2018 in paleoichthyology

Overview of the events of 2018 in paleoichthyology

List of years in paleoichthyology

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This list of fossil fish described in 2018 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fish, bony fish, and other fish of every kind described during the year 2018, as well as other significant discoveries and events related to paleontology occurring in 2018.

## Research

[edit]

Phymolepis cuifengshanensis, life reconstruction[1]

* A study on the fossil fish occurrences and habitat during the middle Paleozoic (480 million to 360 million years ago) is published by Sallan et al. (2018), fish originated in restricted, shallow intertidal-subtidal environments.[2]
* A survey of Devonian fish fauna from Michigan is published by Stack & Sallan (2018).[3]
* A study on the ecological diversification of thelodonts is published by Ferrón et al. (2018).[4]
* The first occurrence of pelvic girdles and intromittent organs in Euphanerops longaevus, associated with a morphologically differentiated region of the axial skeleton, is reported by Chevrinais et al. (2018).[5]
* A study on the identity of the aspidin (a primitive bone-like tissue of heterostracans) is published by Keating et al. (2018), who interpret aspidin as an acellular dermal bone.[6]
* Redescription of Tesseraspis mosaica is published by Blieck, Elliott & Karatajūtė-Talimaa (2018).[7]
* A study on the morphological and taxonomic diversity of pteraspidiforms is published by Romano, Sansom & Randle (2018).[8]
* A study on the diversity of jaw shapes in modern and Paleozoic jawed fishes, evaluating whether the full extent of jaw morphological variation was established early in gnathostome evolutionary history, is published by Hill et al. (2018).[9]
* New specimens of Brindabellaspis stensioi, providing new information on the morphology of the rostral region of the skull, are described from the Lower Devonian of the New South Wales (Australia) by King, Young & Long (2018).[10]
* Redescription of the antiarch placoderm Asterolepis thule and a study on the age of the deposits preserving the fossils of this species is published by Newman & Den Blaauwen (2018).[11]
* Description of bony pelvic plates in 32 specimens of Bothriolepis canadensis from the Upper Devonian Escuminac Formation (Canada) is published by Charest, Johanson & Cloutier (2018), who reject the interpretation of these structures as genital plates (suggested by Long et al., 2015),[12] and identify them as the pelvic girdle instead.[13]
* Redescription of the antiarch placoderm Phymolepis cuifengshanensis and a study on the phylogenetic relationships of this species is published by Wang & Zhu (2018).[1]
* A study on the morphology of the skull, especially the braincase of the petalichthyid placoderm Shearsbyaspis oepiki is published by Castiello & Brazeau (2018).[14]
* Fossil interpreted as placoderm (arthrodiran) egg cases are described from the Devonian (Famennian) Cleveland Shale (Ohio, United States) by Carr & Jackson (2018).[15]
* Circular or near-circular patterned trace fossils, similar to underwater circles produced by male pufferfishes, are described from the Upper Devonian Hongguleleng Formation (China) by Zong & Gong (2018), who consider it possible that these fossils may be structures made by male fish to attract females.[16]
* Redescription of Gladbachus adentatus and a study on the phylogenetic relationships of the species is published by Coates et al. (2018).[17]
* A study on the wear of a tooth whorl of a specimen of Edestus heinrichi, as well as on its implications for inferring the function of the tooth whorls in this species, is published by Itano (2018).[18]
* New description of Edestus, providing new information on the anatomy of this taxon, is published online by Tapanila et al. (2018).[19]
* Two partial specimens of a callorhynchid chimaeroid left in open nomenclature are described from the Upper Kimmeridgian Nusplingen Plattenkalk (Germany) by Duffin (2018).[20]
* A study on the teeth histology and vasculature of the oldest known tooth-bearing sharks, Leonodus carlsi and Celtiberina maderi, is published by Martinez-Perez et al. (2018).[21]
* The first known basicranium of Carcharopsis wortheni is described from the Carboniferous Fayetteville Shale (United States) by Bronson, Mapes & Maisey (2018).[22]
* A study on the morphology of the braincase of Tristychius arcuatus is published by Coates & Tietjen (2018).[23]
* A diverse fauna of Early Triassic cartilaginous fishes is described from the Vikinghøgda Formation (Spitsbergen, Norway) by Bratvold, Delsett & Hurum (2018).[24]
* The first material referable to hybodont shark (a member of the genus Asteracanthus) is described from the Lower Jurassic (Toarcian) Rosso Ammonitico Formation (Italy) by Romano et al. (2018), providing new information on the dispersal of this genus in the Jurassic Tethys.[25]
* The first scroll coprolites from the Mesozoic reported so far, likely produced by euryhaline hybodontid sharks, are described from the Upper Triassic Tiki Formation of India by Rakshit et al. (2018), who name a new coprolite taxon Tikicopros triassicus.[26]
* Description of new remains of the Late Jurassic shark Palaeocarcharias stromeri and a study on the anatomy and phylogenetic relationships of this species is published by Landemaine, Thies & Waschkewitz (2018), who name a new order Palaeocarchariiformes and a new family Palaeocarchariidae.[27]
* Description of an articulated skeleton of a member of the palaeospinacid genus Synechodus from the Lower Cretaceous (Albian) Saint-Pô Formation (France), and a revision of the taxonomic history of the species assigned to the genus Synechodus, is published by Mollen & Hovestadt (2018).[28]
* Late Cretaceous taxon Platylithophycus cretaceus known from the Niobrara Chalk of Kansas (United States), considered to be a green alga or a cuttlefish in earlier publications, is reinterpreted as a member of Elasmobranchii by Bronson & Maisey (2018).[29]
* A study on the microstructure of enameloid in the isolated teeth of archaeobatid batomorphs Toarcibatis elongata, Cristabatis crescentiformis and Doliobatis weisi from the Jurassic (Toarcian) localities of Halanzy (Belgium) and Ginzebierg (Luxembourg) is published by Manzanares, Botella & Delsate (2018).[30]
* A study on the structure of teeth of Myledaphus pustulosus from the Upper Cretaceous (Maastrichtian) Hell Creek Formation (Montana, United States) is published by Hoffman, Jensen & Hageman (2018).[31]
* Isolated teeth of the sand shark Brachycarcharias lerichei are described from the Eocene (Ypresian) La Meseta Formation (Antarctica) by Marramà et al. (2018), representing the southernmost occurrence of the genus Brachycarcharias reported so far.[32]
* A study on the anatomy, paleobiology and paleoecology of the Eocene requiem shark Eogaleus bolcensis is published by Marramà, Carnevale & Kriwet (2018).[33]
* Teeth of members of the genera Galeorhinus and Physogaleus are described from the Lower Eocene sediments of the Khuiala Formation (Jaisalmer basin, India) by Pandey, Chaskar & Case (2018).[34]
* A study on the teeth mineralization process and teeth histology in extant and fossil members of the genus Hemipristis is published by Jambura et al. (2018).[35]
* A study on the global and regional morphological variation of the teeth of the ground sharks and mackerel sharks across the Cretaceous–Paleogene boundary is published by Bazzi et al. (2018).[36]
* A study on the anatomy and evolution of teeth of members of the families Megachasmidae and Cetorhinidae, based on data from recent and fossil teeth, is published by Mitchell, Ciampaglio & Jacquemin (2018).[37]
* A study on the physiological, ecological and life-history traits which influenced the biogeographic distributions of cartilaginous fishes from the Neogene to the present is published by Villafaña & Rivadeneira (2018).[38]
* A review of the present and past (Miocene–Pleistocene) shark and ray diversity in marine waters of Tropical America is published by Carrillo-Briceño et al. (2018).[39]
* A study on the phylogenetic relationships of extant and fossil squalomorph sharks as indicated by teeth morphology is published by Flammensbeck et al. (2018).[40]
