2019 in brachiopod paleontology

Overview of the events of 2019 in paleontology

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Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2019.

## Research

[edit]

* A study aiming to reconstruct the life cycle of the Early Cambrian brachiopods is published by Madison & Kuzmina (2019).[2]
* A study on the anatomy of plectambonitoid specimens from the Middle Ordovician of Russia, and on its implications for the knowledge of life cycles of these brachiopods, is published online by Madison & Kuzmina (2019).[3]
* A study on the petrographic and geochemical preservation of Ordovician dalmanelloid shells from the Lexington Formation of Kentucky, Sheguindah Shale of Ontario and the Stony Mountain Formation of Manitoba, aiming to test the hypothesis of paleo-latitudinal zonation of the shelly benthos, is published by Azmy & Jin (2019).[4]
* A study on the phylogenetic relationships among strophomenoid brachiopods and on the biogeographical changes in the strophomenoids through time (focusing on the impact of the Late Ordovician mass extinction on the evolutionary history of strophomenoids) is published by Congreve, Krug & Patzkowsky (2019).[5]
* A study on the internal structure of the shell of Semiplanella carinthica is published by Pakhnevich (2019), who names a new tribe Semiplanellini in the subfamily Gigantoproductinae.[6]
* A study on the diet of species of Gigantoproductus from the Viséan of Derbyshire (United Kingdom), and on its implications for the knowledge of mechanisms that enabled those brachiopod species to reach large sizes, is published by Angiolini et al. (2019).[7]
* A study on the relative importance of brachiopods and bivalves in the fossil assemblages from the Carboniferous Pennsylvanian Breathitt Formation of Kentucky is published by Hsieh, Bush & Bennington (2019).[8]
* Description of a specimen of a Permian brachiopod species Spiriferella protodraschei bearing ventrally directed spiralia (lophophore-supporting, coiled internal structures), and a study on the morphology of these structures and their implications for the knowledge of the feeding system of this brachiopod, is published by Lee et al. (2019).[9]
* A study on the evolution of the body size of brachiopods from the Late Permian to the Middle Triassic, as indicated by brachiopod specimens from South China, is published by Chen et al. (2019).[10]
* A study on the biogeography of Pliensbachian brachiopods in western Tethys Ocean is published online by Vörös & Escarguel (2019).[11]
* A study on changes in the body size of benthic marine brachiopods and bivalves from the Lusitanian Basin (Portugal) before the Toarcian oceanic anoxic event is published by Piazza et al. (2019).[12]
* A study on the impact of the early Toarcian extinction event on fossil brachiopods and bivalves known from the Iberian Range (Spain) is published by Danise et al. (2019).[13]

## New taxa

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Aegiria apta[14]

Sp. nov

Valid

Huang et al.

Silurian (Rhuddanian)

Zhangwan Formation

China

|  |

Alaskorhynchus[15]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

United States

A member of Rhynchonellida belonging to the family Eatoniidae. The type species is A. sodacreekensis.

Angustothyris dagysi[16]

Sp. nov

Valid

Guo, Chen & Harper

Middle Triassic (Anisian)

Qingyan Formation

China

|  |

Angustothyris qingyanensis[16]

Sp. nov

Valid

Guo, Chen & Harper

Middle Triassic (Anisian)

Qingyan Formation

China

|  |

Anidanthus parvimucronata[17]

Nom. nov

Valid

He & Shen in He et al.

Permian (Changhsingian)

Talung Formation

China

A member of Productida belonging to the family Linoproductidae and the subfamily Anidanthinae; a replacement name for Anidanthus mucronata He & Shen in He et al. (2005).

Anidanthus subquadratus[17]

Sp. nov

Valid

He, Shi & Shen in He et al.

Permian (Changhsingian)

|  |

China

A member of Productida belonging to the family Linoproductidae and the subfamily Anidanthinae.

Aporthophyla prisca[18]

Sp. nov

Valid

Cocks & Popov

Early Ordovician

|  |

United Kingdom

|  |

Arcullina? enokiani[19]

Sp. nov

Valid

Lee & Shi in Lee et al.

Permian (Kungurian)

Kozhim Formation
Kozhim Rudnik Formation

Russia

A member of Spiriferida belonging to the family Spiriferellidae.

Askerina[20]

Gen. et sp. nov

Valid

Baarli

Ordovician (Hirnantian) and Silurian (Aeronian)

Solvik

Norway

A member of the family Atrypidae. The type species is A. cymbula. Announced in 2019; the final version of the article naming was published in 2021.

Aulacothyris stevensi[21]

Sp. nov

Valid

MacFarlan

Early Jurassic (Aratauran)

Diamond Peak Group

New Zealand

A member of Terebratulida belonging to the family Zeilleriidae.

Aulonotreta neptuni[22]

Sp. nov

Valid

Holmer et al.

Ordovician (Darriwilian )

|  |

Sweden

A member of Lingulida belonging to the family Aulonotretidae.

