Endomychidae

Endomychidae, also commonly known as handsome fungus beetles, is a family of coccinelloid beetles with representatives found in all biogeographic realms. As the common name suggests, members of this family feed on fungi.

## Contents

* 1 Description
* 2 Taxonomy
* 2.1 Genera
* 2.2 Fossil genera
* 3 References
* 4 External links

## Description

They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment.

## Taxonomy

The family was established based on the type genus Endomychus, a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea.

Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi.[1] A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae.[2]

This family contains around 120 genera and 1300 species. A list of subfamilies can be seen below:

The subfamilies that are included:

* Danascelinae
* Endomychinae (including Stenotarsinae)
* Epipocinae
* Leiestinae
* Lycoperdininae
* Merophysiinae
* Pleganophorinae
* Xenomycetinae

### Genera

These 70 genera belong to the family Endomychidae as defined in the past (due to revisions, this may not be entirely accurate):

* Acinaces Gerstaecker, 1858 g
* Aclemmysa Reitter, 1904 g
* Agaricophilus Motschulsky, 1838 g
* Amphisternus Germar, 1843 g
* Amphistethus Strohecker, 1964 g
* Ancylopus Costa, 1854 g
* Anidrytus Gerstaecker, 1858 i c g
* Aphorista Gorham, 1873 i c g b
* Archipines Strohecker, 1953 i c g
* Asymbius Gorham, 1896 g
* Atopomychus Tomaszewska & Szawaryn, 2013 g
* Atrichonota Arrow, 1925 g
* Avencymon Strohecker, 1971 g
* Beccariola Arrow, 1943 g
* Bolbomorphus Gorham, 1888 g
* Brachytrycherus Arrow, 1920 g
* Bystodes Strohecker, 1953 g
* Cholovocera Motschulsky, 1838 g
* Chondria Gorham, 1888 g
* Corynomalus Chevrolat in Dejean, 1836 i c g
* Cyclotoma Mulsant, 1851 i g
* Danae Reiche, 1847 i c g b
* Danascelis Tomaszewska, 1999 i c g
* Dapsa Latreille, 1829 g
* Dexialia Sasaji, 1970 g
* Dialexia Gorham, 1891 g
* Discolomopsis Shockley, 2006 g
* Displotera Reitter, 1877 g
* Ectomychus Gorham, 1888 g
* Eidorus g
* Endomychus Panzer, 1795 i c g b
* Ephebus Chevrolat in Dejean, 1836 i c g
* Epipocus Germar, 1843 i c g b
* Epopterus Chevrolat in Dejean, 1836 i c g
* Eucteanus Gerstaecker, 1857 g
* Eucymon g
* Eumorphus Weber, 1801 g
* Geoendomychus Lea, 1922 g
* Glesirhanis Shockley & Alekseev, 2014 g
* Hadromychus Bousquet & Leschen, 2002 i c g b
* Holoparamecus Curtis, 1833 i c g b
* Hylaia Guerin, 1857 g
* Hylaperdina Tomaszewska, 2012 g
* Idiophyes Blackburn, 1896 g
* Indalmus Gerstaecker, 1858 g
* Leiestes Chevrolat, 1836 g
* Lycoperdina Latreille, 1807 i c g b
* Meilichius Gerstaecker, 1857 g
* Merophysia Lucas, 1852 g
* Microxenus Wallaston, 1861 g
* Mycetina Mulsant, 1846 i c g b
* Mychothenus Strohecker, 1953 g
* Natalinus Tomaszewska, 2011 g
* Ohtaius Chûjô, 1938 g
* Palaeoestes Kirejtshuk & Nel, 2009 g
* Paniegena Heller, 1916 g
* Parasymbius Arrow, 1920 g
* Phymaphora Newman, 1838 i c g b
* Phymaphoroides Motschoulsky, 1857 g
* Pleganophorus Hampe, 1855 g
* Polymus Mulsant, 1846 g
* Pseudindalmus Arrow, 1920 g
* Rhanidea Strohecker, 1953 i c g b
* Saula Gerstaecker, 1858 i c g
* Sinocymbachus Strohecker & Chujo, 1970 g
* Sinopanamomus Esser, 2019
* Stenotarsus Perty, 1832 i c g b
* Stethorhanis Blaisdell, 1931 i c g
* Trochoideus Westwood, 1833 i c g b
* Trycherus Gerstaecker, 1857 g
* Xenomycetes Horn, 1880 i c g b