* A study on the morphology and phylogenetic relationships of an early bony fish known from two partial skulls recovered from the Devonian (Emsian) Taemas Limestones of the Burrinjuck area (New South Wales, Australia), possibly belonging to the genus Ligulalepis (described on the basis of isolated scales), is published by Clement et al. (2018).[41]
* Redescription of Elonichthys germari is published by Schindler (2018), who presents the first reconstruction of the skull of this species.[42]
* Redescription of the neotype of the elonichthyid Rhabdolepis macropterus is published by Schindler (2018), who presents new reconstruction of the skull of this species.[43]
* A revision of ray-finned fishes from the Permian locality Buxières-les-Mines (Bourbon-l’Archambault Basin, France) is published by Štamberg (2018).[44]
* A study on the effect of Permian–Triassic and Triassic–Jurassic extinction events on ray-finned fishes is published by Smithwick & Stubbs (2018).[45]
* A study on the evolutionary history of ray-finned fishes across the Cretaceous–Paleogene extinction event, as indicated by isolated fossil teeth preserved in a South Pacific sediment core spanning 72–43 Ma, is published by Sibert et al. (2018).[46]
* A study on the morphological variation of the dorsal finlets in extant bichirs, testing the viability of these anatomic structures as a tool for taxonomic diagnoses in the study of fossil members of this group, is published by Coelho, Cupello & Brito (2018).[47]
* New data on the reproductive biology of the species Saurichthys curionii and Saurichthys macrocephalus from the Middle Triassic Meride Limestone (Monte San Giorgio, Switzerland) is presented by Maxwell et al. (2018), who identify six specimens as unambiguously gravid.[48]
* A study on the internal anatomy of the skulls of two Early Triassic specimens of Saurichthys, as well as on the phylogenetic relationships of saurichthyiforms, is published by Argyriou et al. (2018).[49]
* A comparative study on the bony labyrinth of early neopterygians, including relatives of gars and teleosts, is published by Giles, Rogers & Friedman (2018).[50]
* A study on the diversity of body shapes of neopterygians from the Triassic to the Early Cretaceous is published by Clarke & Friedman (2018).[51]
* Redescription and taxonomic reassessment of the pycnodontiform genus Cosmodus is published by Vullo et al. (2018).[52]
* A study on 52 specimens of Pycnodus from the Eocene Monte Bolca Lagerstätte (Italy), evaluating whether the morphological variability within the sample might be related to inter- or intraspecific variation, is published by Cawley et al. (2018).[53]
* A study on the anatomical structure and possible function of the flank bar-scales of members of Pycnodontiformes is published by Capasso (2018).[54]
* A revision of the phylogenetic relationships of the fossils fishes belonging to the group Halecomorphi is published by Ebert (2018).[55]
* A redescription of Asialepidotus shingyiensis and a study on the phylogenetic relationships of the species is published by Xu & Ma (2018).[56]
* A study on the phylogenetic relationships of the Triassic neopterygian Ticinolepis, as well as on the relationships of the fossil neopterygians in general, is published by López-Arbarello & Sferco (2018).[57]
* A study on the locomotion energetics of Leedsichthys problematicus, possible factors that drove the gigantism in pachycormiforms and the metabolic limits of body size in ray-finned fishes is published by Ferrón et al. (2018).[58]
* An ichthyodectiform fossil specimen preserving a small skull and anterior part of the trunk is described from a core recovered from a well drilled in the Cape Verde Basin, ca. 400 km offshore from the West African Atlantic Margin, by Casson et al. (2018).[59]
* A study on the evolutionary history of the family Catostomidae, based on data from molecules, morphology and fossil record, is published by Bagley, Mayden & Harris (2018).[60]
* A study on the phylogenetic relationships of members of Acanthomorpha and on the timescale of the radiation of this group is published by Alfaro et al. (2018), who report that crown ages for five of the six major percomorph subclades, and for the bulk of the species diversity in the sixth, coincide with the Cretaceous–Paleogene boundary.[61]
* A study on the morphology of the skeleton of Pholidophorus latiusculus, as well as on the phylogenetic relationships of this species, is published by Taverne (2018).[62]
* A study on the gill-arch anatomy in Late Cretaceous–early Paleogene members of Aulopiformes is published by Beckett, Giles & Friedman (2018).[63]
* A new specimen of Spinocaudichthys with preserved intestinal tract is described from the Cretaceous (Cenomanian) Jbel Oum Tkout Lagerstätte (Morocco) by Davesne et al. (2018).[64]
* A study on the bone histology of extant opahs, comparing it with bones of their extant and fossil relatives (including "Aipichthys" velifer), and testing the hypothesized link between endothermy and cellular bone (bone containing embedded osteocyte cells) in teleosts, is published by Davesne et al. (2018).[65]
* An articulated skeleton of a juvenile toadfish distinct from Louckaichthys novosadi is described from the Oligocene Bituminous Marls Formation (Romania) by Pikryl et al. (2018).[66]
* A study on the dynamics of diversification, phenotypic evolution and habitat transitions in the ray-finned fish group Carangaria after the Cretaceous–Paleogene extinction event is published by Ribeiro et al. (2018).[67]
* A study on the phylogenetic relationships of the fossil snake mackerels and cutlassfishes from the Eocene (Ypresian) London Clay Formation (United Kingdom) is published by Beckett et al. (2018).[68]
* Detailed description of the caudal skeleton of the Paleogene surgeonfish Arambourgthurus scombrurus is published by Carnevale & Tyler (2018).[69]
* A study on the morphology of the Oligocene percoid fish Oligoserranoides budensis is published by Bieńkowska-Wasiluk & Pałdyna (2018).[70]
* A study on the Pliocene fish fossils from the Kanapoi site (Kenya) and their implications for reconstructing lake and river environments in the Kanapoi Formation is published online by Stewart & Rufolo (2018).[71]
* A metacarpal bone of a specimen of Pteranodon, bearing teeth marks likely produced by a shark and by a saurodontid fish, is described from the Campanian Mooreville Chalk (Alabama, United States) by Ehret & Harrell (2018).[72]
* A series of neck vertebrae of Pteranodon associated with a tooth of the lamniform shark Cretoxyrhina mantelli is described from the Upper Cretaceous Niobrara Formation (Kansas, United States) by Hone, Witton & Habib (2018), who interpret the specimen as evidence of Cretoxyrhina biting Pteranodon.[73]
* A mawsoniid coelacanth specimen is described from Rhaetian deposits of the Var Department (France) by Deesri et al. (2018), representing the first known coelacanth from the marine Triassic of France.[74]
* A study on both newly collected and earlier fossil material of Ventalepis ketleriensis from the Devonian (Famennian) of Latvia and central and northwestern Russia is published by Lebedev & Lukševičs (2018), who interpret the fossils as supporting the porolepiform affinities of this species, and name a new family Ventalepididae.[75]
* Anatomical description of the endocast of "Chirodipterus" australis from the Upper Devonian Gogo Formation (Australia) is published by Henderson & Challands (2018).[76]
* A revision of the lungfish remains from the Triassic of the Świętokrzyskie Mountains and from the northeastern Poland is published by Skrzycki, Niedźwiedzki & Tałanda (2018), who report the first known Middle Triassic finding of Arganodus worldwide and the oldest known occurrence of Ptychoceratodus in Europe.[77]
* Lungfish burrows are reported for the first time from the Lower and Middle Triassic deposits of the Southern Cis-Urals by Sennikov (2018).[78]
* A study on the anatomy of the lungfish Mioceratodus gregoryi from the Eocene Redbank Plains Formation (Australia) is published by Kemp (2018).[79]
* Description of well-preserved pelvic fin skeleton of a specimen of Rhizodus hibberti from the Carboniferous (Viséan) Asbian Wardie Shales (United Kingdom) is published by Jeffery et al. (2018).[80]
* A rediagnosis and redescription of Hyneria lindae based on new remains from the Catskill Formation (Pennsylvania, United States) is published by Daeschler & Downs (2018).[81]

