2018 in paleobotany

Overview of the events of 2018 in paleobotany

List of years in paleobotany

* …
* 2015
* 2016
* 2017
* 2018
* 2019
* 2020
* 2021
* …

In paleontology
2015
2016
2017
2018
2019
2020
2021

In arthropod paleontology
2015
2016
2017
2018
2019
2020
2021

In paleoentomology
2015
2016
2017
2018
2019
2020
2021

In paleomalacology
2015
2016
2017
2018
2019
2020
2021

In reptile paleontology
2015
2016
2017
2018
2019
2020
2021

In archosaur paleontology
2015
2016
2017
2018
2019
2020
2021

In mammal paleontology
2015
2016
2017
2018
2019
2020
2021

In paleoichthyology
2015
2016
2017
2018
2019
2020
2021

* Art
* Archaeology
* Architecture
* Literature
* Music
* Philosophy
* Science +...

This article records new taxa of plants described during the year 2018, as well as other significant discoveries and events related to paleobotany that occurred in 2018.

## Flowering plants

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Notes  | Images

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Alloberberis axelrodii[1]

Sp. nov

Valid

Doweld

Miocene

|  |

United States
( Nevada)

A member of the family Berberidaceae; a replacement name for the previously invalidly published Mahonia sinuata Axelrod (1985), lacking holotype designation when published.

Alloberberis caeruleomontana[1]

Nom. nov

Valid

Doweld

Miocene

|  |

United States
( Oregon)

A member of the family Berberidaceae; a replacement name for Ilex sinuata Chaney & Axelrod (1959).

Anacolosidites eosenonicus[2]

Sp. nov

Valid

Arai & Dias-Brito

Late Cretaceous (Santonian)

São Carlos Formation

Brazil

A pollen taxon, possibly a member of the family Loranthaceae.

Aniba caucasica[3]

Nom. nov

Valid

Doweld

Pliocene

|  |

Abkhazia

A species of Aniba; a replacement name for Aniba longifolia Kolakovsky & Schakryl (1958).

Anisodromum upchurchii[4]

Sp. nov

Valid

Wang & Dilcher

Early Cretaceous (Albian)

Dakota Formation

United States
( Kansas)

A rosid described on the basis of fossil leaves.

Annona nepalensis[5]

Sp. nov

Valid

Prasad et al.

Miocene

Churia Formation

Nepal

A species of Annona.

Araliaephyllum popovii[6]

Sp. nov

Valid

Golovneva

Early Cretaceous (Albian)

|  |

Russia

A member of Laurales described on the basis of fossil leaves.

Archeampelos betulifolia[7]

Sp. nov

Valid

Moiseeva, Kodrul & Herman

Paleocene

Zeya–Bureya Basin

Russia

A flowering plant described on the basis of fossil leaves, similar to leaves of members of the family Betulaceae.

Austrovideira[8]

Gen. et sp. nov

Valid

Rozefelds & Pace

Early Oligocene

|  |

Australia

A member of Vitaceae. Genus includes new species A. dettmannae.

Berberis miopannonica[1]

Nom. nov

Valid

Doweld

Miocene

|  |

Romania

A species of Berberis; a replacement name for Berberis lanceolata Givulescu (1985).

Berberis notata[1]

Nom. nov

Valid

Doweld

Miocene

|  |

Austria

A species of Berberis; a replacement name for Ilex ambigua Unger (1847) and Berberis ambigua Kovar-Eder & Kvaček (2004).

Berryoxylon[9]

Gen. et sp. nov

Valid

Awasthi, Mehrotra & Shukla

Late Miocene–early Pliocene

Cuddalore Sandstone Formation

India

A fossil wood showing affinities with members of the genus Berrya. Genus includes new species B. cuddalorensis.

Bignonioxylon[10]

Gen. et sp. nov

Valid

Moya & Brea

Late Pleistocene

Arroyo Feliciano Formation

Argentina

A member of Bignoniaceae described on the basis of fossil wood. Genus includes new species B. americanum.

Burretiodendron guangxiensis[11]

Sp. nov

Valid

Dong & Sun in Dong et al.

Oligocene

Ningming Formation

China

A species of Burretiodendron.

Buxus pliosinica[12]

Sp. nov

Valid

Huang, Su & Zhou

Late Pliocene

Sanying Formation

China

A species of Buxus.

Canarium guangxiensis[13]

Sp. nov

Valid

Han & Manchester in Han et al.

Late Oligocene to late Miocene

Erzitang Formation
Foluo Formation
Yongning Formation

China

A species of Canarium

Carlquistoxylon australe[14]

Sp. nov

Valid

Pujana et al.

Early Cretaceous (late Albian)

Cerro Barcino Formation

Argentina

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil wood.

Castanopsis guangxiensis[15]

Sp. nov

Valid

Huang et al.

Late Oligocene

Yongning Formation

China

A species of Castanopsis.

Castanopsis nanningensis[15]

Sp. nov

Valid

Huang et al.

Late Oligocene

Yongning Formation

China

A species of Castanopsis.

Chenocybus[16]

Gen. et sp. nov

Valid

Poinar

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A flowering plant of uncertain phylogenetic placement. Genus includes new species C. allodapus.

Chisochetonoxylon vastanensis[17]

Sp. nov

Valid

Shukla & Mehrota

Early Eocene

Cambay Shale Formation

India

A member of the family Meliaceae described on the basis of fossil wood.

Cladium transdnestrovicum[18]

Nom. nov

Valid

Doweld

Miocene (Serravallian)

|  |

Transnistria

A species of Cladium; a replacement name for Cladium crassum Negru (1972), preoccupied by extant C. crassum (Thwaites) Kükenthal.

Clerodendrum sarmatiacum[3]

Nom. nov

Valid

Doweld

Miocene

|  |

Russia
( Rostov Oblast)

A species of Clerodendrum; a replacement name for Clerodendrum ovalifolium Baikovskaja in Kryshtofovich & Baikovskaja (1965).

Cobbania pharao[19]

Sp. nov

Valid

Coiffard & Mohr

Late Cretaceous (Campanian)

Quseir Formation

Egypt

A member of the family Araceae belonging or related to the subfamily Aroideae.

Concavistylon[20]

Gen. et 2 sp. nov

Valid

Manchester, Pigg & Devore

Early Eocene to Middle Miocene

Little Butte Volcanic Series

United States( Oregon)

A Trochodendraceae genus. Type species C. kvacekii Manchester, Pigg & Devore (2018) from Oregon
C. wehrii Manchester et al. (2018) from Washington state and British Columbia was originally described as a second species of this genus,[21] but subsequently it was transferred to the separate genus Paraconcavistylon.[22]

Craspedodromophyllum boguchanicum[7]

Sp. nov

Valid

Moiseeva, Kodrul & Herman

Paleocene

Zeya–Bureya Basin

Russia

A member of the family Betulaceae.

Cretaceoxylon[23]

Gen. et sp. nov

Valid

Pujana in Pujana et al.

Late Cretaceous (Campanian)

Santa Marta Formation

Antarctica
(James Ross Island)

A eudicot of uncertain phylogenetic placement, described on the basis of fossil wood. Genus includes new species C. heteropunctatum.

Cryptocaryoxylon lemnium[24]

Sp. nov

Valid

Mantzouka

Early Miocene

|  |

Greece

A member of the family Lauraceae.

Cryptocaryoxylon lesbium[24]

Sp. nov

Valid

Mantzouka

Early Miocene

|  |

Greece

A member of the family Lauraceae.

Cussoniophyllum[25]

Nom. nov

Valid

Doweld

Late Cretaceous (Cenomanian)

|  |

Czech Republic

A flowering plant described on the basis of fossil leaves; a replacement name for the invalidly published Cussoniphyllum Velenovský (1889). Genus includes "Cussonia" partita Velenovský (1882).

Cyperus maii[18]

Nom. nov

Valid

Doweld

Miocene

|  |

Germany

A species of Cyperus; a replacement name for Dichostylis macrocarpa Mai (1987).

Cyperus waltheri[18]

Nom. nov

Valid

Doweld

Miocene

|  |

Germany

A species of Cyperus; a replacement name for Dichostylis minor Mai in Mai & Walther (1991).

Dakotanthus[26]

Gen. et comb. nov

Valid

Manchester et al.

Cretaceous (late Albian to Cenomanian)

Dakota Formation
Woodbine Formation

United States
( Kansas
Nebraska
Texas)

An early eudicot; a new genus for "Carpites" cordiformis Lesquereux (1892).

Dalbergioxylon biseriatensis[27]

Sp. nov

Valid

Cheng et al.

Pliocene

Yuanmou Basin

China

A member of the family Fabaceae described on the basis of fossil wood.

Diaphoranthus[16]

Gen. et sp. nov

Junior homonym

Poinar

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A flowering plant of uncertain phylogenetic placement. Genus includes new species D. burmensis. The generic name is preoccupied by Diaphoranthus Meyen (1834); Poinar (2019) coined a replacement name Exalloanthum.[28]

Dicotylophyllum skogii[4]

Sp. nov

Valid

Wang & Dilcher

Early Cretaceous (Albian)

Dakota Formation

United States
( Kansas)

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil leaves.

Dioscorites palauensis[29]

Sp. nov

Valid

Guzmán-Vázquez, Calvillo-Canadell & Sánchez-Beristain

Late Cretaceous

Olmos Formation

Mexico

A member of the family Dioscoreaceae.

Diplosophyllum[25]

Nom. nov

Valid

Doweld

Late Cretaceous (Cenomanian)

|  |

Czech Republic
Germany

A flowering plant described on the basis of fossil leaves; a replacement name for the preoccupied Diplophyllum Velenovský & Viniklář (1929). Genus includes "Inga" cottae Ettingshausen (1867), "Diplophyllum" cretaceum Velenovský & Viniklář (1929), "Hymenaea" elongata Velenovský (1884), "Hymenaea" inaequalis Velenovský (1884) and "Hymenaea" primigenia de Saporta in Velenovský (1884).

Dipterocarpuspollenites cretacea[30]

Sp. nov

Valid

Prasad et al.

Late Cretaceous (Maastrichtian)

|  |

India

A pollen taxon belonging to the family Dipterocarpaceae.

Donlesia cheyennensis[31]

Sp. nov

Valid

Wang & Dilcher

Early Cretaceous (Albian)

Cheyenne Sandstone

United States
( Kansas)

A member of the family Ceratophyllaceae.

Ebenoxylon cuddalorensis[9]

Sp. nov

Valid

Awasthi, Mehrotra & Shukla

Late Miocene–early Pliocene

Cuddalore Sandstone Formation

India

A fossil wood showing affinities with members of the family Ebenaceae.

Edencarpa[32]

Gen. et sp. nov

Valid

Atkinson, Stockey & Rothwell

Late Cretaceous (early Coniacian)

|  |

Canada
( British Columbia)

A member of Cornales. Genus includes new species E. grandis.

Endobeuthos[33]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A flowering plant of uncertain phylogenetic placement. Originally described as a possible relative of members of the family Dilleniaceae; Chambers & Poinar (2023) subsequently reinterpreted it as a member of the family Proteaceae,[34] but this interpretation was rejected by Lamont & Ladd (2024).[35] Genus includes new species E. paleosum.

