. ↑ 14.0 14.1 Miller, Alden H.; Ashley, James F. (1934). "Goose Footprints on a Pliocene Mud-flat". The Condor 36 (4): 178–179. doi:10.2307/1363416. http://sora.unm.edu/sites/default/files/journals/condor/v036n04/p0178-p0179.pdf.
15. ↑ Howard, Hildegarde (1931). "Pliocene Bird Remains from Santa Barbara, California". The Condor 33 (1): 30–31. doi:10.2307/1363932. http://sora.unm.edu/sites/default/files/journals/condor/v033n01/p0030-p0031.pdf.
16. ↑ Miller, Loye (1931). "Bird Remains from the Kern River Pliocene of California". The Condor 33 (2): 70–72. doi:10.2307/1363312. http://sora.unm.edu/sites/default/files/journals/condor/v033n02/p0070-p0072.pdf.
17. ↑ Cisneros, Juan Carlos (2005). "New Pleistocene vertebrate fauna from El Salvador" (in English, Portuguese). Revista Brasileira de Paleontologia 8 (3): 239–255. doi:10.4072/rbp.2005.3.09. Bibcode: 2005RvBrP...8..239C. http://www.sbpbrasil.org/revista/edicoes/8_3/cisneros.pdf.

## Further reading

* Carboneras, Carles (1992). "Family Anatidae (Ducks, Geese and Swans)". in del Hoyo, Josep. Handbook of Birds of the World. Volume 1: Ostrich to Ducks. Barcelona: Lynx Edicions. pp. 536–629, plates 40–50. ISBN 84-87334-10-5.

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*[†]: Extinct