## New taxa

[edit]

### Jawless vertebrates

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Altigibbaspis[82]

Gen. et sp. nov

Valid

Liu, Gai & Zhu

Devonian (early Lochkovian)

Xishancun

China

A member of Galeaspida belonging to the group Polybranchiaspiformes and the family Polybranchiaspidae. The type species is A. huiqingae.

Elgaia[83]

Gen. et comb. nov

Valid

Glinskiy

Devonian

|  |

Estonia

A member of the family Psammosteidae. The type species is "Tartuosteus" luhai Mark-Kurik (1965).

Faberaspis[84]

Gen. et sp. nov

Valid

Elliott, Lassiter & Blieck

Devonian (Lochkovian)

| Drake Bay |

Canada
( Nunavut)

A member of the family Cyathaspididae. The type species is F. elgae.

Kalanaspis[85]

Gen. et sp. nov

Valid

Tinn & Märss

Silurian (Aeronian)

|  |

Estonia

An early osteostracan. Genus includes new species K. delectabilis.

Nanningaspis[86]

Gen. et sp. nov

Valid

Gai et al.

Devonian (Pragian)

Nakaolin

China

A member of Galeaspida belonging to the group Polybranchiaspiformes and the family Gumuaspidae. The type species is N. zengi.

Paralogania denisoni[87]

Sp. nov

Valid

Turner & Burrow

Silurian–Devonian boundary

| Eastport |

United States
( Maine)

A thelodont.

Platylomaspis[86]

Gen. et sp. nov

Valid

Gai et al.

Silurian (Telychian)

Tataertag

China

A member of Galeaspida belonging to the group Polybranchiaspiformes and the family Gumuaspidae. The type species is P. serratus.

Pseudolaxaspis[86]

Gen. et comb. nov

Valid

Gai et al.

Devonian (early Lochkovian)

Tataertag

China

A member of Galeaspida belonging to the group Polybranchiaspiformes and the family Gumuaspidae. The type species is "Laxaspis" rostrata Liu (1975).

Vladimirolepis[83]

Gen. et comb. nov

Valid

Glinskiy

Devonian

|  |

Estonia

A member of the family Psammosteidae. The type species is "Psammolepis" proia Mark-Kurik (1965).

### Placoderms

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Hlavinichthys[88]

Gen. et sp. nov

Valid

Carr

Devonian (Famennian)

Ohio Shale

United States
( Ohio)

A member of Aspinothoracidi. The type species is H. jacksoni.

Microbrachius kedoae[89]

Sp. nov

Valid

Mark-Kurik et al.

Devonian (Givetian)

Moroch Beds

Belarus

|  |

Wufengshania[90]

Gen. et sp. nov

Valid

Pan et al.

Devonian (Emsian)

|  |

China

A member of Antiarchi belonging to the family Bothriolepididae. The type species is W. magniforaminis.

### Acanthodians

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Notes  | Images

|
|
|
|
|
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|
|

Cheiracanthus flabellicostatus[91]

Sp. nov

Valid

Pinakhina

Middle Devonian

|  |

Russia
United Kingdom[92]

|  |

Drygantacanthus[93]

Gen. et sp. nov

Valid

Voichyshyn & Szaniawski

Early Devonian

|  |

Ukraine

A member of Ischnacanthiformes belonging to the new family Podoliacanthidae. The type species is D. semirotunda.

Ginkgolepis[94]

Gen. et sp. et comb. nov

Valid

Pinakhina & Märss

Devonian (Eifelian, Givetian, probably Famennian)

Härma Beds

Estonia
Russia

A member of Acanthodiformes belonging to the family Cheiracanthidae. The type species is G. tenericostatus; genus also includes "Cheiracanthus" talimae Valiukevičius (1985).

Jolepis[95]

Gen. et comb. nov

Valid

Burrow & Turner

Devonian (Lochkovian)

|  |

Germany
United Kingdom

A possible acanthodian of uncertain phylogenetic placement. The type species is "Diplacanthoides" robustus Brotzen (1934).

Kasperacanthus[93]

Gen. et sp. nov

Valid

Voichyshyn & Szaniawski

Early Devonian

|  |

Ukraine

A member of Ischnacanthiformes belonging to the new family Podoliacanthidae. The type species is K. serratus.

Rhadinacanthus deltosquamosus[94]

Sp. nov

Valid

Pinakhina & Märss

Devonian (Givetian)

Härma Beds

Estonia
Russia

A member of Diplacanthiformes belonging to the family Diplacanthidae.

### Cartilaginous fishes

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Acutalamna[96]

Gen. et sp. nov

Valid

Guinot & Carrillo-Briceño

Cretaceous (Albian and Cenomanian)

La Luna

Ecuador
France
Lithuania
Peru
Venezuela

A mackerel shark of uncertain phylogenetic placement. Genus includes new species A. karsteni.

Alopias palatasi[97]

Sp. nov

Valid

Kent & Ward

Miocene

Calvert

Malta
United States
( Maryland
North Carolina
South Carolina
Virginia)

A thresher shark.

Altholepis salopensis[95]

Sp. nov

Valid

Burrow & Turner

Devonian (Lochkovian)

|  |

United Kingdom

A member of Altholepidiformes belonging to the family Altholepididae.