Eucalyptoxylon cuddalorensis[9]

Sp. nov

Valid

Awasthi, Mehrotra & Shukla

Late Miocene–early Pliocene

Cuddalore Sandstone Formation

India

A fossil wood showing affinities with members of the genus Eucalyptus.

Euphorbia pontiana[3]

Nom. nov

Valid

Doweld

Miocene

|  |

Ukraine

A species of Euphorbia; a replacement name for Euphorbia cylindrica Negru (1979).

Eydeia vancouverensis[32]

Sp. nov

Valid

Atkinson, Stockey & Rothwell

Late Cretaceous (early Coniacian)

|  |

Canada
( British Columbia)

A member of Cornales.

Ficophyllum angustifolium[36]

Nom. nov

Valid

Doweld

Late Cretaceous (Campanian)

|  |

Germany

A replacement name for Ficus angustifolia Hosius (1869).

Ficophyllum antiquum[36]

Nom. nov

Valid

Doweld

Late Cretaceous (Campanian)

|  |

Germany

A replacement name for Ficus crassinervis Hosius (1869).

Ficophyllum hosii[36]

Nom. nov

Valid

Doweld

Late Cretaceous (Santonian)

|  |

Germany

A replacement name for Ficus laurifolia Hosius & Marck (1880).

Ficophyllum magnolioides[36]

Nom. nov

Valid

Doweld

Early Cretaceous (Albian)

Dakota Formation

United States
( Kansas)

A replacement name for Ficus magnoliifolia Lesquereux (1883).

Ficophyllum marckii[36]

Nom. nov

Valid

Doweld

Late Cretaceous (Campanian)

|  |

Germany

A replacement name for Ficus elongata Hosius (1869).

Ficus aenigmatica[36]

Nom. nov

Valid

Doweld

Eocene

Wilcox Formation

United States
( Mississippi)

A species of Ficus; a replacement name for Ficus schimperi Lesquereux (1868).

Ficus microtrivia[37]

Sp. nov

Valid

Huang & Zhou in Huang et al.

Miocene

Wenshan Basin

China

A species of Ficus.

Ficus myrtoides[36]

Nom. nov

Valid

Doweld

Eocene

|  |

United States
( Mississippi)

A species of Ficus; a replacement name for Ficus myrtifolius Berry (1916).

Ficus slovenica[36]

Nom. nov

Valid

Doweld

Eocene

|  |

Slovenia

A species of Ficus; a replacement name for Ficus pilosa Ettingshausen (1872).

Ficus venustoides[36]

Nom. nov

Valid

Doweld

Oligocene (Chattian)

|  |

France

A species of Ficus; a replacement name for Ficus venusta Saporta (1861).

Ficus venustula[36]

Nom. nov

Valid

Doweld

Eocene

|  |

Croatia

A species of Ficus; a replacement name for Malpighiastrum venustum Unger (1860).

Ficus yellowstonica[36]

Nom. nov

Valid

Doweld

Paleocene

|  |

United States
( Wyoming)

A species of Ficus; a replacement name for Ficus densifolia Knowlton (1899).

Fissistigma nanningense[38]

Sp. nov

Valid

Li et al.

Oligocene

Yongning Formation

China

A species of Fissistigma.

Gardenia eocenicus[39]

Sp. nov

Valid

Shukla, Mehrotra & Nawaz Ali

Early Eocene

Palana Formation

India

A species of Gardenia.

Gastonispermum[40]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

|  |

Portugal

A flowering plant with affinities to Austrobaileyales or Nymphaeales. Genus includes new species G. portugallicum.

Gleditsioxylon jiangsuensis[41]

Sp. nov

Valid

Cheng et al.

Early Miocene

|  |

China

A member of Leguminosae described on the basis of fossil wood.

Gmelina siwalika[42]

Sp. nov

Valid

Khan, Bera & Bera in Khan et al.

Late Pliocene or early Pleistocene

Kimin Formation

India

A species of Gmelina.

Goniothalamus miocenicus[5]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Formation

Nepal

A species of Goniothalamus.

Gouania miocenica[43]

Sp. nov

Valid

Hernandez-Hernández & Castañeda-Posadas

Early Miocene

Mexican amber

Mexico

A species of Gouania.

Hederago[25]

Nom. nov

Valid

Doweld

Late Cretaceous (Cenomanian)

|  |

Czech Republic

A flowering plant described on the basis of fossil leaves; a replacement name for the invalidly published Hederophyllum Velenovský (1889). Genus includes "Hedera" credneriifolia Velenovský (1882) and "Hedera" primordialis de Saporta (1879).

Hemitrapa alpina[44]

Sp. nov

Valid

Su & Zhou in Su et al.

Early Oligocene

|  |

China

A member of the family Lythraceae. Originally described as a species of Hemitrapa, but subsequently transferred to the genus Primotrapa by Li et al. (2020).[45]

Hibiscus sarmatiacus[3]

Sp. nov

Valid

Doweld

Miocene

|  |

Russia
( Rostov Oblast)

A species of Hibiscus; a replacement name for the invalidly named Hibiscus splendens Baikovskaja.

Holigarna palaeograhamii[39]

Sp. nov

Valid

Shukla, Mehrotra & Nawaz Ali

Early Eocene

Palana Formation

India

A species of Holigarna.

Hopenium tertiarum[9]

Sp. nov

Valid

Awasthi, Mehrotra & Shukla

Late Miocene–early Pliocene

Cuddalore Sandstone Formation

India

A fossil wood showing affinities with members of the genus Hopea.

Ipomoea meghalayensis[46]

Sp. nov

Valid

Srivastava, Mehrotra & Dilcher

Paleocene (Thanetian)

|  |

India

A species of Ipomoea.

Kirchheimeria[47]

Gen. et comb. nov

Valid

Kowalski in Kowalski & Worobiec

Oligocene to Pliocene

|  |

Denmark
Germany
Poland
Russia
( Kaliningrad Oblast)

A member of Ericaceae of uncertain phylogenetic placement. Genus includes "Elaeocarpus" globulus Menzel (1906).

Kvacekispermum[48]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

Figueira da Foz Formation

Portugal

A member of the family Chloranthaceae. Genus includes new species K. rugosum.

Lachnociona camptostylus[49]

Sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A flowering plant of uncertain phylogenetic placement, most similar to members of the families Brunelliaceae and Cunoniaceae.

Lacinipetalum[50]

Gen. et sp. nov

Valid

Jud et al.

Paleocene (early Danian)

Upper Salamanca Formation

Argentina

A member of Cunoniaceae. Genus includes new species L. spectabilum.

Laurinoxylon rennerae[51]

Sp. nov

Valid

Estrada-Ruiz et al.

Late Cretaceous (late Campanian)

McRae Formation

United States
( New Mexico)

A member of Lauraceae described on the basis of fossil wood.

Laurus ficoides[36]

Nom. nov

Valid

Doweld

Eocene

|  |

France

A species of Laurus; a replacement name for Ficus reticulata Saporta (1863).

Lefipania[52]

Gen. et sp. nov

Valid

Martínez, Gandolfo & Cúneo

Late Cretaceous (Maastrichtian)

Lefipán Formation

Argentina

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil leaves. Genus includes new species L. padillae.

Leguminocarpum oguruiensis[53]

Sp. nov

Valid

Yabe & Nakagawa

Miocene

Shimo Formation

Japan

A fossil legume fruit.

Ligustrum miovulgare[3]

Sp. nov

Valid

Doweld

Miocene

|  |

Russia
( Rostov Oblast)

A species of Ligustrum; a replacement name for the invalidly named Ligustrum vulgare var. fossilis Baikovskaja.

Lijinganthus[54]

Gen. et sp. nov

Valid

Liu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Pentapetalae of uncertain phylogenetic placement. Genus includes new species L. revoluta.

Limnobiophyllum stockeyana[19]

Sp. nov

Valid

Coiffard & Mohr

Late Cretaceous (Campanian)

Quseir Formation

Egypt

A member of the family Araceae belonging to the subfamily Lemnoideae.

Liquidambar fujianensis[55]

Sp. nov

Valid

Dong et al.

Middle Miocene

Fotan Group

China

A species of Liquidambar.

Lithocarpoxylon microporosum[27]

Sp. nov

Valid

Cheng et al.

Pliocene

Yuanmou Basin

China

A member of the family Fagaceae described on the basis of fossil wood.

Lithocarpoxylon nanningensis[15]

Sp. nov

Valid

Huang et al.

Late Oligocene

Yongning Formation

China

A member of Fagaceae described on the basis of fossil wood.

Litseoxylon[56]

Gen. et sp. nov

Valid

Huang et al.

Late Oligocene

Yongning Formation

China

A member of the family Lauraceae. Genus includes new species L. nanningensis.

Luckowcarpa[57]

Gen. et sp. nov

Valid

Martínez

Late Eocene

Esmeraldas Formation

Colombia

A member of Fabaceae belonging to the group Dalbergieae. Genus includes new species L. gunnii.

Lusitanispermum[40]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

|  |

Portugal

A flowering plant with affinities to Austrobaileyales or Nymphaeales. Genus includes new species L. choffatii.

Lycopus europleistocenicus[3]

Sp. nov

Valid

Doweld

Pleistocene

|  |

Belarus

A species of Lycopus; a replacement name for the invalidly named Lycopus intermedius Dorofeev (1963).

Malus antiqua[3]

Nom. nov

Valid

Doweld

Miocene

|  |

Romania

A species of Malus; a replacement name for Malus pulcherrima Givulescu (1980).

Maytenoxylon[58]

Gen. et sp. nov

Valid

Franco

Late Cenozoic

Ituzaingó Formation

Argentina

A member of Celastraceae described on the basis of fossil wood. Genus includes new species M. perforatum.

Mcraeoxylon[51]

Gen. et sp. nov

Valid

Estrada-Ruiz et al.

Late Cretaceous (late Campanian)

McRae Formation

United States
( New Mexico)

A flowering plant described on the basis of fossil wood, with a suite of features seen in several families of Malpighiales, Myrtales and Oxalidales. Genus includes new species M. waddellii.

Meliosma antiqua[3]

Nom. nov

Valid

Doweld

Oligocene

|  |

United Kingdom

A species of Meliosma; a replacement name for Calvarinus reticulatus Reid & Reid (1910).

Menispermites calderensis[59]

Sp. nov

Valid

Jud et al.

Eocene (Ypresian)

Huitrera Formation

Argentina

A member of the family Menispermaceae described on the basis of fossil leaves.

Menispermites olmosensis[29]

Sp. nov

Valid

Guzmán-Vázquez, Calvillo-Canadell & Sánchez-Beristain

Late Cretaceous

Olmos Formation

Mexico

A member of the family Menispermaceae.

Nelumbo jiayinensis[60]

Sp. nov

Valid

Liang et al.

Late Cretaceous (Santonian)

Yong'ancun Formation

China

A species of Nelumbo.

Neofructus[61]

Gen. et sp. nov

Valid

Liu & Wang

Early Cretaceous (Barremian–Aptian)

Yixian Formation

China

An early flowering plant. Genus includes new species N. lingyuanensis.

Nitaspermum[62]

Gen. et 5 sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

United States
( Maryland
Virginia)

A fossil seed with affinities to Austrobaileyales and Nymphaeales. Genus includes new species N. taylorii, N. hopewellense, N. crassum, N. virginiense and N. marylandense.

Nyssa givulescui[3]

Nom. nov

Valid

Doweld

Oligocene

|  |

Romania

A tupelo; a replacement name for Nyssa maxima Givulescu, Petrescu & Barbu (1997).