Data sources: i = ITIS,[3] c = Catalogue of Life,[4] g = GBIF,[5] b = Bugguide.net[6]

### Fossil genera

* Burmalestes Tomaszewska and Ślipiński, 2018[7] Burmese amber, Myanmar, Late Cretaceous (Cenomanian)
* Cretolestes Tomaszewska, Ślipiński and Ren, 2018[7] Burmese amber, Myanmar, Cenomanian
* Cretaparamecus Tomaszewska, Ślipiński, Bai and Zhang, 2018[7] Burmese amber, Myanmar, Cenomanian
* Palaeomycetes Tomaszewska, Ślipiński and Ren, 2018[7] Burmese amber, Myanmar, Cenomanian
* Zemyna Tomaszewska, 2018[7] (replaced name Laima Alekseev and Tomaszewska, 2018): Baltic amber, Eocene

## References

1. ↑ Tomaszewska, K.Wioletta (2000). "Morphology, phylogeny and classification of adult endomychidae (Coleoptera: Cucujoidea)". Annales Zoologici 50 (4): 449–558. http://rcin.org.pl/Content/56456/WA058_74047_P255-T50_Annal-Zool-nr-4-1.pdf.
2. ↑ Robertson, James A; Ślipiński, Adam; Moulton, Matthew; Shockley, Floyd W; Giorgi, Adriano; Lord, Nathan P; McKenna, Duane D; Tomaszewska, Wioletta et al. (2015). "Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)". Systematic Entomology 40 (4): 745. doi:10.1111/syen.12138. Bibcode: 2015SysEn..40..745R. https://repository.si.edu/bitstream/handle/10088/21810/ent_Shockley_Tomaszewska_McHugh_2009b.pdf.
3. ↑ "Endomychidae Report". https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=114369.
4. ↑ "Browse Endomychidae". http://www.catalogueoflife.org/col/browse/tree/id/a4a75dd67401cc5ba261577dec2e7526.
5. ↑ "Endomychidae". https://www.gbif.org/species/7828.
6. ↑ "Endomychidae Family Information". https://bugguide.net/node/view/13860.
7. ↑ 7.0 7.1 7.2 7.3 7.4 Tomaszewska, Wioletta; Ślipiński, Adam; Bai, Ming; Zhang, Weiwei; Ren, Dong (November 2018). "The oldest representatives of Endomychidae (Coleoptera: Coccinelloidea) from the Upper Cretaceous Burmese amber" (in en). Cretaceous Research 91: 287–298. doi:10.1016/j.cretres.2018.07.001. Bibcode: 2018CrRes..91..287T. https://linkinghub.elsevier.com/retrieve/pii/S0195667118301605.

## External links

* "Strohecker Collection — Amphix". University of Georgia. http://www.ent.uga.edu/mchugh/Strohecker Files/Strohecker Genera Pages/Amphix.htm.
* "Endomychidae of Wisconsin". University of Wisconsin. http://entomology.wisc.edu/~mbprice/wibeetles/Price/Index.htm.