Altusmirus[98]

Gen. et sp. nov

Valid

Fuchs et al.

Early Cretaceous (Valanginian)

|  |

Austria

A ground shark. Genus includes new species A. triquetrus.

Amaradontus[99]

Gen. et sp. nov

Valid

Hodnett & Elliott

Carboniferous (Serpukhovian)

Surprise Canyon

United States
( Arizona)

A member of Neoselachii belonging to the family Anachronistidae. The type species is A. santuccii.

Arcuodus[100]

Gen. et sp. et comb. nov

Valid

Itano & Lambert

Carboniferous (late Tournaisian to Viséan)

Bangor
Burlington
Keokuk
Salem

United States
( Alabama
Illinois
Indiana
Iowa)

A member of Holocephali belonging to the group Cochliodontiformes. The type species is A. multicuspidatus; genus also includes "Deltodopsis" bialveatus St. John & Worthen (1883).

Carcharoides lipsiensis[101]

Sp. nov

Valid

Reinecke et al.

Oligocene (Rupelian)

Böhlen

Germany

A sand shark.

Caucasochasma[102]

Gen. et sp. nov

Valid

Prokofiev & Sychevskaya

Early Oligocene

|  |

Russia
( Krasnodar Krai)

A relative of the basking shark. The type species is C. zherikhini.

Cooleyella platera[99]

Sp. nov

Valid

Hodnett & Elliott

Carboniferous (Serpukhovian)

Surprise Canyon

United States
( Arizona)

A member of Neoselachii belonging to the family Anachronistidae.

Cretacladoides[103]

Gen. et 2 sp. nov

Valid

Feichtinger et al.

Early Cretaceous (Valanginian)

|  |

Austria

A cartilaginous fish of uncertain affinities. Originally described as a possible member of the family Falcatidae; Ivanov (2022) considered its teeth to be only superficially similar to the teeth of Paleozoic falcatids, and considered it to be a possible neoselachian.[104] Genus includes new species C. ogiveformis and C. noricum.

Cretalamna bryanti[105]

Sp. nov

Valid

Ebersole & Ehret

Late Cretaceous (late Santonian and early Campanian)

Eutaw
Mooreville Chalk

United States
( Alabama)

|  |

Deltodus tubineus[106]

Sp. nov

Valid

Richards et al.

Carboniferous (Tournaisian)

Ballagan

United Kingdom

A member of Cochliodontiformes belonging to the family Cochliodontidae.

Fornicatus[98]

Gen. et sp. nov

Valid

Fuchs et al.

Early Cretaceous (Valanginian)

|  |

Austria

A ground shark. Genus includes new species F. austriacus.

Hokomata[99]

Gen. et sp. nov

Valid

Hodnett & Elliott

Carboniferous (Bashkirian)

Watahomigi

United States
( Arizona)

A member of Xenacanthiformes belonging to the family Diplodoselachidae. The type species is H. parva.

Hybodus chuanjieensis[107]

Sp. nov

|  |

Sun in Sun et al.

Middle Jurassic

Chuanjie Basin

China

|  |

Hybodus xinzhuangensis[107]

Sp. nov

|  |

Sun in Sun et al.

Middle Jurassic

Chuanjie Basin

China

|  |

Igdabatis marmii[108]

Sp. nov

Valid

Blanco

Late Cretaceous (Maastrichtian)

|  |

Spain

|  |

Lamarodus[109]

Gen. et sp. nov

Valid

Ivanov in Ivanov et al.

Permian (Guadalupian)

Bell Canyon

United States
( Texas)

A member of Hybodontiformes belonging to the superfamily Hybodontoidea. Genus includes new species L. triangulus. Announced in 2018; the final version of the article naming it was published in 2020.

Marambioraja[110]

Gen. et sp. nov

Valid

Engelbrecht et al.

Eocene (Ypresian)

La Meseta

Antarctica
(Seymour Island)

A skate. The type species is M. leiostemma.

Mesetaraja[110]

Gen. et sp. nov

Valid

Engelbrecht et al.

Eocene (Ypresian)

La Meseta

Antarctica
(Seymour Island)

A skate. The type species is M. maleficapelli.

Microcarcharias[96]

Gen. et comb. nov

Valid

Guinot & Carrillo-Briceño

Late Cretaceous

Britton
Niobrara

Canada
United States
Venezuela

A sand shark; a new genus for "Odontaspis" saskatchewanensis Case, Tokaryk & Baird (1990).

Microklomax[99]

Gen. et sp. nov

Valid

Hodnett & Elliott

Carboniferous (Serpukhovian)

Surprise Canyon

United States
( Arizona)

A member of Euselachii belonging to the family Protacrodontidae. The type species is M. carrieae.

Mooreodontus jaini[111]

Sp. nov

Valid

Bhat, Ray & Datta

Late Triassic

Tiki

India

A xenacanthid.

Natarapax[103]

Gen. et sp. nov

Valid

Feichtinger et al.

Early Cretaceous (Valanginian)

|  |

Austria

A possible member of the family Ctenacanthidae. Genus includes new species N. trivortex.

Novaculodus[99]

Gen. et sp. nov

Valid

Hodnett & Elliott

Carboniferous (Serpukhovian)

Surprise Canyon

United States
( Arizona)

A member of Euselachii belonging to the family Protacrodontidae. The type species is N. billingsleyi.

Ostarriraja[112]

Gen. et sp. nov

Valid

Marramà, Schultz & Kriwet

Miocene (Burdigalian)

|  |

Austria

A member of Rajiformes of uncertain phylogenetic placement. The type species is O. parva.

Palaeocentroscymnus[113]

Gen. et comb. nov

Valid

Pollerspöck, Flammensbeck & Straube

Miocene

Lakšárska Nová Ves

Austria
Germany
Japan
Slovakia
Spain

A sleeper shark; a new genus for "Paraetmopterus" horvathi Underwood & Schlögl (2013).

Palaeoheterodontus[114]

Gen. et comb. nov

Valid

Hovestadt

Jurassic

|  |

Germany

A bullhead shark. Genus includes "Heterodontus" sarstedtensis Thies (1983).

Pastinachus kebarensis[115]

Sp. nov

Valid

Adnet et al.

Eocene (late Bartonian)

|  |

Tunisia

A cowtail stingray of Pastinachus. Announced in 2018; the final version of the article naming it was published in 2019.

Pristrisodus[116]

Gen. et comb. nov

Valid

Bhat, Ray & Datta

Late Triassic

Tiki

India

A member of Hybodontiformes belonging to the family Lonchidiidae. Genus includes "Parvodus" tikiensis Prasad et al. (2008).

Procestracion[114]

Gen. et comb. nov

Valid

Hovestadt

Jurassic

|  |

Germany

A bullhead shark. Genus includes "Acrodus" semirugosus Plieninger (1847), "Hemipristis" bidens Quenstedt (1852) and "Strophodus" semirugosus Quenstedt (1852).