Obamacarpa[32]

Gen. et sp. nov

Valid

Atkinson, Stockey & Rothwell

Late Cretaceous (early Coniacian)

|  |

Canada
( British Columbia)

A member of Cornales. Genus includes new species O. edenensis.

Ocotea undulatoides[3]

Nom. nov

Valid

Doweld

Miocene

|  |

Germany

A species of Ocotea; a replacement name for Laurophyllum undulatum Weyland & Kilpper (1963).

Paisia[63]

Gen. et sp. nov

Valid

Friis, Mendes & Pedersen

Early Cretaceous (late Barremian–early Albian)

Almargem Formation

Portugal

An early eudicot. Genus includes new species P. pantoporata.

Palaeocarya huashanensis[64]

Sp. nov

Valid

Chen et al.

Oligocene

Ningming Formation

China

A member of the family Juglandaceae.

Paleoallium[65]

Gen. et sp. nov

Valid

Pigg, Bryan & DeVore

Eocene
Ypresian

Okanagan Highlands
Klondike Mountain Formation

United States
Washington

A monocot similar to members of Amaryllidaceae. Genus includes new species P. billgenseli.

|  Paleoallium billgenseli

Paliurus hirsuta[66]

Sp. nov

Valid

Dong & Sun in Dong et al.

Middle Miocene

Fotan Group

China

A species of Paliurus.

Palmoxylon araneus[67]

Sp. nov

Valid

Nour-El-Deen, El-Saadawi & Thomas

Oligocene (Rupelian)

Jebel Qatrani Formation

Egypt

|  |

Palmoxylon elsaadawii[67]

Sp. nov

Valid

Nour-El-Deen & Thomas in Nour-El-Deen, Thomas & El-Saadawi

Oligocene (Rupelian)

Jebel Qatrani Formation

Egypt

|  |

Palmoxylon qatraniense[67]

Sp. nov

Valid

Nour-El-Deen, El-Saadawi & Thomas

Oligocene (Rupelian)

Jebel Qatrani Formation

Egypt

|  |

Paraalbizioxylon sinica[27]

Sp. nov

Valid

Cheng et al.

Pliocene

Yuanmou Basin

China

A member of the family Fabaceae described on the basis of fossil wood.

Paraalbizioxylon yunnanensis[27]

Sp. nov

Valid

Cheng et al.

Pliocene

Yuanmou Basin

China

A member of the family Fabaceae described on the basis of fossil wood.

Parahancornioxylon[68]

Gen. et comb. nov

Valid

Moya, Brea & Lutz

Pliocene

Andalhualá Formation

Argentina

A member of Apocynaceae described on the basis of fossil wood; a new genus for "Menendoxylon" piptadiensis Lutz (1987).

Paraphyllanthoxylon antarcticum[23]

Sp. nov

Valid

Pujana in Pujana et al.

Late Cretaceous (Campanian)

Santa Marta Formation

Antarctica
(James Ross Island)

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil wood.

Pazlia[40]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

|  |

Portugal

A flowering plant with affinities to Austrobaileyales or Nymphaeales. Genus includes new species P. hilaris.

Pazliopsis[40]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

Almargem Formation

Portugal

A flowering plant with affinities to Austrobaileyales or Nymphaeales. Genus includes new species P. reyi.

Pentacentron[21]

Gen. et sp. nov

Valid

Manchester et al.

Eocene
Ypresian

Okanagan Highlands
Klondike Mountain Formation

United States
Washington

A member of the family Trochodendraceae. The type species is P. sternhartae.

|  Pentacentron sternhartae

Photinia sarmatiaca[3]

Sp. nov

Valid

Doweld

Miocene

|  |

Russia
( Rostov Oblast)

A species of Photinia; a replacement name for the invalidly named Photinia acuminata Baikovskaja in Kryshtofovich & Baikovskaja (1965).

Pistacia miolentiscus[3]

Nom. nov

Valid

Doweld

Miocene

|  |

Hungary

A species of Pistacia; a replacement name for Pistacia lentiscoides Andreánszky & Cziffery in Andreánszky (1959).

Pistacia pliolentiscus[3]

Nom. nov

Valid

Doweld

Pliocene

|  |

Netherlands

A species of Pistacia; a replacement name for Pistacia acuminata Reid & Reid (1915).

Pistacioxylon ufuki[69]

Sp. nov

Valid

Akkemik & Poole in Akkemik et al.

Early Miocene

Haymana Basin

Turkey

A Pistacia-like plant described on the basis of fossil wood.

Polyalthioxylon arunachalensis[70]

Sp. nov

Valid

Srivastava, Mehrotra & Srikarni

Late Pliocene–Early Pleistocene

Kimin Formation

India

A member of the family Annonaceae described on the basis of fossil wood.

Priscophyllum[25]

Nom. nov

Valid

Doweld

Late Cretaceous (Cenomanian)

|  |

Czech Republic

A flowering plant described on the basis of fossil leaves; a replacement name for the invalidly published Grevilleophyllum Velenovský (1889). Genus includes "Grevillea" constans Velenovský (1883).

Prunus hirsutipetala[71]

Sp. nov

Valid

Sokoloff, Remizowa & Nuraliev in Sokoloff et al.

Eocene

Rovno amber

Ukraine

A species of Prunus.

Pseudoanacardium[72]

Gen. et comb. nov

Valid

Manchester & Balmaki

Early Oligocene

|  |

Peru

A fossil fruit of uncertain phylogenetic placement; a new genus for "Anacardium" peruvianum Berry (1924).

Pseudolimnobiophyllum[19]

Gen. et sp. nov

Valid

Coiffard & Mohr

Late Cretaceous (Campanian)

Quseir Formation

Egypt

A member of the family Araceae belonging to the subfamily Lemnoideae. Genus includes new species P. simile.

Pseudowinterapollis agatdalensis[73]

Sp. nov

Valid

Grímsson & Zetter in Grímsson et al.

Paleocene (Danian)

Agatdal Formation

Greenland

A pollen taxon, a member of the family Winteraceae.

Pterocaryoxylon huxii[27]

Sp. nov

Valid

Cheng et al.

Pliocene

Yuanmou Basin

China

A member of the family Juglandaceae described on the basis of fossil wood.

Pterygota eocenica[39]

Sp. nov

Valid

Shukla, Mehrotra & Nawaz Ali

Early Eocene

Palana Formation

India

A species of Pterygota.

Ranunculus eoreptans[3]

Nom. nov

Valid

Doweld

Pliocene

|  |

Belarus

A species of Ranunculus; a replacement name for Ranunculus pusillus Dorofeev (1987).

Retiacolpites pigafettaensis[30]

Sp. nov

Valid

Prasad et al.

Late Cretaceous (Maastrichtian)

|  |

India

A pollen taxon resembling pollen of members of the genus Pigafetta.

Reyispermum[40]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

Figueira da Foz Formation

Portugal

A flowering plant with affinities to Austrobaileyales or Nymphaeales. Genus includes new species R. parvum.

Rhododendron maii[3]

Nom. nov

Valid

Doweld

Pliocene

|  |

Germany

A species of Rhododendron; a replacement name for Rhododendron germanicum Mai & Walther (1988).

Rightcania[48]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

United States
( Virginia)

A member of the family Chloranthaceae. Genus includes new species R. kvacekii.

Ripogonum palaeozeylandiae[74]

Sp. nov

Valid

Conran, Kennedy & Bannister

Early Eocene

|  |

New Zealand

A species of Ripogonum.

Ruprechtioxylon breae[58]

Sp. nov

Valid

Franco

Late Cenozoic

Ituzaingó Formation

Argentina

A member of Polygonaceae described on the basis of fossil wood.

Ryparosa churiaensis[5]

Sp. nov

Valid

Prasad et al.

Miocene

Churia Formation

Nepal

A species of Ryparosa.

Salacia lombardii[75]

Sp. nov

Valid

Hernández-Damián, Gómez-Acevedo & Cevallos-Ferriz

Miocene

|  |

Mexico

A species of Salacia.

Sambucus sarmatiaca[3]

Sp. nov

Valid

Doweld

Miocene

|  |

Russia
( Rostov Oblast)

A species of Sambucus; a replacement name for the invalidly named Sambucus palaeoracemosa Baikovskaja in Kryshtofovich & Baikovskaja (1965).

Sapindopsis retallackii[4]

Sp. nov

Valid

Wang & Dilcher

Early Cretaceous (Albian)

Dakota Formation

United States
( Kansas)

A member or a relative of the family Platanaceae described on the basis of fossil leaves.

Schoenoplectiella isolepioides[18]

Sp. nov

Valid

Doweld

Pliocene

|  |

Germany

A member of the family Cyperaceae; a replacement name for the invalidly named Scirpus (Schoenoplectus) isolepioides Mai & Walther (1988).

Scirpus novorossicus[18]

Nom. nov

Valid

Doweld

Miocene (Tortonian)

|  |

Ukraine

A species of Scirpus; a replacement name for Scirpus leptocarpus Negru (1986), preoccupied by extant Scirpus leptocarpus Mueller (1855).

Setitheca[76]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Laurales of uncertain phylogenetic placement. Genus includes new species S. lativalva.

Silutanispermum[40]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

|  |

Portugal

A flowering plant with affinities to Austrobaileyales or Nymphaeales. Genus includes new species S. kvacekiorum.

Sloanea siwalika[77]

Sp. nov

Valid

More et al.

Pliocene

Geabdat Sandstone Formation

India

A species of Sloanea.

Soepadmoa[78]

Gen. et sp. nov

Valid

Nixon, Crepet, Gandolfo & Grimaldi

Late Cretaceous (Turonian)

Raritan Formation
(New Jersey amber)

United States
( New Jersey)

A member of Fagales of uncertain phylogenetic placement. Genus includes new species S. cupulata.

Staphylea spinosa[79]

Sp. nov

Valid

Huang & Momohara in Huang, Momohara & Wang

Pleistocene

Shobudani Formation

Japan

A species of Staphylea.

Stafylioxylon[8]

Gen. et comb. nov

Valid

Rozefelds & Pace

Eocene

London Clay

United Kingdom

A member of Vitaceae; a new genus for "Vitaceoxylon" ramunculiformis Poole & Wilkinson (2000).

Stellatia[2]

Gen. et comb. nov

Valid

Arai & Dias-Brito

Late Cretaceous (Santonian)

São Carlos Formation

Brazil

A phytoclast, possibly a member of Nymphaeaceae. Genus includes S. furcata (Duarte & Arai, 2010).

Stephania auriformis[80]

Comb nov

valid

(Hollick) Manchester & Han

Paleocene/Eocene

"King Salmon Lake flora"

USA
Alaska

A moonseed species.
Moved from Diploclisia auriformis (1994)[81]

Stephania jacquesii[80]

Sp. nov

Valid

Han & Manchester in Han et al.

Late Eocene to late Oligocene

Clarno Formation
Yongning Formation

China
United States
( Oregon)

A species of Stephania.

Stephania psittaca[59]

Sp. nov

Valid

Jud & Gandolfo in Jud et al.

Paleocene (Danian)

Salamanca Formation

Argentina

A species of Stephania.

Stephania wilfii[80]

Sp. nov

Valid

Han & Manchester in Han et al.

Paleocene to Eocene

Green River Formation

United States
( Wyoming)

A species of Stephania.