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Extant Coleoptera families

* Kingdom: Animalia
* Phylum: Arthropoda
* Class: Insecta
* Subclass: Pterygota
* Infraclass: Neoptera
* Superorder: Holometabola

Suborder Archostemata

* Crowsoniellidae (Crowsoniella relicta)
* Cupedidae (reticulated beetles)
* Jurodidae (Sikhotealinia zhiltzovae)
* Micromalthidae
* Ommatidae

Suborder Adephaga

| Extant families|

* Amphizoidae (trout-stream beetles)
* Aspidytidae
* Carabidae (ground beetles)
* Cicindelidae (tiger beetles)
* Dytiscidae (predaceous diving beetles)
* Gyrinidae (whirligig beetles)
* Haliplidae (crawling water beetles)
* Hygrobiidae
* Meruidae (Meru phyllisae)
* Noteridae (burrowing water beetles)
* Trachypachidae (false ground beetles)

|

Suborder Myxophaga

* Hydroscaphidae (skiff beetles)
* Lepiceridae
* Sphaeriusidae
* Torridincolidae

Suborder Polyphaga

| Bostrichiformia|

| Bostrichoidea|

* Bostrichidae (auger beetles)
* Dermestidae (skin beetles)
* Endecatomidae
* Jacobsoniidae (Jacobson's beetles)
* Nosodendridae (wounded-tree beetles)
* Ptiniidae (furniture beetles, death watch beetles, spider beetles)

|

Derodontoidea|

* Derodontidae (tooth-necked fungus beetles)

Cucujiformia|

| Chrysomeloidea|

* Cerambycidae (longhorn beetles)
* Chrysomelidae (leaf beetles)
* Disteniidae
* Megalopodidae
* Orsodacnidae
* Oxypeltidae
* Vesperidae

|

Cleroidea|

* Acanthocnemidae (Acanthocnemus nigricans)
* Biphyllidae (false skin beetles)
* Byturidae (fruitworm beetles)
* Chaetosomatidae
* Cleridae (checkered beetles)
* Lophocateridae
* Mauroniscidae
* Melyridae (soft-wing flower beetles)
* Metaxinidae (Metaxina ornata)
* Peltidae
* Phloiophilidae (Phloiophilus edwardsi)
* Phycosecidae
* Prionoceridae
* Protopeltidae
* Rentoniidae
* Rhadalidae
* Thanerocleridae
* Thymalidae
* Trogossitidae (bark-gnawing beetles)

Coccinelloidea|

* Akalyptoischiidae
* Alexiidae
* Anamorphidae
* Bothrideridae (dry bark beetles)
* Cerylonidae (minute bark beetles)
* Coccinellidae (lady beetles, or God's cows)
* Corylophidae (minute fungus beetles)
* Discolomatidae
* Endomychidae (handsome fungus beetles)
* Eupsilobiidae
* Euxestidae (well polished beetles)
* Latridiidae (minute brown scavenger beetles)
* Murmidiidae
* Mycetaeidae
* Teredidae

Cucujoidea|

* Agapythidae (Agapytho foveicollis)
* Boganiidae
* Cavognathidae
* Cryptophagidae (silken fungus beetles)
* Cucujidae (flat bark beetles)
* Cybocephalidae
* Cyclaxyridae
* Erotylidae (pleasing fungus beetles)
* Helotidae
* Hobartiidae
* Kateretidae (short-winged flower beetles)
* Laemophloeidae (lined flat bark beetles)
* Lamingtoniidae (Lamingtonium binnaberrense)
* Monotomidae (root-eating beetles)
* Myraboliidae
* Nitidulidae (sap beetles)
* Passandridae (parasitic flat bark beetles)
* Phalacridae (shining flower beetles)
* Phloeostichidae
* Priasilphidae
* Propalticidae
* Protocucujidae
* Silvanidae (silvanid flat bark beetles)
* Smicripidae (palmetto beetles)
* Sphindidae (dry-fungus beetles)
* Tasmosalpingidae

Curculionoidea
(weevils)|

* Anthribidae (fungus weevils)
* Attelabidae (leaf-rolling weevils)
* Belidae (primitive weevils)
* Brentidae (straight snout weevils, New York weevil)
* Caridae
* Curculionidae (true weevils, bark beetles, ambrosia beetles)
* Nemonychidae (pine flower weevils)

Lymexyloidea|

* Lymexylidae (ship-timber beetles)