Protoheterodontus[114]

Gen. et comb. nov

Valid

Hovestadt

Late Cretaceous

|  |

France
United Kingdom

A bullhead shark. Genus includes "Heterodontus" boussioni Guinot et al. (2013).

Protohimantura[117]

Gen. et comb. nov

Valid

Marramà et al.

Miocene (Burdigalian?)

Tonasa

Indonesia

A whiptail stingray belonging to the subfamily Urogymninae. The type species is "Trygon" vorstmani de Beaufort (1926).

Raja amphitrita[110]

Sp. nov

Valid

Engelbrecht et al.

Eocene (Ypresian and Lutetian)

La Meseta
Submeseta

Antarctica
(Seymour Island)

A skate, a species of Raja.

Raja manitaria[110]

Sp. nov

Valid

Engelbrecht et al.

Eocene (Ypresian and Lutetian)

La Meseta
Submeseta

Antarctica
(Seymour Island)

A skate, a species of Raja.

Similiteroscyllium[98]

Gen. et sp. et comb. nov

Valid

Fuchs et al.

Early Cretaceous (Valanginian)

|  |

Austria
France

A carpet shark. Genus includes new species S. iniquus, as well as "Ornatoscyllium" rugasimulatum Guinot, Cappetta & Adnet (2014).

Squalicorax acutus[118]

Sp. nov

Valid

Siversson et al.

Late Cretaceous (Cenomanian)

Gearle

Australia

|  |

Squalicorax bazzii[118]

Sp. nov

Valid

Siversson et al.

Late Cretaceous (Cenomanian)

Gearle

Australia

|  |

Squalicorax lalunaensis[96]

Sp. nov

Valid

Guinot & Carrillo-Briceño

Late Cretaceous (Cenomanian)

La Luna

Venezuela

|  |

Squalicorax moodyi[96]

Sp. nov

Valid

Guinot & Carrillo-Briceño

Late Cretaceous (Cenomanian)

La Luna

Venezuela

|  |

Squalicorax mutabilis[118]

Sp. nov

Valid

Siversson et al.

Late Cretaceous (Cenomanian and Turonian)

Haycock

Australia

|  |

Stethacanthus concavus[119]

Sp. nov

Valid

Ginter

Carboniferous (Gzhelian)

Indian Cave

United States
( Nebraska)

|  |

Tamiobatis elgae[120]

Sp. nov

Valid

Ivanov in Ivanov & Plax

Devonian (Famennian) and Carboniferous (Tournaisian)

|  |

Belarus

A member of Ctenacanthiformes belonging to the family Ctenacanthidae.

Tikiodontus[111]

Gen. et sp. nov

Valid

Bhat, Ray & Datta

Late Triassic

Tiki

India

A xenacanthid. Genus includes new species T. asymmetricus.

Truyolsodontos[121]

Gen. et sp. nov

Valid

Bernárdez

Late Cretaceous (Cenomanian)

|  |

Spain

A mackerel shark. Genus includes new species T. estauni.

Whitropus[106]

Gen. et sp. nov

Valid

Richards et al.

Carboniferous (Tournaisian)

Ballagan

United Kingdom

A member of Cochliodontiformes of uncertain phylogenetic placement. The type species is W. longicalcus.

### Ray-finned fishes

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Allogenartina[122]

Gen. et sp. nov

Valid

Schwarzhans, Huddleston & Takeuchi

Late Cretaceous (Santonian and Campanian)

Eutaw

United States
( Alabama
New Jersey)

Possibly a member of Stomiiformes belonging to the group Gonostomatoidei, of uncertain phylogenetic placement within the latter group. The type species is A. muscogeei.

Amakusaichthys[123]

Gen. et sp. nov

Valid

Yabumoto, Hirose & Brito

Late Cretaceous (Santonian)

Hinoshima

Japan

A member of Ichthyodectiformes. Genus includes new species A. goshouraensis. Announced in 2018; the final version of the article naming it was published in 2020.

Ameiurus grangerensis[124]

Sp. nov

Valid

Smith, Martin & Carpenter

Miocene

Ellensburg

United States
( Washington)

A species of Ameiurus.

Amphiarius paleoorinocoensis[125]

Sp. nov

Valid

Aguilera & Marceniuk

Late Miocene

Urumaco

Venezuela

A species of Amphiarius.

Anchiacipenser[126]

Gen. et sp. nov

Valid

Sato et al.

Late Cretaceous (Campanian)

Dinosaur Park

Canada
( Alberta)

A sturgeon. Genus includes new species A. acanthaspis.

Andrewsolepis[127]

Gen. et sp. nov

Valid

Elliott

Carboniferous (Bashkirian)

Lower Coal Measures

United Kingdom

A member of Actinopteri belonging to the family Haplolepidae. The type species is A. lochlani.

Apateodus? assisi[122]

Sp. nov

Valid

Schwarzhans, Huddleston & Takeuchi

Late Cretaceous (Santonian)

Eutaw

United States
( Alabama)

A member of Aulopiformes belonging to the family Ichthyotringidae.

Aphanolebias sarmaticus[128]

Sp. nov

Valid

Reichenbacher, Filipescu & Miclea

Middle Miocene

|  |

Romania

|  |

Apsopelix ?berlinensis[129]

Sp. nov

Valid

Schwarzhans

Early Cretaceous (Berriasian to Valanginian)

|  |

Germany
Israel

A member of Crossognathiformes belonging to the family Crossognathidae.

Archaeglossus[129]

Gen. et comb. et sp. nov

Valid

Schwarzhans

Jurassic (Bathonian to Tithonian)

|  |

United Kingdom

A member of Osteoglossiformes of uncertain phylogenetic placement. The type species is "Otolithus (Leptolepidarum)" pentangulatus Frost (1924); genus also includes new species A. torrensi.

Ariopsis ariopsilus[125]

Sp. nov

Valid

Aguilera & Marceniuk

Late Miocene

Urumaco

Venezuela

A species of Ariopsis.

Atherina carnevalei[128]

Sp. nov

Valid

Reichenbacher, Filipescu & Miclea

Middle Miocene

|  |

Romania

A species of Atherina.

Aulothrissus[129]

Gen. et 2 sp. nov

Valid

Schwarzhans

Early Cretaceous (Berriasian to Valanginian)

Gevaram

Israel

A member of Elopiformes of uncertain phylogenetic placement. The type species is A. avitus; genus also includes A. heletzensis.

Avonichthys[130]

Gen. et sp. nov

|  |

Wilson, Pardo & Anderson

Carboniferous (Tournaisian)

Horton Bluff

Canada
( Nova Scotia)

An early ray-finned fish. The type species is A. manskyi.

Bagre urumacoensis[125]

Sp. nov

Valid

Aguilera & Marceniuk

Late Miocene

Urumaco

Venezuela

A species of Bagre.

Benthalbella praecessor[131]

Sp. nov

Valid

Nazarkin & Carnevale

Miocene (late Langhian–early Serravallian)

Kurasi

Russia
( Sakhalin Oblast)

A species of Benthalbella.