Sterculia acerina[3]

Nom. nov

Valid

Doweld

Eocene

|  |

Czech Republic

A species of Sterculia; a replacement name for Acer crassinervium Ettingshausen (1869).

Symplocos hitchcockii[82]

Sp. nov

Valid

Tiffney, Manchester & Fritsch

Early Miocene

Brandon Lignite

United States
( Vermont)

A species of Symplocos.

Syzygium christophelii[83]

Sp. nov

Valid

Tarran et al.

Early Miocene

|  |

Australia

A species of Syzygium.

Syzygium gurhaensis[39]

Sp. nov

Valid

Shukla, Mehrotra & Nawaz Ali

Early Eocene

Palana Formation

India

A species of Syzygium.

Tanispermum[84]

Gen. et 4 sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (early Aptian to early to middle Albian)

Potomac Group

United States
( Maryland
Virginia)

A flowering plant with affinities to Austrobaileyales or Nymphaeales. Genus includes new species T. hopewellense, T. marylandense, T. drewriense and T. antiquum.

Teuschestanthes[85]

Gen. et sp. nov

Valid

Crepet, Nixon & Weeks

Late Cretaceous (Turonian)

Lower Magothy Formation

United States
( New Jersey)

A member of Ericales of uncertain phylogenetic placement. Genus includes new species T. squamata.

Trichomites[2]

Gen. et 3 sp. nov

Valid

Arai & Dias-Brito

Late Cretaceous (Santonian)

São Carlos Formation

Brazil

A phytoclast. Genus includes new species T. brevifurcatus (probably a member of Campanulaceae), T. duplihelicoidus (affinity unknown) and T. simplex (a dicotyledon of uncertain affinity).

Tricolpites joelcastroi[2]

Sp. nov

Valid

Arai & Dias-Brito

Late Cretaceous (Santonian)

São Carlos Formation

Brazil

A pollen taxon, an indeterminate dicotyledon.

Trochodendroides sittensis[86]

Sp. nov

Valid

Golovneva in Golovneva & Zolina

Early Cretaceous

|  |

Russia

Taxon described on the basis of fossil leaves resembling leaves of members of the family Cercidiphyllaceae.

Trochodendron postnastae[20]

Sp. nov

Valid

Manchester, Pigg & Devore

Middle Miocene

Little Butte Volcanic Series

United States
( Oregon)

A species of Trochodendron.

Trochodendron rosayi[20]

Sp. nov

Valid

Manchester, Pigg & Devore

Middle Miocene

Little Butte Volcanic Series

United States
( Idaho
Oregon)

A species of Trochodendron.

Turneroxylon[51]

Gen. et sp. nov

Valid

Estrada-Ruiz et al.

Late Cretaceous (late Campanian)

McRae Formation

United States
( New Mexico)

A eudicot with similarities to members of Dilleniaceae, described on the basis of fossil wood. Genus includes new species T. newmexicoense.

Ulmus maguanensis[87]

Sp. nov

Valid

Zhang & Xing in Zhang et al.

Miocene

Huazhige Formation

China

An elm.

Ulmus prelanceaefolia[87]

Sp. nov

Valid

Zhang & Xing in Zhang et al.

Miocene

Huazhige Formation

China

An elm.

Ulmus priamurica[88]

Sp. nov

Valid

Blokhina & Bondarenko

Miocene

Sazanka Formation

Russia
( Amur Oblast)

An elm.

Unona miocenica[5]

Sp. nov

Valid

Prasad et al.

Miocene

Churia Formation

Nepal

A member of the family Annonaceae.

Viburnum pliolantana[3]

Nom. nov

Valid

Doweld

Pliocene

|  |

Russia
( Bashkortostan)

A species of Viburnum; a replacement name for Viburnum lantanoides Dorofeev (1977).

Weinmannioxylon trichospermoides[23]

Sp. nov

Valid

Pujana in Pujana et al.

Late Cretaceous (Campanian)

Santa Marta Formation

Antarctica
(James Ross Island)

A member of Cunoniaceae described on the basis of fossil wood.

Wilkinsoniphyllum[59]

Gen. et sp. nov

Valid

Jud et al.

Paleocene (Danian)

Salamanca Formation

Argentina

A member of the family Menispermaceae described on the basis of fossil leaves. Genus includes new species W. menispermoides.

Wingia[4]

Gen. et comb. nov

Valid

Wang & Dilcher

Early Cretaceous (Albian)

Dakota Formation

United States
( Kansas
Nebraska)

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil leaves. Genus includes "Dicotylophyllum" expansolobum Upchurch & Dilcher (1990).

Zanthoxylum pilari[3]

Nom. nov

Valid

Doweld

Miocene

|  |

Croatia

A species of Zanthoxylum; a replacement name for Zanthoxylum affine Pilar (1883).

Zanthoxylum tethyca[3]

Nom. nov

Valid

Doweld

Eocene

|  |

United Kingdom

A species of Zanthoxylum; a replacement name for Rutaspermum rugosum Chandler (1964).

Zelkovoxylon yesimae[69]

Sp. nov

Valid

Akkemik & Poole in Akkemik et al.

Early Miocene

Haymana Basin

Turkey

A Zelkova-like plant described on the basis of fossil wood.

Zygogynum poratus[89]

Sp. nov

Valid

Liang & Zhou in Liang et al.

Middle Miocene

|  |

China

A species of Zygogynum.

## Pinales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Agathis immortalis[90]

Sp. nov

Valid

Escapa et al.

Paleocene (Danian)

|  |

Argentina

A species of Agathis.

Agathoxylon crasseradiatum[91]

Sp. nov

Valid

Lignier ex Philippe et al.

Early Cretaceous (late Aptian-Albian)

|  |

France

A member of Araucariaceae described on the basis of fossil wood.

Agathoxylon holbavicum[92]

Sp. nov

Valid

Iamandei, Iamandei & Grădinaru

Early Jurassic

|  |

Romania

|  |

Agathoxylon santacruzense[93]

Sp. nov

Valid

Kloster & Gnaedinger

Middle Jurassic

La Matilde Formation

Argentina

|  |

Araucaria lefipanensis[94]

Sp. nov

Valid

Andruchow-Colombo et al.

Late Cretaceous

Lefipán Formation

Argentina

A species of Araucaria.

Atlanticoxylon ibiratinum[95]

Sp. nov

Valid

Faria et al.

Permian (Artinskian)

Irati Formation

Brazil

A conifer described on the basis of fossil wood.

Brachyoxylon cristianicum[92]

Sp. nov

Valid

Iamandei, Iamandei & Grădinaru

Early Jurassic

|  |

Romania

|  |

Brachyoxylon holbavicum[92]

Sp. nov

Valid

Iamandei, Iamandei & Grădinaru

Early Jurassic

|  |

Romania

|  |

Brachyoxylon zhejiangense[96]

Sp. nov

Valid

Tian, Zhu & Wang in Tian et al.

Early Cretaceous

Guantou Formation

China

A coniferous wood.

Chimaerostrobus[97]

Gen. et sp. nov

Valid

Atkinson et al.

Early Jurassic(Pliensbachian-Toarcian)

Mawson Formation

Antarctica

A conifer pollen cone. Genus includes new species C. minutus.

Cryptomeria yunnanensis[98]

Sp. nov

Valid

Ding & Zhou in Ding et al.

Oligocene (Rupelian)

Lühe Basin

China

A member of Cupressaceae, a species of Cryptomeria.

Cunninghamia shangcunica[99]

Sp. nov

Valid

Kodrul et al.

Early Oligocene

Shangcun Formation

China

A species of Cunninghamia.

Cyclusphaera annularis[100]

Sp. nov

Valid

Perez Loinaze & Llorens

Early Cretaceous (Aptian)

Anfiteatro de Ticó Formation

Argentina

A pollen taxon with affinities with the family Araucariaceae.

Cyclusphaera punnulosa[100]

Sp. nov

Valid

Perez Loinaze & Llorens

Early Cretaceous (Aptian)

Anfiteatro de Ticó Formation

Argentina

A pollen taxon with affinities with the family Araucariaceae.

Elatides laiyangensis[101]

Sp. nov

Valid

Jin & Sun in Jin et al.

Early Cretaceous

Laiyang Formation

China

A conifer.

Hirandubia[102]

Gen. et sp. nov

Valid

Ghosh et al.

Early Cretaceous

Rajmahal Basin

India

A member of Cupressaceae. Genus includes new species H. cupressoides.

Kirketapel salamanquensis[103]

Sp. nov

Valid

Andruchow-Colombo et al.

Paleocene (Danian)

Salamanca Formation

Argentina

The oldest member of a scale-leaved clade of Podocarpaceae.

Marskea heeriana[104]

Sp. nov

Valid

Nosova & Kiritchkova

Middle Jurassic

Irkutsk Coal Basin

Russia

|  |

Morinostrobus[105]

Gen. et sp. nov

Valid

Stockey et al.

Early Cretaceous (Valanginian)

|  |

Canada
( British Columbia)

A member of Cupressaceae described on the basis of pollen cones. Genus includes new species M. holbergensis.

Pinus daflaensis[106]

Nom. nov

Valid

Khan & Bera

Miocene

Dafla Formation

India

A pine; a replacement name for Pinus arunachalensis Khan & Bera (2017) (preoccupied by Pinus arunachalensis Srivastava, 2017).

Pinus enochii[107]

Sp. nov

Valid

Huerta Vergara & Cevallos-Ferriz

Late Cretaceous (late Campanian)

Lutita Packard Formation

Mexico

A pine.

Pinus leiophylloides[108]

Nom. nov

Valid

Doweld

Oligocene (Chattian)

|  |

France

A pine; a replacement name for Pinus pseudotaeda Saporta (1865).

Pinus microstrobus[108]

Nom. nov

Valid

Doweld

Oligocene (Chattian)

|  |

France

A pine; a replacement name for Pinus microcarpa Saporta (1865).

Pinus notata[108]

Nom. nov

Valid

Doweld

Oligocene (Chattian)

|  |

France

A pine; a replacement name for Pinus divaricata Saporta (1865).

Pinus pentaphylloides[108]

Nom. nov

Valid

Doweld

Late Cretaceous (Santonian)

|  |

Japan

A pine; a replacement name for Pinus hokkaidoensis Stockey & Ueda (1986).

Pinus tetraphylloides[108]

Nom. nov

Valid

Doweld

Oligocene (Chattian)

|  |

France

A pine; a replacement name for Pinus deflexa Saporta (1865).

Pinus uxui[107]

Sp. nov

Valid

Huerta Vergara & Cevallos-Ferriz

Late Cretaceous (late Campanian)

Lutita Packard Formation

Mexico

A pine.

Platycladus preorientalis[109]

Sp. nov

Valid

He et al.

Early Miocene

|  |

China

A species of Platycladus.

Podocarpospermum podocarpoides[102]

Sp. nov

Valid

Ghosh et al.

Early Cretaceous

Rajmahal Basin

India

A member of Podocarpaceae.

Protocedroxylon zhalantunense[110]

Sp. nov

Valid

Zhang, Tian & Wang in Zhang et al.

Middle Jurassic

Wanbao Formation

China

A member of the family Pinaceae.

Protocedroxylon zhangii[110]

Sp. nov

Valid

Zhang, Tian & Wang in Zhang et al.