Tenebrionoidea|

* Aderidae (ant-like leaf beetles)
* Anthicidae (ant-like flower beetles)
* Archeocrypticidae (cryptic fungus beetles)
* Boridae (conifer bark beetles)
* Chalcodryidae
* Ciidae (minute tree-fungus beetles)
* Melandryidae (false darkling beetles)
* Meloidae (blister beetles)
* Mordellidae (tumbling flower beetles)
* Mycetophagidae (hairy fungus beetles)
* Mycteridae (palm and flower beetles)
* Oedemeridae (false blister beetle)
* Perimylopidae, or Promecheilidae
* Prostomidae (jugular-horned beetles)
* Pterogeniidae
* Pyrochroidae (fire-coloured beetles)
* Pythidae (dead log bark beetles)
* Ripiphoridae (wedge-shaped beetles)
* Salpingidae (narrow-waisted bark beetles)
* Scraptiidae (false flower beetles)
* Stenotrachelidae (false longhorn beetles)
* Synchroidae (synchroa bark beetles)
* Tenebrionidae (darkling beetles)
* Tetratomidae (polypore fungus beetles)
* Trictenotomidae
* Ulodidae
* Zopheridae (ironclad beetles, cylindrical bark beetles)

Elateriformia|

| Buprestoidea|

* Buprestidae (jewel beetles, or metallic wood-boring beetles)
* Schizopodidae

|

Byrrhoidea|

* Byrrhidae (pill beetles)
* Callirhipidae (cedar beetles)
* Chelonariidae (turtle beetles)
* Cneoglossidae
* Dryopidae (long-toed water beetles)
* Elmidae (riffle beetles)
* Eulichadidae (forest stream beetles)
* Heteroceridae (variegated mud-loving beetles)
* Limnichidae (minute mud beetles)
* Lutrochidae (travertine beetles)
* Psephenidae (water-penny beetles)
* Ptilodactylidae

Dascilloidea|

* Dascillidae (soft bodied plant beetles)
* Rhipiceridae (cicada beetle, cicada parasite beetles)

Elateroidea|

* Artematopodidae (soft-bodied plant beetles)
* Brachypsectridae (Texas beetles)
* Cantharidae (soldier beetles)
* Cerophytidae (rare click beetles)
* Elateridae (click beetles)
* Eucnemidae (false click beetles)
* Jurasaidae
* Lampyridae (fireflies)
* Lycidae (net-winged beetles)
* Omethidae (false fireflies, long-lipped beetles)
* Phengodidae (glowworm beetles)
* Rhagophthalmidae
* Sinopyrophoridae
* Throscidae (false metallic wood-boring beetles)

Rhinorhipoidea|

* Rhinorhipidae (Rhinorhipus tamborinensis)

Scirtoidea|

* Clambidae
* Decliniidae (Declinia relicta)
* Eucinetidae (plate-thigh beetles)
* Scirtidae

Scarabaeiformia|

| Scarabaeoidea|

* Belohinidae (Belohina inexpectata)
* Bolboceratidae
* Diphyllostomatidae (false stag beetles)
* Geotrupidae (dor beetles)
* Glaphyridae (bumble bee scarab beetles)
* Glaresidae (enigmatic scarab beetles)
* Hybosoridae (scavenger scarab beetles)
* Lucanidae (stag beetles)
* Ochodaeidae (sand-loving scarab beetles)
* Passalidae (betsy beetles)
* Pleocomidae (rain beetles)
* Scarabaeidae (scarabs)
* Trogidae (hide beetles)

|

Staphyliniformia|

| Histeroidea|

* Histeridae (clown beetles)
* Sphaeritidae (false clown beetles)
* Synteliidae

|

Hydrophiloidea|

* Epimetopidae
* Georissidae (minute mud-loving beetles)
* Helophoridae
* Hydrochidae
* Hydrophilidae (water scavenger beetles)
* Spercheidae

Staphylinoidea|

* Agyrtidae (primitive carrion beetles)
* Hydraenidae
* Leiodidae (round fungus beetles)
* Ptiliidae (feather-winged beetles)
* Staphylinidae (rove beetles)

List of subgroups of the order Coleoptera

Wikidata ☰ Q1076939 entry

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