Bicavolithus[129]

Gen. et sp. nov

Valid

Schwarzhans

Early Cretaceous (Albian)

|  |

Germany

A teleost of uncertain phylogenetic placement. Genus includes new species B. cavatus.

Blairolepis wallacei[127]

Sp. nov

Valid

Elliott

Carboniferous (Bashkirian)

Lower Coal Measures

United Kingdom

A member of Actinopteri belonging to the family Haplolepidae.

Blennius? martinii[128]

Sp. nov

Valid

Reichenbacher, Filipescu & Miclea

Middle Miocene

|  |

Romania

A combtooth blenny.

Bluefieldius[132]

Gen. et sp. nov

Valid

Mickle

Carboniferous (late Mississippian)

Bluefield

United States
( West Virginia)

An early ray-finned fish. The type species is B. mercerensis.

Braccohaplolepis[127]

Gen. et sp. nov

Valid

Elliott

Carboniferous (Bashkirian)

Lower Coal Measures

United Kingdom

A member of Actinopteri belonging to the family Haplolepidae. The type species is B. fenestratum.

Cerinichthys[55]

Gen. et sp. nov

Valid

Ebert

Late Jurassic

|  |

France

A member of Halecomorphi belonging to the new order Ophiopsiformes. Genus includes new species C. koelblae.

Champsodon tethensis[133]

Sp. nov

Valid

Bannikov

Eocene (Bartonian)

|  |

Russia

A species of Champsodon

Chaoia[134]

Gen. et comb. nov

Valid

Bannikov, Schwarzhans & Carnevale

Neogene

|  |

Austria
Croatia
Romania

A member of the family Sciaenidae. The type species is "Sciaena" moguntiniformis Pana (1977).

Clupea macrocephala[135]

Sp. nov

Junior homonym

Yabumoto & Nazarkin

Miocene

Bessho

Japan

A species of Clupea. The specific name is preoccupied by Clupea macrocephala Lacépède (1803); Yabumoto & Nazarkin (2020) coined a replacement name Clupea hanishinaensis.[136]

Coryphaenoides scrupus[137]

Sp. nov

Valid

Brzobohatý & Nolf

Middle Miocene

|  |

Czech Republic

A species of Coryphaenoides.

Cowetaichthys[122]

Gen. et 2 sp. et comb. nov

Valid

Schwarzhans, Huddleston & Takeuchi

Late Cretaceous (Santonian to Maastrichtian) and Paleocene

Eutaw

Denmark
Germany
Greenland
United States
( Alabama)

A beardfish. The type species is C. alabamae; genus also includes new species C. amberi, as well as "genus Polymixiidarum" beaury Schwarzhans (2010), "genus Veliferidarum" groenlandicus Schwarzhans (2004) and "genus Veliferidarum" harderi Schwarzhans (2003).

Cretolepis[138]

Gen. et sp. nov

Valid

Wang et al.

Early Cretaceous

Longjiang

China

A member of Palaeonisciformes. The type species is C. dongbeiensis.

Croatosciaena[134]

Gen. et sp. nov

Valid

Bannikov, Schwarzhans & Carnevale

Middle Miocene

|  |

Croatia

A member of the family Sciaenidae. The type species is C. krambergeri.

Cubariomma[139]

Gen. et sp. nov

Valid

Bannikov

Early Oligocene

|  |

Russia
( Adygea)

A member of Stromateoidei of uncertain phylogenetic placement. Genus includes new species C. yanakuzminae.

Dapedium ballei[140]

Sp. nov

Valid

Maxwell & López-Arbarello

Middle Jurassic (Aalenian)

Opalinuston

Germany

|  |

Doggerichthys[129]

Gen. et sp. nov

Valid

Schwarzhans

Middle Jurassic (Aalenian

|  |

Germany

A member of Leptolepidiformes sensu lato of uncertain phylogenetic placement. Genus includes new species D. anguilliformis.

Eleogobius prochazkai[128]

Sp. nov

Valid

Reichenbacher, Filipescu & Miclea

Middle Miocene

|  |

Romania

|  |

Elops eutawanus[122]

Sp. nov

Valid

Schwarzhans, Huddleston & Takeuchi

Late Cretaceous (Santonian)

Eutaw

United States
( Alabama)

A species of Elops.

Eoalosa[141]

Gen. et sp. nov

Valid

Marramà & Carnevale

Eocene (Ypresian)

Monte Bolca

Italy

A member of the family Clupeidae. The type species is E. janvieri.

Eoellimmichthys[142]

Gen. et sp. nov

Valid

Marramà et al.

Eocene (late Ypresian)

Monte Bolca

Italy

A member of Clupeomorpha belonging to the group Ellimmichthyiformes and the family Paraclupeidae. The type species is E. superstes.

Eutawichthys[122]

Gen. et 2 sp. et comb. nov

Valid

Schwarzhans, Huddleston & Takeuchi

Late Cretaceous (Santonian to Maastrichtian)

Eutaw

United States
( Alabama
Maryland
Mississippi
New Jersey
Tennessee)

A member of Beryciformes belonging to the group Berycoidei, of uncertain phylogenetic placement within the latter group. The type species is E. stringeri; genus also includes new species E. compressus, as well as “genus Apogonidarum” maastrichtiensis Nolf & Stringer (1996) and “genus Apogonidarum” zideki Nolf & Stringer (1996).

Furloichthys[143]

Gen. et sp. nov

Valid

Taverne & Capasso

Late Cretaceous (Cenomanian)

|  |

Italy

A member of the family Ichthyodectidae. The type species is F. bonarellii.

Fuyuanichthys[144]

Gen. et sp. nov

Valid

Xu, Ma & Ren

Middle Triassic (Ladinian)

Falang

China

An early member of Ginglymodi. The type species is F. wangi.

Genartina cretacea[122]

Sp. nov

Valid

Schwarzhans, Huddleston & Takeuchi

Late Cretaceous (Santonian)

Eutaw

United States
( Alabama)

Possibly a member of Stomiiformes.

Gobius apuseni[128]

Sp. nov

Valid

Reichenbacher, Filipescu & Miclea

Middle Miocene

|  |

Romania

A species of Gobius.

Gobius holcovae[128]

Sp. nov

Valid

Reichenbacher, Filipescu & Miclea

Middle Miocene

|  |

Romania

A species of Gobius.

Gobius manfredi[128]

Sp. nov

Valid

Reichenbacher, Filipescu & Miclea

Middle Miocene

|  |

Romania

A species of Gobius.

Grimmenichthys[145]

Gen. et sp. nov

Valid

Konwert & Hörnig

Early Jurassic (Toarcian)

|  |

Germany

A member of Pholidophoriformes. Genus includes new species G. ansorgei.

Guimarotaichthys[129]

Gen. et sp. nov

Valid

Schwarzhans

Late Jurassic

Camadas de Guimarota

Portugal

A non-teleost bony fish of uncertain phylogenetic placement. Genus includes new species G. problematicus.

Haqelpycnodus[146]

Gen. et sp. nov

Valid

Taverne & Capasso

Late Cretaceous (Cenomanian)

|  |

Lebanon

A member of the family Pycnodontidae. The type species is H. picteti.