Middle Jurassic

Wanbao Formation

China

A member of the family Pinaceae.

Protophyllocladoxylon holbavicum[92]

Sp. nov

Valid

Iamandei, Iamandei & Grădinaru

Early Jurassic

|  |

Romania

|  |

Pseudofrenelopsis salesii[111]

Sp. nov

Valid

Batista et al.

Early Cretaceous (Albian)

Romualdo Member

Brazil

A member of Cheirolepidiaceae.

Rabagostrobus[112]

Gen. et sp. nov

Valid

Kvaček et al.

Early Cretaceous (Albian)

|  |

Spain

An araucarian pollen cone. Genus includes new species R. hispanicus.

Sequoioxylon carneyvillense[113]

Sp. nov

Valid

Li, Jin & Manchester

Paleocene

Fort Union Formation

United States
( Wyoming)

Fossil wood resembling Sequoia.

Sequoioxylon zhangii[114]

Sp. nov

Valid

Tian et al.

Late Cretaceous

|  |

China

A member of Sequoioideae described on the basis of fossil wood.

Taxocladus czeremchoviensis[115]

Sp. nov

Valid

Frolov & Mashchuk

Early Jurassic

Czeremkhovskaya Formation

Russia

Possibly a member of the family Taxaceae.

Yanliaoa daohugouensis[116]

Sp. nov

Valid

Tan et al.

Middle Jurassic

Daohugou Beds

China

A member of Cupressaceae sensu lato.

## Other seed plants

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Baiera telmensis[115]

Sp. nov

Valid

Frolov in Frolov & Mashchuk

Early Jurassic

Prisayanskaya Formation

Russia

A member of Ginkgoales.

Calycosperma[117]

Gen. et sp. nov

Valid

Liu et al.

Late Devonian

Wutong Formation

China

An early seed plant. Genus includes new species C. qii.

Carpolithes kurminensis[115]

Sp. nov

Valid

Frolov in Frolov & Mashchuk

Middle Jurassic

Taltsy Formation

Russia

Seed of a gymnosperm of uncertain affinities.

Cordaites daviessensis[118]

Sp. nov

Valid

Šimůnek

Carboniferous (early Westphalian D)

Staunton Formation

United States
( Indiana)

|  |

Cordaites kinneyensis[118]

Sp. nov

Valid

Šimůnek

Carboniferous (Stephanian B)

Atrasado Formation

United States
( New Mexico)

|  |

Cordaites minshallensis[118]

Sp. nov

Valid

Šimůnek

Carboniferous (Bolsovian)

Brazil Formation

United States
( Indiana)

|  |

Cordaites olneyensis[118]

Sp. nov

Valid

Šimůnek

Carboniferous (late Pennsylvanian)

Mattoon Formation

United States
( Illinois)

|  |

Cycadolepis ferrugineus[119]

Sp. nov

Valid

McLoughlin, Pott & Sobbe

Jurassic (Pliensbachian–Aalenian)

|  |

Australia

A member of Bennettitales belonging to the family Williamsoniaceae.

Cycadopites grossus[100]

Sp. nov

Valid

Perez Loinaze & Llorens

Early Cretaceous (Aptian)

Anfiteatro de Ticó Formation

Argentina

A pollen taxon, similar to many of the modern cycad pollen types.

Czekanowskia ottenii[120]

Sp. nov

Valid

Kiritchkova, Kostina & Nosova

Jurassic

|  |

Russia

|  |

Eamesia[121]

Gen. et sp. nov

Valid

Yang et al.

Early Cretaceous (Aptian)

Yixian Formation

China

A member of Ephedraceae. Genus includes new species E. chinensis.

Eretmophyllum neimengguensis[122]

Sp. nov

Valid

Li et al.

Middle Jurassic

Yan'an Formation

China

A member of Ginkgoales.

Eretmophyllum odintsovae[115]

Sp. nov

Valid

Frolov & Mashchuk

Middle Jurassic

Taltsy Formation

Russia

A member of Ginkgoales.

Eretmophyllum olchaense[120]

Sp. nov

Valid

Kiritchkova, Kostina & Nosova

Jurassic

|  |

Russia

|  |

Ginkgo cuneifolia[123]

Sp. nov

Valid

Tan, Dilcher, Wang & Sun in Sun et al.

Middle Jurassic

Jiulongshan Formation

China

A species of Ginkgo.

Ginkgo daohugouensis[123]

Sp. nov

Valid

Tan, Dilcher, Wang & Sun in Sun et al.

Middle Jurassic

Jiulongshan Formation

China

A species of Ginkgo.

Ginkgo glinkiensis[115]

Sp. nov

Valid

Frolov & Mashchuk

Early Jurassic

Czeremkhovskaya Formation

Russia

Originally described as a species of Ginkgo, but subsequently transferred to the genus Ginkgoites.[124]

Ginkgo parvifolia[123]

Sp. nov

Valid

Tan, Dilcher, Wang & Sun in Sun et al.

Middle Jurassic

Jiulongshan Formation

China

A species of Ginkgo.

Ginkgophyllum rhipidomorphum[125]

Sp. nov

Valid

Gomankov

Late Permian

|  |

Russia

|  |

Hexianthus[126]

Gen. et sp. nov

Valid

Wang & Sun in Wang et al.

Early Permian

Taiyuan Formation

China

A cone fossil belonging to the group Cordaitopsida and the family Cordaitaceae. Genus includes new species H. shenii.

Jugasporites vellicoites[127]

Sp. nov

Valid

Zavattieri, Gutiérrez & Ezpeleta

Permian (Lopingian)

La Veteada Formation

Argentina

A member of Voltziales described on the basis of fossil pollen grains.

Nanjinganthus[128]

Gen. et sp. nov

Valid

Fu et al.

Early Jurassic

South Xiangshan Formation

China

A seed plant of uncertain phylogenetic placement. Interpreted as an early fossil flower by Fu et al. (2018);[128] Coiro, Doyle & Hilton (2019) considered known specimens of this plant to be more similar to conifer cones.[129] Genus includes new species N. dendrostyla.

Nilssoniopteris crassiaxis[130]

Sp. nov

Valid

Zhao & Deng in Zhao et al.

Middle Jurassic

Xishanyao Formation

China

A member of Bennettitales.

Nilssoniopteris hamiensis[130]

Sp. nov

Valid

Zhao & Deng in Zhao et al.

Middle Jurassic

Xishanyao Formation

China

A member of Bennettitales.

Nilssoniopteris neimenguensis[131]

Nom. nov

Valid

Zhao & Deng in Zhao et al.

Early and Middle Jurassic

Hongqi Formation
Mentougou Formation

China

A member of Bennettitales; a replacement name for Nilssoniopteris angustifolia Wang (1984), preoccupied by Nilssoniopteris angustifolia Doludenko and Svanidze (1969).

Nilssoniopteris shiveeovoensis[132]

Sp. nov

Valid

Herrera et al.

Early Cretaceous (Aptian–Albian)

Khukhteeg Formation

Mongolia

A member of Bennettitales.

Nilssoniopteris tomentosa[132]

Sp. nov

Valid

Herrera et al.

Early Cretaceous (Aptian–Albian)

Tevshiingovi Formation

Mongolia

A member of Bennettitales.

Otozamites toshioensoi[133]

Sp. nov

Valid

Yamada, Legrand & Nishida

Early Cretaceous (Albian)

Sasayama Group

Japan

|  |

Ovalocarpus[134]

Gen. et sp. nov

Valid

Naugolnykh

Early Permian

|  |

Russia

A member of Ginkgoales belonging to the family Cheirocladaceae. Genus includes new species O. ovoides.

Pachytestopsis[135]

Gen. et sp. nov

Valid

McLoughlin, Bomfleur & Drinnan

Permian (Changhsingian)

Fort Cooper Coal Measures

Australia

A member of Glossopteridales. Genus includes new species P. tayloriorum.

Phoenicopsis kurminensis[136]

Sp. nov

Valid

Frolov in Frolov & Mashchuk

Middle Jurassic

Irkutsk Basin

Russia

A member of Leptostrobales (= Czekanowskiales).

Podozamites harrisii[137]

Sp. nov

Valid

Shi et al.

Early Cretaceous (Aptian–Albian)

Tevshiin Govi Formation

Mongolia

A conifer belonging to the family Podozamitaceae, described on the basis of leaves.

Pseudotorellia kiensis[138]

Sp. nov

Valid

Nosova & Golovneva

Late Cretaceous

|  |

Russia

A member of Ginkgoales, described on the basis of leaves.

Pseudotorellia palustris[137]

Sp. nov

Valid

Shi et al.

Early Cretaceous (Aptian–Albian)

Tevshiin Govi Formation

Mongolia

A member of Ginkgoales, described on the basis of leaves.

Pseudotorellia parvifolia[138]

Sp. nov

Valid

Nosova & Golovneva

Early Cretaceous

|  |

Russia

A member of Ginkgoales, described on the basis of leaves.

Pseudotorellia resinosa[137]

Sp. nov

Valid

Shi et al.

Early Cretaceous (Aptian–Albian)

Tevshiin Govi Formation

Mongolia

A member of Ginkgoales, described on the basis of leaves.

Pterophyllum philippoviae[139]

Sp. nov

Valid

Gnilovskaya & Golovneva

Late Cretaceous (Turonian–Coniacian)

|  |

Russia
( Magadan Oblast)

A member of Bennettitales.

Pterophyllum terechoviae[139]

Sp. nov

Valid

Gnilovskaya & Golovneva

Late Cretaceous (Maastrichtian)

Kakanaut Formation

Russia
( Koryak Okrug)

A member of Bennettitales.

Ptilozamites longifolia[140]

Sp. nov

Valid

Cariglino, Monti & Zavattieri

Middle Triassic

Quebrada de los Fósiles Formation

Argentina

A seed fern.

Rufloria glabra[141]

Sp. nov

Valid

Gomankov

Permian

|  |

Russia

A member of Pinopsida belonging to the group Cordaitanthales and to the family Rufloriaceae.

Samaropsis shenii[126]

Sp. nov

Valid

Wang & Sun in Wang et al.

Early Permian

Taiyuan Formation

China

A seed fossil belonging to the group Cordaitopsida and the family Cordaitaceae.

Solenites haojiagouensis[142]

Sp. nov

Valid

Yang et al.

Late Triassic

Haojiagou Formation

China

A member of Czekanowskiales.

Sphenopteris valentinii[143]

Sp. nov

Valid

Forte & Kerp in Forte et al.

Permian (Kungurian)

Tregiovo Formation

Italy

A fern-like plant, probably a seed fern.

Trisquama[144]

Gen. et sp. nov

Valid

Gordenko & Broushkin

Middle Jurassic (Bathonian)

|  |

Russia
( Kursk Oblast)

A gymnosperm of uncertain phylogenetic placement, belonging to the new order Trisquamales. Genus includes new species T. valentinii.

Williamsonia durikaiensis[119]

Sp. nov

Valid

McLoughlin, Pott & Sobbe

Jurassic (Pliensbachian–Aalenian)

|  |

Australia

A member of Bennettitales belonging to the family Williamsoniaceae.

Williamsonia eskensis[119]

Sp. nov

Valid

McLoughlin, Pott & Sobbe

Middle Triassic

Esk Formation

Australia

A member of Bennettitales belonging to the family Williamsoniaceae.

Williamsonia gracilis[119]

Sp. nov

Valid

McLoughlin, Pott & Sobbe

Early Cretaceous (Berriasian–Hauterivian)

Lees Sandstone

Australia

A member of Bennettitales belonging to the family Williamsoniaceae.

Williamsonia ipsvicensis[119]

Sp. nov

Valid

McLoughlin, Pott & Sobbe

Late Triassic (Carnian or earliest Norian)

Blackstone Formation

Australia

A member of Bennettitales belonging to the family Williamsoniaceae.

Williamsonia rugosa[119]

Sp. nov

Valid

McLoughlin, Pott & Sobbe

Middle Jurassic (Aalenian–Bajocian)

|  |

Australia

A member of Bennettitales belonging to the family Williamsoniaceae.

Wintucycas beatrizae[145]

Sp. nov

Valid

Martínez, Ottone & Artabe

Paleocene

Pichaihue Limestone

Argentina

A cycad belonging to the group Encephalartoideae.

Zamia nelliae[146]

Sp. nov

Valid

Erdei & Calonje in Erdei et al.

Late Eocene

Gatuncillo Formation

Panama

A cycad, a species of Zamia.

## Other plants

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Notes  | Images

|
|
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|
|
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|
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Acitheca murphyi[147]

Sp. nov

Valid

Correia et al.

Carboniferous (Gzhelian)

Douro Basin

Portugal

A marattialean fern.

Adoketophyton pingyipuensis[148]

Sp. nov

Valid

Edwards & Li

Early Devonian

Pingyipu Group

China

|  |

Apiculatasporites ruptus[149]

Sp. nov

Valid

Noetinger, di Pasquo & Starck

Devonian

|  |

Argentina

A trilete spore.

Aptychellites[150]

Gen. et sp. nov

Valid

Schäfer-Verwimp, Hedenäs, Ignatov & Heinrichs in Kaasalainen et al.

Miocene

Dominican amber

Dominican Republic

A moss resembling members of the extant genus Aptychella of the family Pylaisiadelphaceae. Genus includes new species A. fossilis.

Arthropitys taoshuyuanensis[151]

Sp. nov

Valid

Chen et al.

Permian (Wuchiapingian)

Wutonggou Formation

China

A member of Calamitaceae.

Asinisetum plaatkopensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Azolla coloniensis[153]

Sp. nov

Valid

De Benedetti et al.

Late Cretaceous

|  |

Argentina

A species of Azolla.

Balenosetum[152]

Gen. et sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Echinostachyales. Genus includes new species B. candlewaxia.

Baoyinia[154]

Gen. et sp. nov

Valid

Edwards & Li

Early Devonian

Pingyipu Group

China

A zosterophyll. Genus includes new species B. sichuanensis.

Calamospora fissurata[155]

Sp. nov

Valid

Gutiérrez & Balarino

Carboniferous (Pennsylvanian)

Ordóñez Formation

Argentina

A spore taxon.

Cetistachys[152]

Gen. et sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Echinostachyales. Genus includes new species C. cetenis.

Cheilolejeunea lamyi[156]

Sp. nov

Valid

Heinrichs et al.

Miocene

Dominican amber

Dominican Republic

A member of Lejeuneaceae.

Cladophlebis akulovii[115]

Sp. nov

Valid

Frolov in Frolov & Mashchuk

Middle Jurassic

Taltsy Formation

Russia

A fern of uncertain affinities.

Cladophlebis odintsovае[115]

Sp. nov

Valid

Frolov & Mashchuk

Middle Jurassic

Prisayanskaya Formation

Russia

A fern of uncertain affinities.

Cooksonia barrandei[157]

Sp. nov

Valid

Libertín et al.

Early Silurian

Motol Formation

Czech Republic

|  |

Coptospora santacrucensis[100]

Sp. nov

Valid

Perez Loinaze & Llorens

Early Cretaceous (Aptian)

Anfiteatro de Ticó Formation

Argentina

A spore taxon similar to spores of extant members of the families Sphaerocarpaceae, Ricciaceae and Riellaceae.

Crybelosporites corrugatus[100]

Sp. nov

Valid

Perez Loinaze & Llorens

Early Cretaceous (Aptian)

Anfiteatro de Ticó Formation

Argentina

A spore taxon related to the family Marsileaceae.

Culcita remberi[158]

Sp. nov

Valid

Pinson, Manchester & Sessa

Miocene

Clarkia fossil beds

United States
( Idaho)

A species of Culcita.

Cymatiosphaera robusta[149]

Sp. nov

Valid

Noetinger, di Pasquo & Starck

Devonian

|  |

Argentina

A prasinophyte.

Densoisporites patagonicus[100]

Sp. nov

Valid

Perez Loinaze & Llorens

Early Cretaceous (Aptian)

Anfiteatro de Ticó Formation

Argentina

A spore taxon with affinities with the Lycopsida.

Dictyophyllum menendezii[159]

Sp. nov

Valid

Bodnar et al.

Middle Triassic (Ladinian)

Cortaderita Formation

Argentina

A fern belonging to the family Dipteridaceae.

Digitopteris[160]

Gen. et sp. nov

Valid

Pott & Bomfleur in Pott et al.

Late Triassic (Carnian)

|  |

Austria

A fern belonging to the family Dipteridaceae. Genus includes new species D. repanda.

Echinostachys tinensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Echinostachyales and the family Echinostachyaceae.

Eddianna[161]

Gen. et sp. nov

Valid

Pfeiler & Tomescu

Devonian (Emsian)

Battery Point Formation

Canada
( Quebec)

A member of Rhyniopsida. Genus includes new species E. gaspiana

Electorotheca[162]

Gen. et sp. nov

Valid

Morris, Edwards & Pedersen

Devonian (Lochkovian)

Freshwater West Formation

United Kingdom

A plant of uncertain phylogenetic placement. Genus includes new species E. enigmatica.

Emphanisporites genselae[163]

Sp. nov

Valid

Wellman

Devonian (Pragian-earliest Emsian)

Val d'Amour Formation

Canada
( New Brunswick)

A plant described on the basis of fossil spores.

Emphanisporites morrisae[163]

Sp. nov

Valid

Wellman

Devonian (Pragian-earliest Emsian)

Campbellton Formation
Val d'Amour Formation

Canada
( New Brunswick)

A plant described on the basis of fossil spores.

Emphanisporites? tenuis[164]

Sp. nov

Valid

García Muro, Rubinstein & Steemans

Silurian (Přídolí)

Los Espejos Formation

Argentina

A plant described on the basis of fossil spores.

Endosporites menendezi[155]

Nom. nov

Valid

Gutiérrez & Balarino

Carboniferous (Pennsylvanian)

Agua Colorada Formation

Argentina

A spore taxon; a replacement name for Endosporites parvus Menéndez (1965).

Equisetites budagaevae[115]

Sp. nov

Valid

Frolov in Frolov & Mashchuk

Middle Jurassic

Prisayanskaya Formation

Russia

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetites greenensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetites kanensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetites kapokensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetites nuwensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetites pentapenensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetites umkensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetostachys boesmansensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetostachys calensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetostachys cervensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetostachys jaarensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetostachys kroonensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetostachys laggensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetostachys luziensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetostachys penensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Equisetostachys pokensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Escapia[165]

Gen. et sp. nov

Valid

Rothwell, Millay & Stockey

Early Cretaceous

|  |

Canada
( British Columbia)

A member of Marattiales. Genus includes new species E. christensenioides.

Frederica kurdistanensis[166]

Sp. nov

Valid

Bucur et al.

Paleogene

Khurmala Formation

Iraq

A green alga belonging to the group Dasycladales.

Frullania grabenhorstii[167]

Sp. nov

Valid

Feldberg et al.

Eocene

Bitterfeld amber

Germany

A liverwort, a species of Frullania.

Frullania zerovii[168]

Sp. nov

Valid

Mamontov, Ignatov & Perkovsky

Eocene

Rovno amber

Ukraine

A liverwort, a species of Frullania.

Geocalyx heinrichsii[169]

Sp. nov

Valid

Katagiri

Eocene

Baltic amber

Europe (Baltic Sea region)

A liverwort.

Gleicheniorachis sinensis[170]

Sp. nov

Valid

Tian et al.

Late Jurassic

Manketouebo Formation

China

A fern belonging to the family Gleicheniaceae.

Groenlandia pescheri[171]

Sp. nov

Valid

Uhl & Poschmann

Oligocene (Chattian)

Enspel Formation

Germany

A species of Groenlandia.

Heilongjiangcaulis[172]

Gen. et sp. nov

Valid

Cheng & Yang

Cretaceous

Songliao Basin

China

A tree fern. Genus includes new species H. keshanensis.

Holttumopteris[173]

Gen. et sp. nov

Valid

Regalado et al.

Cretaceous (Albian–Cenomanian )

Burmese amber

Myanmar

A eupolypod fern. Genus includes new species H. burmensis.

Horriditriletes chacoparanensis[155]

Sp. nov

Valid

Gutiérrez & Balarino

Carboniferous (Pennsylvanian)

Ordóñez Formation

Argentina

A spore taxon.

Hypnites lycopodioides[174]

Nom. nov

Valid

Ignatov & Váňa in Winterscheid et al.

Late Oligocene

Rott Formation

Germany

A member of Hypnales of uncertain phylogenetic placement; a replacement name for Hypnum lycopodioides Weber in Wessel & Weber.

Jaffrezocodium[175]

Gen. et sp. nov

Valid

Granier

Cretaceous (Albian-Cenomanian)

|  |

France
Spain

A green alga belonging to the group Bryopsidales. Genus includes new species J. bipennatus.

Jiangyounia[154]

Gen. et sp. nov

Valid

Edwards & Li

Early Devonian

Pingyipu Group

China

A rhyniophyte. Genus includes new species J. gengi.

Knorripteris taylorii[176]

Sp. nov

Valid

Galtier et al.

Triassic

|  |

Germany

A pteridophyte of uncertain phylogenetic placement.

Kowieria[177]

Gen. et sp. nov

Valid

Gess & Prestianni

Devonian (Famennian)

Witpoort Formation

South Africa

A lycopsid. Genus includes new species K. alveoformis.

Kraaiostachys[152]

Gen. et sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation
Santa Clara Formation

Mexico
South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae. Genus includes new species K. plaatkopensis.

Leiosphaeridia ibateensis[2]

Sp. nov

Valid

Arai & Dias-Brito

Late Cretaceous (Santonian)

São Carlos Formation

Brazil

An acritarch, probably a prasinophyte.

Leiotriletes malanzanensis[155]

Nom. nov

Valid

Gutiérrez & Balarino

Carboniferous (Pennsylvanian)

Malanzán Formation

Argentina

A spore taxon; a replacement name for Leiotriletes tenuis Azcuy (1975).

Lejeunea miocenica[150]

Sp. nov

Valid

Heinrichs, Schäfer-Verwimp, Renner & Lee in Kaasalainen et al.

Miocene

Dominican amber

Dominican Republic

A liverwort, a species of Lejeunea.

Lilingostrobus[178]

Gen. et sp. nov

Valid

Gerrienne et al.

Devonian (Famennian)

Xikuangshan Formation

China

A member of Lycopsida of uncertain phylogenetic placement. Genus includes new species L. chaloneri.

Marsilea mascogos[179]

Sp. nov

Valid

Estrada-Ruiz et al.