Ijimaia rara[137]

Sp. nov

Valid

Brzobohatý & Nolf

Middle Miocene

|  |

Czech Republic

A species of Ijimaia.

Italoalbula[147]

Gen. et sp. nov

Valid

Taverne & Capasso

Early Cretaceous (Albian)

| Pietraroja |

Italy

A bonefish. The type species is I. pietrarojae.

Joffrichthys tanyourus[148]

Sp. nov

Valid

Murray et al.

Paleocene

Paskapoo

Canada
( Alberta)

A member of Osteoglossomorpha of uncertain phylogenetic placement.

Kalabisia[149]

Gen. et sp. nov

Valid

Přikryl & Carnevale

Oligocene–early Miocene

Ždánicko-Hustopeče

Czech Republic

A viviparous brotula belonging to the subfamily Brosmophycinae. The type species is K. krumvirensis.

Lagocephalus striatus[150]

Sp. nov

Valid

Aguilera et al.

Miocene (Serravallian)

Valiente Peninsula

Panama

A species of Lagocephalus.

Lanarkichthys[127]

Gen. et sp. nov

Valid

Elliott

Carboniferous (Bashkirian)

Lower Coal Measures

United Kingdom

A member of Actinopteri of uncertain phylogenetic placement. The type species is L. gardineri.

Landinisciaena[134]

Gen. et sp. et comb. nov

Valid

Bannikov, Schwarzhans & Carnevale

Miocene (late Burdigalian or early Langhian)

|  |

Belgium
Germany
Russia

A member of the family Sciaenidae. The type species is L. popovi; genus also includes "Atractoscion" elongatissimus Schwarzhans (1993).

Lanxangichthys[151]

Gen. et sp. nov

Valid

Cavin et al.

Early Cretaceous (Aptian)

Grès supérieurs
Khok Kruat?

Laos
Thailand?

A gar. Genus includes new species L. alticephalus.

Lasalichthys otischalkensis[152]

Sp. nov

Valid

Gibson

Late Triassic

Dockum Group

United States
( Texas)

A member of Redfieldiiformes.

Leptoelops[129]

Gen. et comb. nov

Valid

Schwarzhans

Middle Jurassic

|  |

Germany

A member of Elopiformes of uncertain phylogenetic placement. The type species is Otolithus (Lycopteridarum) rhenanus Weiler (1954); genus also includes Otolithus (Lycopteridarum) acutus Weiler (1954).

Leptosciaena[134]

Gen. et comb. et sp. nov

Valid

Bannikov, Schwarzhans & Carnevale

Neogene

|  |

Azerbaijan
Italy
Kazakhstan

A member of the family Sciaenidae. The type species is "Genyonemus" karagiensis Bratishko, Schwarzhans & Reichenbacher (2015); genus also includes "Otolithus (Mugilidarum)" azerbaidjanicus Djafarova (2006), as well as new species L. caputoi.

Leuroglossus kobylianskyi[153][154]

Sp. nov

Valid

Nazarkin

Middle–Late Miocene

Kurasi

Russia
( Sakhalin Oblast)

A species of Leuroglossus

Libanopycnodus[155]

Gen. et sp. nov

Valid

Taverne & Capasso

Late Cretaceous (Cenomanian)

|  |

Lebanon

A member of the family Pycnodontidae. The type species is L. wenzi.

Lopadichthys[148]

Gen. et sp. nov

Valid

Murray et al.

Paleocene

Paskapoo

Canada
( Alberta)

A member of Osteoglossiformes. The type species is L. colwellae.

Macroynis[68]

Gen. et sp. nov

Valid

Beckett et al.

Eocene

London Clay

United Kingdom

A relative of snake mackerels and cutlassfishes. Genus includes new species M. casieri.

Motlayoichthys[156]

Gen. et sp. nov

Valid

Arratia, González-Rodríguez & Hernández-Guerrero

Cretaceous (Albian-Cenomanian)

El Doctor

Mexico

A member of Crossognathiformes belonging to the family Pachyrhizodontidae. The type species is M. sergioi.

Mylocheilus kevinmeeksi[124]

Sp. nov

Valid

Smith, Martin & Carpenter

Miocene

Ellensburg

United States
( Washington)

A relative of the peamouth.

Nardoglossus[157]

Gen. et sp. nov.

Valid

Taverne & Capasso

Late Cretaceous (Campanian-Maastrichtian)

Altamura

Italy

A member of Gonorynchiformes belonging to the group Gonorynchoidei. The type species is N. sanctibernardini.

Nipponocypris takayamai[158]

Sp. nov

Valid

Miyata, Yabumoto & Hirano

Middle Pleistocene

Nogami

Japan

A species of Nipponocypris.

Palealbula declivis[129]

Sp. nov

Valid

Schwarzhans

Early Cretaceous (Valanginian)

|  |

Germany

A member of Albuliformes of uncertain phylogenetic placement.

Palealbula depressidorsalis[129]

Sp. nov

Valid

Schwarzhans

Early Cretaceous (Albian)

|  |

United Kingdom

A member of Albuliformes of uncertain phylogenetic placement.

Paleoschizothorax[159]

Gen. et sp. nov

Valid

Yang et al.

Oligocene

Qaidam Basin

China

A member of the family Cyprinidae related to members of the genus Schizothorax. Genus includes new species P. qaidamensis.

Paleoserranus[160]

Gen. et sp. nov

Valid

Cantalice, Alvarado-Ortega & Alaniz-Galvan

Paleocene

|  |

Mexico

A member of the family Serranidae. Genus includes new species P. lakamhae.

Parahaplolepis poppaea[127]

Sp. nov

Valid

Elliott

Carboniferous (Bashkirian)

Lower Coal Measures

United Kingdom

A member of Actinopteri belonging to the family Haplolepidae.

Peltopleurus tyrannos[161]

Sp. nov

Valid

Xu, Ma & Zhao

Middle Triassic (Ladinian)

Falang

China

|  |

Pickeringius[162]

Gen. et sp. nov

Valid

Choo et al.

Late Devonian

Gogo

Australia

An early ray-finned fish. Genus includes new species P. acanthophorus.

Piranhamesodon[163]

Gen. et sp. nov

Valid

Kölbl-Ebert et al.

Late Jurassic (late Kimmeridgian to early Tithonian)

|  |

Germany

A member of Pycnodontiformes of uncertain phylogenetic placement. The type species is P. pinnatomus.

Pontosciaena[134]

Gen. et comb. nov

Valid

Bannikov, Schwarzhans & Carnevale

Neogene

|  |

Crimean Peninsula  Azerbaijan?
Romania?

A member of the family Sciaenidae. The type species is "Serranus" acuterostratus Rückert-Ülkümen (1996); genus might also include "Otolithus (Percidarum)" sigmoilinoides Pobedina (1956).

Protalbula websteri[129]

Sp. nov

Valid

Schwarzhans

Late Jurassic (Kimmeridgian)

|  |

United Kingdom

A member of Albuliformes of uncertain phylogenetic placement.