Late Cretaceous (late Campanian)

Olmos Formation

Mexico

A species of Marsilea.

Molaspora aspera[180]

Sp. nov

Valid

Zavialova & Batten

Late Cretaceous (Cenomanian)

|  |

France

A member of Marsileaceae described on the basis of megaspores.

Moltenomites[152]

Gen. et 2 sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Echinostachyales. Genus includes new species M. linearifolia and M. attenuatifolia.

Naybandoporella[181]

Gen. et sp. et comb. nov

Valid

Senowbari-Daryan

Late Triassic (Rhaetian)

Nayband Formation

Greece
Iran

A green alga belonging to the group Dasycladales, possibly a member of the family Triploporellaceae. Genus includes new species N. rhaetica, as well as "Probolocupsis" sarmeikensis Senowbari-Daryan (2014).

Oleandra bangmaii[182]

Sp. nov

Valid

Xie et al.

Late Miocene

|  |

China

A species of Oleandra.

Ornicephalum[148]

Gen. et comb. nov

Valid

Edwards & Li

Early Devonian

Pingyipu Group

China

A member of Lycophytina; a new genus for "Zosterophyllum" sichuanensis Geng (1992).

?Osmunda weylandii[174]

Sp. nov

Valid

Kvaček & Winterscheid in Winterscheid et al.

Late Oligocene

Rott Formation

Germany

A fern, possibly a species of Osmunda.

Osmundopsis zunigai[183]

Sp. nov

Valid

Coturel et al.

Late Triassic (Carnian)

Potrerillos Formation

Argentina

A fern belonging to the family Osmundaceae.

Paraschizoneura fredensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Echinostachyales and the family Echinostachyaceae.

Paraschizoneura quadripenensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Echinostachyales and the family Echinostachyaceae.

Paraschizoneura rooipoortensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Echinostachyales and the family Echinostachyaceae.

Paraschizoneura telensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Echinostachyales and the family Echinostachyaceae.

Peromonolites globosum[100]

Sp. nov

Valid

Perez Loinaze & Llorens

Early Cretaceous (Aptian)

Anfiteatro de Ticó Formation

Argentina

A spore taxon with affinities with the Filicales.

Pleurorhizoxylon[184]

Gen. et sp. nov

Valid

Zhang et al.

Late Devonian

|  |

China

An early euphyllophyte. Genus includes new species P. yixingense.

Polycladophyton[154]

Gen. et sp. nov

Valid

Edwards & Li

Early Devonian

Pingyipu Group

China

A rhyniophyte. Genus includes new species P. gracilis.

Pterospermella simplex[149]

Sp. nov

Valid

Noetinger, di Pasquo & Starck

Devonian

|  |

Argentina

A prasinophyte.

Radula intecta[150]

Sp. nov

Valid

Renner, Schäfer-Verwimp & Heinrichs in Kaasalainen et al.

Miocene

Dominican amber

Dominican Republic

A species of Radula

Rafaherbstia[185]

Ge. et sp. nov

Valid

Vera & Césari

Early Cretaceous (Aptian)

Cerro Negro Formation

Antarctica
(Livingston Island)

A cyathealean tree fern. Genus includes new species R. nishidai.

Retitriletes ornatus[100]

Sp. nov

Valid

Perez Loinaze & Llorens

Early Cretaceous (Aptian)

Anfiteatro de Ticó Formation

Argentina

A spore taxon with affinities with the Lycopodiales.

Retusotriletes archangelskyi[155]

Sp. nov

Valid

Gutiérrez & Balarino

Carboniferous (Pennsylvanian)

Ordóñez Formation

Argentina

A spore taxon.

Schizoneura cucumis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Echinostachyales and the family Echinostachyaceae.

Schizoneura koningensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Echinostachyales and the family Echinostachyaceae.

Scleropteris iljiniana[120]

Sp. nov

Valid

Kiritchkova, Kostina & Nosova

Jurassic

|  |

Russia

|  |

Sichuania[154]

Gen. et sp. nov

Valid

Edwards & Li

Early Devonian

Pingyipu Group

China

A zosterophyll. Genus includes new species S. uskielloides.

Sphenophyllum changxingense[186]

Sp. nov

Valid

Huang et al.

Late Devonian

Wutong Formation

China

|  |

Suppiluliumaella tarburensis[187]

Sp. nov

Valid

Rashidi & Schlagintweit

Late Cretaceous (Maastrichtian)

Tarbur Formation

Iran

A green alga belonging to the group Dasycladales.

Tauridium elongatum[188]

Sp. nov

Valid

Jia & Song

Late Permian

Changxing Formation

China

A member of Gymnocodiaceae.

Taurocusporites inaequalis[100]

Sp. nov

Valid

Perez Loinaze & Llorens

Early Cretaceous (Aptian)

Anfiteatro de Ticó Formation

Argentina

A spore taxon with affinities with the Bryophyta sensu lato.

Tempskya zhangii[189]

Sp. nov

Valid

Xiaonan, Fengxiang & Yeming

Cretaceous

|  |

China

A tree fern

Tiaomaphyton[190]

Gen. et sp. nov

Valid

Xu, Fu & Wang

Middle Devonian

Tiaomachian Formation

China

A Colpodexylon-like lycopsid. Genus includes new species T. fui.

Townroviamites multifoliata[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Townroviamites petfredae[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Townroviamites stellata[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Tricarinella[191]

Gen. et sp. nov

Valid

Savoretti et al.

Early Cretaceous (Valanginian)

|  |

Canada
( British Columbia)

A moss belonging to the family Grimmiaceae. Genus includes new species T. crassiphylla.

Viridistachys[152]

Gen. et 2 sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae. Genus includes new species V. moltenensis and V. gypsensis.

Zonulamites annumensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Zonulamites collensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Zonulamites elandensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Zonulamites viridensis[152]

Sp. nov

Valid

Anderson & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A member of Equisetopsida belonging to the group Equisetales and the family Equisetaceae.

Zosterophyllum ovatum[148]

Sp. nov

Valid

Edwards & Li

Early Devonian

Pingyipu Group

China

|  |

## General research

[edit]

* A study attempting to establish a timescale of early land plant evolution is published by Morris et al. (2018).[192][193][194]
* Assemblage of putative Ordovician (Hirnantian) land plants is described from the Zbrza locality in the southern Świętokrzyskie Mountains (Poland) by Salamon et al. (2018).[195]
* A study on the structure and variation of areolation patterns in leaves of Paleozoic protosphagnalean mosses is published by Ivanov, Maslova & Ignatov (2018).[196]
* A study on the phylogenetic relationships of the Cretaceous mosses Meantoinea alophosioides and Eopolytrichum antiquum within Polytrichaceae is published by Bippus, Escapa & Tomescu (2018).[197]
* Meristems of rooting axes belonging to Asteroxylon mackiei are described from the Rhynie chert (United Kingdom) by Hetherington & Dolan (2018).[198]
* A study re-examining the evidence on the speed of growth and life cycle of the tree-like lycophytes from the Carboniferous (Pennsylvanian) coal swamps, and in particular addressing an earlier study by Boyce & DiMichele (2016),[199] is published by Thomas & Cleal (2018).[200][201]
* A study on the impact of increased ultraviolet irradation (caused by volcanism-induced ozone shield deterioration) on plants during the Permian–Triassic extinction event is published by Benca, Duijnstee & Looy (2018).[202]
* A study on the composition of the Late Triassic flora of the American Southwest, based on palynological data from the Chinle Formation, and indicative of a floral turnover occurring in the middle Norian, is published by Baranyi et al. (2018).[203]
* A study on the Middle Jurassic flora from Yorkshire (United Kingdom) as indicated by pollen and spores, and on the possible dinosaur-plant interactions in the area is published by Slater et al. (2018).[204]
* Occurrence of the characean genus Tolypella is reported from the Lower Cretaceous of the Garraf Massif (Catalonia, Spain) by Martín-Closas et al. (2018), representing the oldest known record of the genus reported so far.[205]
* A study on the spore wall structure and development in Psilophyton dawsonii is published by Noetinger, Strayer & Tomescu (2018).[206]
* Lycopsid megaspores preserved with fossil starch, probably used to attract and reward animals for megaspore dispersal, are described from the Permian of north China by Liu et al. (2018).[207]
* A study on the phylogenetic relationships of extant and fossil members of Equisetales is published by Elgorriaga et al. (2018).[208]
* A study on the anatomy of the Devonian fern-like plant Shougangia bella is published by Wang et al. (2018).[209]
* A study on the phylogenetic relationships of a putative Triassic fern Pekinopteris, based on evaluation of specimens preserving fertile pinnae, is published by Axsmith, Skog & Pott (2018).[210]
* A study on the anatomical structure of Coniopteris hymenophylloides (a fossil fern belonging to the family Dicksoniaceae) based on well-preserved materials from the Middle Jurassic Yaojie Formation (China), including sterile and fertile pinnae, sporangia and in situ spores, epidermal cuticles and stomatal complexes, is published by Xin et al. (2018).[211]
* A study on the phylogenetic relationships of extant and fossil marattialean ferns is published by Rothwell, Millay & Stockey (2018).[212]
* A study on the phylogenetic relationships of members of Dipteridaceae based on data from extant and fossil taxa is published by Choo & Escapa (2018).[213]
* A study on the phylogenetic relationships of early seed plants, aneurophytalean progymnosperms, Stenokoleales and several Devonian plants of uncertain affinities is published by Toledo, Bippus & Tomescu (2018).[214]
* Plant fossils representing the genera Glossopteris, Vertebraria, Samaropsis, Paracalamites, Sphenophyllum and Dichotomopteris are described from the Permian strata in the Tabbowa Basin of Sri Lanka by Edirisooriya, Dharmagunawardhane & McLoughlin (2018), thus being the first representatives of the distinctive Permian Glossopteris flora reported from that country.[215]
* Fossils of member of the genus Glossopteris related to the species Glossopteris communis from India are described from the Permian deposits of southeastern Gobi (Mongolia) by Naugolnykh & Uranbileg (2018).[216]
* A study on the fossils of glossopterids from the Permian (Lopingian) Buckley Formation (Antarctica) will be published by DeWitt et al. (2018), who present evidence of glossopterids shedding their pollen organs during a different time of the season than Glossopteris leaves.[217]
* Blomenkemper et al. (2018) report the discovery of mixed plant-fossil assemblages in Late Permian deposits on the margins of the Dead Sea in Jordan, including fossils of seed ferns, members of Bennettitales and the earliest records of conifers reported so far.[218]
* A study on the phylogeny of conifers, comparing the inferred phylogenetic relationships and estimated divergence ages with the paleobotanical record, is published by Leslie et al. (2018).[219]
* A study on the atmospheric carbon dioxide concentration levels in the Early Cretaceous based on data from specimens of the fossil conifer species Pseudofrenelopsis papillosa is published by Jing & Bainian (2018).[220]
* A study on the phylogenetic relationships of members of Pinaceae based on data from extant and fossil taxa is published by Gernandt et al. (2018).[221]
* A study on the epidermis of the leaves of the fossil pine Pinus mikii and on the phylogenetic relationships of the species is published by Yamada & Yamada (2018).[222]
* A study on the anatomy and phylogenetic relationships of Austrohamia acanthobractea, based on data from leafy twigs with attached pollen cones and seed cones from the Middle Jurassic Daohugou Lagerstätte (China), is published by Dong et al. (2018).[223]
* Rediscovery of the holotype specimen of Weltrichia fabrei is reported by Moreau & Thévenard (2018).[224]
* Revision of gymnosperm species known from the Eocene Baltic amber is published by Alekseev (2018).[225]
* A study on the phylogenetic relationships of the vascular plants and the timescale of their evolution, attempting to establish when the flowering plants originated, is published by Barba-Montoya et al. (2018).[226]
* A study on the early evolution of Chloranthaceae, focusing on the phylogenetic relationships of the Cretaceous taxa Canrightiopsis and Pseudoasterophyllites, is published by Doyle & Endress (2018).[227]
* Fossil assemblage including plant and vertebrate remains is described from the Turonian Ferron Sandstone Member of the Mancos Shale Formation (Utah, United States) by Jud et al. (2018), who report turtle and crocodilian remains and an ornithopod sacrum, as well as a large silicified log assigned to the genus Paraphyllanthoxylon, representing the largest known pre-Campanian flowering plant reported so far and the earliest documented occurrence of an angiosperm tree more than 1.0 m in diameter.[228]
* A study on the phylogenetic relationships of extant and fossil members of Zingiberales is published by Smith et al. (2018).[229]
* A study on the phylogenetic relationships of Cornales based on data from extant and fossil taxa is published by Atkinson (2018).[230]
* A study on the microstructure of the fossils assigned to the genus Operculifructus, and on its implications for inferring the phylogenetic relationships of this genus, is published by Hayes et al. (2018).[231]
* A study on the phylogenetic relationships of the flowering plants and Gnetales, as indicated by morphological data from extant and fossil taxa, is published by Coiro, Chomicki & Doyle (2018).[232]
* Revision of the taxonomy of the Cretaceous monocot genus Viracarpon is published by Matsunaga et al. (2018), who transfer the species Coahuilocarpon phytolaccoides known from the Campanian Cerro del Pueblo Formation (Mexico) to the genus Viracarpon, thus rejecting the hypothesis that Viracarpon was endemic to India.[233]
* Microfossil remains of early grasses extracted from a specimen of the Early Cretaceous dinosaur species Equijubus normani from China are described by Wu, You & Li (2018).[234]
* Cantisolanum daturoides from the Eocene London Clay Formation, previously suggested to be a member of the family Solanaceae, is reinterpreted as more likely to be a commelinid monocot by Särkinen et al. (2018).[235]
* A study on the lower threshold of extant palm temperature tolerance, as well as on the potential of using the presence of palm fossils to infer past climate, is published by Reichgelt, West & Greenwood (2018).[236]
* A study on the human use of rainforest plant resources of prehistoric Sri Lanka, as indicated by data from phytoliths from the Fahien Rock Shelter sediments, is published by Premathilake & Hunt (2018).[237]
* A study on the occurrence of bananas in the archaeological sequence at Fahien Rock Shelter (south-west Sri Lanka), as indicated by seed and leaf phytolith evidence, is published by Premathilake & Hunt (2018).[238]
* A study on the macroevolutionary dynamics of extinction and adaptation of palms with megafaunal fruits in the late Cenozoic is published by Onstein et al. (2018), who interpret their findings as indicating that progressive loss of megafaunal frugivores during the late Cenozoic likely resulted in increased extinction rates of palms with megafaunal fruits.[239]
* A study on the floral and fruit morphology of the early eudicot species Ranunculaecarpus quinquecarpellatus is published by Manchester et al. (2018).[240]
* A study on the principal morphological characters distinguishing shade and sun leaves in modern species of Liquidambar, and on their implications for identifying leaf polymorphisms in fossil members of this genus that could otherwise be used to establish unwarranted new species, is published by Maslova et al. (2018).[241]
* A study on fossil pollen of members of the group Ericales from five Eocene localities in the United Kingdom, Austria, Germany and China, aiming to describe fossil pollen types and compare them with the most similar looking pollen of modern species, is published by Hofmann (2018).[242]
* A new fossil Loranthaceae pollen type (the first representative of this family in the fossil record of Africa) is described from the earliest Miocene of Saldanha Bay (South Africa) by Grímsson et al. (2018).[243]
* A study on the types of fossil oak pollen grains from the Last Glacial Maximum sediments from the northern South China Sea, and on their implications for inferring regional climatic conditions in this area during the Last Glacial Maximum, is published by Dai, Hao & Mao (2018).[244]
* A pistillate partial inflorescence of a member of the genus Castanopsis is described from Baltic amber by Sadowski, Hammel & Denk (2018), representing the first record of this genus from Baltic amber and the first pistillate inflorescence of Fagaceae from Eurasia reported so far.[245]
* A study on factors which influenced the diversification processes and diversity dynamics of Cenozoic woody flowering plants is published by Shiono et al. (2018).[246]
* Description of plant remains and palynomorphs preserved in the coprolites produced by large dicynodonts from the Triassic Chañares Formation (Argentina), and a study on the affinities of the plants preserved in those coprolites, is published by Perez Loinaze et al. (2018).[247]
* A study on the nutritional value of plants grown under elevated CO2 levels, evaluating the hypothesis that constraints on sauropod diet quality were driven by Mesozoic CO2 concentration, is published by Gill et al. (2018).[248]
* A study on the diversity, frequency and representation of insect damage of fossil plant specimens from the Permian La Golondrina Formation (Argentina) is published by Cariglino (2018).[249]
* A study on the insect herbivory on fossil ginkgoalean and bennettitalean leaves from the Middle Jurassic Daohugou Beds (China), and on defenses of these plants against insect herbivory, is published by Na et al. (2018).[250]
* Diverse gymnosperm and angiosperm fossils, displaying affinities with the flora of the Araripe Basin (Santana Formation) as well as those identified in deposits from the North America (Potomac Group), are described from the Lower Cretaceous Codó Formation (Brazil) by Lindoso et al. (2018).[251]
* A study on the impact of the Cenomanian-Turonian boundary event on the continental flora, as indicated by spore-pollen fossil record, is published by Heimhofer et al. (2018).[252]
* Insect and plant inclusions are reported from amber from the uppermost Campanian Kabaw Formation of Tilin (Myanmar) by Zheng et al. (2018).[253]
* Grimaldi et al. (2018) report biological inclusions (fungi, plants, arachnids and insects) in amber from the Paleogene Chickaloon Formation of Alaska, representing the northernmost deposit of fossiliferous amber from the Cenozoic.[254]
* Organically preserved plant fossils, including leaves with cuticular preservation, are described from the Paleogene Ligorio Márquez Formation (Argentina) by Carpenter, Iglesias & Wilf (2018).[255]
* A study on changes in Eocene plant diversity and floristic composition at Messel (Germany) is published by Lenz & Wilde (2018).[256]
* An amber layer is reported from the lower part of the Dingqing Formation (late Oligocene) in Lunpola of central Tibet (representing the first record of amber from Tibet) by Wang et al. (2018), who interpret this amber as derived from dipterocarp trees, and who interpret the amber layer as remains of the northernmost dipterocarp forest discovered so far.[257]
* A study on CO2 concentrations during the early Miocene, as indicated by stomatal characteristics of fossil leaves from a late early Miocene assemblage from Panama and a leaf gas-exchange model, is published by Londoño et al. (2018).[258]
* A study evaluating when the plants using the C4 photosynthetic pathway initially expanded on the Australian continent, as indicated by carbon isotope ratios of plant waxes from scientific ocean drilling sediments off north-western Australia, is published by Andrae et al. (2018).[259]
* A study on the role of fire during the expansion of C4 grassland ecosystems in the Mio-Pliocene, based on data from molecular proxies from paleosol samples of the Siwalik Group (Pakistan), is published by Karp, Behrensmeyer & Freeman (2018).[260]
* A study on the macroevolutionary responses of noctuid moths from the group Sesamiina and their associated host-grasses to environmental changes during the Neogene is published by Kergoat et al. (2018).[261]
* A study on the abundance of the C3 and C4 grasses in the central interior of southern Africa in the Early Pleistocene, as indicated by enamel stable carbon and oxygen isotope data, associated faunal abundance and phytolith evidence from the site of Wonderwerk Cave (South Africa), is published by Ecker et al. (2018).[262]
* A study on the changes of vegetation in the temperate zone of Asia during an interval containing the Mid-Pleistocene Transition, ~1.2–0.7 million years ago, as indicated by pollen data from a drilling core from the North China Plain, as well as on their effect on the large mammal fauna is published by Xinying et al. (2018).[263]
* A study on the use of plants by early modern humans during the Middle Stone Age as indicated by analyses of phytoliths from the Pinnacle Point locality (South Africa) is published by Esteban et al. (2018).[264]
* A study on the distance of seed dispersal by extant and extinct mammalian frugivores and on the impact of the extinction of Pleistocene megafauna on seed dispersal is published by Pires et al. (2018).[265]
* A study evaluating how mega-herbivore animal species controlled plant community composition and nutrient cycling, relative to other factors during and after the Late Quaternary extinction event in Great Britain and Ireland, is published by Jeffers et al. (2018).[266]
* A study on the seeds preserved in moa coprolites is published by Carpenter et al. (2018), who question the hypothesis that some of the largest-seeded plants of New Zealand were dispersed by moas.[267]
* A study on the plant–insect interactions in the European forest plant communities in the Upper Pliocene Lagerstätte of Willershausen (Lower Saxony, Germany), the Upper Pliocene locality of Berga (Thuringia, Germany) and the Pleistocene locality of Bernasso (France) is published by Adroit et al. (2018).[268]
* A study on pollen recovered from hyaena coprolites from Vanguard Cave (Gibraltar), and on its implications for reconstructing the vegetation landscapes in the environment inhabited by southern Iberian Neanderthals during the MIS 3, is published by Carrión et al. (2018).[269]
* A study on the inner structure of cuticles and carbonaceous compressions of Early Jurassic plants from Argentinian Patagonia, using Focused Ion Beam Scanning Electron Microscopy, is published by Sender et al. (2018).[270]
* A study on the changing ecology of woodland vegetation of southern mainland Greece during the late Pleistocene and the early-mid Holocene, and on the ecological context of the first introduction of crop domesticates in the southern Greek mainland, as indicated by data from carbonized fuel wood waste from the Franchthi Cave, is published by Asouti, Ntinou & Kabukcu (2018).[271]
* Evidence of plant domestication and food production from the early and middle Holocene site of Teotonio (southwestern Amazonia, Brazil) is presented by Watling et al. (2018).[272]
* A study on changes in plant pathogen communities (fungi and oomycetes) in response to changing climate during late Quaternary, as indicated by data from solidified deposits of rodent coprolites and nesting material from the central Atacama Desert spanning the last ca. 49,000 years, is published by Wood et al. (2018).[273]
* A study on the timing of the origination of the East Asian flora (including Sino-Japanese Flora Metasequoia Flora and Sino-Himalayan Rhododendron Flora), as indicated by molecular and fossil data, is published by Chen et al. (2018).[274]

## References

[edit]

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5. ^ a b c d Mahesh Prasad; Somlata Gautam; Nupur Bhowmik; Sanjeev Kumar; Sanjai Kumar Singh (2018). "New fossil leaves of Annonaceae and Achariaceae from Churia Group of Nepal and their phytogeographical implications". The Palaeobotanist. 67 (1): 47–66\. doi:10.54991/jop.2018.47. S2CID 252304015.
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