Protarpon boualii[164]

Sp. nov

Valid

Khalloufi et al.

Paleocene (Danian)

Oulad Abdoun Basin

Morocco

A relative of tarpons.

Protoelops[129]

Gen. et comb. nov

Valid

Schwarzhans

Middle Jurassic to Early Cretaceous (Bathonian to Aptian)

|  |

Spain
United Kingdom

A member of the family Elopidae. The type species is "Otolithus (Leptolepidarum)" cuneiformis Frost (1924); genus also includes "Leptolepis" tenuirostris Stinton (1968) and "genus Protacanthopterygiorum" scalpellum Nolf (2004).

Protosciaena kirbyorum[165]

Sp. nov

Valid

Stringer & Bell

Early Pliocene

|  |

United States
( Georgia (U.S. state))

A member of the family Sciaenidae.

Pseudogonatodus aurulentum[127]

Sp. nov

Valid

Elliott

Carboniferous (Bashkirian)

Lower Coal Measures

United Kingdom

A member of Actinopteri belonging to the family Gonatodidae.

Pseudohaplolepis[127]

Gen. et sp. nov

Valid

Elliott

Carboniferous (Bashkirian)

Lower Coal Measures

United Kingdom

A member of Actinopteri of uncertain phylogenetic placement. The type species is P. argentatum.

Pseudotrichiurus[122]

Gen. et sp. nov

Valid

Schwarzhans, Huddleston & Takeuchi

Late Cretaceous (Santonian)

Eutaw

United States
( Alabama)

A member of Aulopiformes of uncertain phylogenetic placement. The type species is P. sagax.

Pterothrissus carolinensis[166]

Sp. nov

Valid

Stringer et al.

Late Cretaceous (early Campanian)

Tar Heel

United States
( North Carolina)

A relative of the Japanese gissu.

Quasimodichthys[167]

Gen. et comb. nov

Valid

Paiva & Gallo

Late Jurassic (Oxfordian)

Pastos Bons

Brazil

A member of Semionotiformes. The type species is "Lepidotus" piauhyensis Roxo & Löfgren (1936)

Rhadinichthys glabrolepis[127]

Sp. nov

Valid

Elliott

Carboniferous (Bashkirian)

Lower Coal Measures

United Kingdom

A member of Actinopteri belonging to the family Rhadinichthyidae.

Rhadinichthys? ornatocephalum[127]

Sp. nov

Valid

Elliott

Carboniferous (Bashkirian)

Lower Coal Measures

United Kingdom

A member of Actinopteri belonging to the family Rhadinichthyidae.

Rhadinichthys? plumosum[127]

Sp. nov

Valid

Elliott

Carboniferous (Bashkirian)

Lower Coal Measures

United Kingdom

A member of Actinopteri belonging to the family Rhadinichthyidae.

Sanjuanableps[168]

Gen. et sp. nov

Valid

Bogan et al.

Late Miocene

|  |

Argentina

A member of the family Anablepidae. Genus includes new species S. calingasta.

Saurichthys spinosa[169]

Sp. nov

Valid

Wu, Sun & Fang

Middle Triassic (Anisian)

Guanling

China

|  |

Sciades latissimum[125]

Sp. nov

Valid

Aguilera & Marceniuk

Late Miocene

Urumaco

Venezuela

A species of Sciades.

Sciades peregrinus[125]

Sp. nov

Valid

Aguilera & Marceniuk

Late Miocene

Urumaco

Venezuela

A species of Sciades.

Scopulipiscis[170]

Gen. et sp. nov

Valid

Latimer & Giles

Late Triassic

Kössen

Switzerland

A relative of Dapedium. Genus includes new species S. saxciput.

Sigmapycnodus[155]

Gen. et sp. nov

Valid

Taverne & Capasso

Late Cretaceous (Cenomanian)

|  |

Lebanon

A member of the family Pycnodontidae. The type species is S. giganteus.

Sphaeronchus rundlei[129]

Sp. nov

Valid

Schwarzhans

Late Jurassic (Kimmeridgian)

Kimmeridge Clay

United Kingdom

A member of Leptolepidiformes sensu lato of uncertain phylogenetic placement.

Vinctifer ferrusquiai[171]

Sp. nov

Valid

Cantalice, Alvarado-Ortega & Brito

Late Jurassic (Kimmeridgian)

Sabinal

Mexico

A member of Aspidorhynchiformes belonging to the family Aspidorhynchidae.

Vox thlotlo[122]

Gen. et sp. nov

Valid

Schwarzhans, Huddleston & Takeuchi

Late Cretaceous (Santonian)

Eutaw

United States
( Alabama)

A teleost of uncertain phylogenetic placement. The type species is V. thlotlo.

Wettonius[172]

Gen. et sp. nov

Valid

Carnevale & Bannikov

Eocene

Monte Bolca
Nanjemoy Formation

Italy
United States
( Virginia)[173]

A member of the family Veliferidae. Genus includes new species W. angeloi.

Xenoleptolepis[129]

Gen. et comb. nov

Valid

Schwarzhans

Early Jurassic to Early Cretaceous (Sinemurian to Berriasian)

|  |

Germany
United Kingdom

A member of the family Leptolepididae sensu lato. The type species is Otolithus (incertae sedis) withersi Frost (1926); genus also includes Otolithus (Salmonoidei) oncorhynchoides Weiler (1954).

Yuskaichthys[174]

Gen. et sp. nov

Valid

Bogan, Agnolin & Scanferla

Eocene

Maíz Gordo

Argentina

A member of the family Andinichthyidae. The type species is Y. eocenicus.

Zaprora koreana[175]

Sp. nov

Valid

Nam & Nazarkin

Neogene

|  |

South Korea

A relative of the prowfish.

Zurupleuropholis[176]

Gen. et 2 sp. nov

Valid

Giordano et al.

Early Cretaceous (Albian)

Lagarcito

Argentina

A relative of Pleuropholis. Genus includes new species Z. quijadensis and Z. decollavi.

### Lobe-finned fishes

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Celsiodon[177]

Gen. et sp. nov

Valid

Clack et al.

Devonian (late Famennian)

|  |

Greenland

A lungfish related to Ctenodus. Genus includes new species is C. ahlbergi.

Ceratodus tunuensis[178]

Sp. nov

Valid

Agnolin et al.

Late Triassic

Fleming Fjord

Greenland

A lungfish.

Eusthenopteron jenkinsi[179]

Sp. nov

Valid

Downs et al.

Devonian (Frasnian)

Fram

Canada
( Nunavut)

|  |

Mawsonia soba[180]

Sp. nov

Valid

Brito et al.

Early Cretaceous

Babouri Figuil Basin

Cameroon

A coelacanth.

Ptychoceratodus oldhami[181]

Sp. nov

Valid

Bhat & Ray

Late Triassic (Carnian)

Tiki

India

A lungfish. Announced in 2018; the final version of the article naming it was published in 2020.

## References

[edit]

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