Australocoelia boucoti[23]

Sp. nov

Valid

De Rezende, Machado & Ponciano

Devonian

|  |

Brazil

|  |

Austriellula iordanae[24]

Sp. nov

Valid

Gaetani in Grădinaru & Gaetani

Triassic

|  |

Romania

A member of Rhynchonellida belonging to the family Norellidae and the subfamily Norellinae.

Bellistrophia[25]

Gen. et comb. nov

Valid

Holmer et al.

Cambrian

Atei Formation
Meagher Formation

Kazakhstan
United States

A member of Kutorginida. The type species is "Nisusia" deissei Bell (1941); genus also includes "Nisusia" montanensis Bell (1941).

Bernousia[26]

Gen. et sp. nov

Valid

Gourvennec

Paleozoic

Tindouf Basin

Algeria

A member of Spiriferida. Genus includes new species B. brevedorsata.

Biernatia rhapsody[27]

Sp. nov

Valid

Lavié, Serra & Feltes

Middle and Late Ordovician

Las Aguaditas Formation
Las Chacritas Formation

Argentina

A member of Acrotretida belonging to the family Biernatidae.

Borellithyris[28]

Gen. et sp. nov

Valid

Dulai

Miocene (Tortonian)

Sant'Agata Fossili Formation

Italy

A member of Terebratulida belonging to the family Megathyrididae. The type species is B. gaetanii.

Cathaysiorthis xichuanensis[14]

Sp. nov

Valid

Huang et al.

Silurian (Rhuddanian)

Zhangwan Formation

China

|  |

Caucasorhynchella[16]

Gen. nov

Valid

Guo, Chen & Harper

Middle Triassic (Anisian)

|  |

China

|  |

Chonosteges cooperi[29]

Sp. nov

Valid

Torres-Martínez et al.

Permian

Paso Hondo Formation

Mexico

|  |

Cleiothyridina pfaffenbergensis[30]

Sp. nov

Valid

Mottequin et al.

Carboniferous (Tournaisian)

Gleitsch Formation

Germany

A member of Athyridida belonging to the family Athyrididae.

"Clorinda" symmetrica[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Conotreta andina[27]

Sp. nov

Valid

Lavié, Serra & Feltes

Ordovician (Darriwilian)

Las Aguaditas Formation
Las Chacritas Formation

Argentina

A member of Acrotretida belonging to the family Acrotretidae.

Coolinia diabolica[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Cordatia[32]

Gen. et sp. nov

Valid

Brock & Claybourn in Betts et al.

Early Cambrian

|  |

Australia

A member of Paterinata possibly belonging to the family Paterinidae. The type species is C. erinae.

Craniops popovi[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Crassispirifer galinae[33]

Sp. nov

Valid

Manankov

Permian

|  |

Mongolia

A member of Spiriferida.

Crassumbo germanicus[30]

Sp. nov

Valid

Mottequin et al.

Carboniferous (Tournaisian)

Gleitsch Formation

Germany

A member of Spiriferida belonging to the family Crassumbidae.

Cyrtia wrekinensis[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Cyrtina evanescens[26]

Sp. nov

Valid

Gourvennec

Paleozoic

Tindouf Basin

Algeria

A member of Spiriferinida.

Cyrtospirifer aouinetensis[26]

Sp. nov

Valid

Gourvennec

Paleozoic

Tindouf Basin

Algeria

|  |

Cyrtospirifer (Elasmospirifer)[26]

Subgen. et sp. nov

Valid

Gourvennec

Paleozoic

Tindouf Basin

Algeria

The subgenus includes new species C. (E.) djebiletensis.

Cyrtospirifer robardeti[26]

Sp. nov

Valid

Gourvennec

Paleozoic

Tindouf Basin

Algeria

|  |

Dictyothyropsis vogli[34]

Sp. nov

Valid

Bujtor & Vörös

Early Cretaceous

|  |

Hungary

|  |

Dihelictera askeriensis[20]

Sp. nov

Valid

Baarli

Ordovician (Hirnantian) and Silurian (Aeronian)

Solvik

Norway

A member of the family Atrypidae. Announced in 2019; the final version of the article naming was published in 2021.

Dinobolus athelstani[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Eleutherokomma djezairensis[26]

Sp. nov

Valid

Gourvennec

Paleozoic

Tindouf Basin

Algeria

A member of Spiriferida.

Eospirifer majesticus[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Erbotreta elankii[35]

Sp. nov

Valid

Ushatinskaya & Korovnikov

Cambrian

|  |

Russia

A member of Acrotretida.

Eucalathis giulioi[28]

Sp. nov

Valid

Dulai

Miocene (Tortonian)

Sant'Agata Fossili Formation

Italy

A member of Terebratulida belonging to the family Chlidonophoridae.

Farewellirhynchia[15]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

United States

A member of Rhynchonellida belonging to the subfamily Leiorhynchinae. The type species is F. kulkovi.

Fascistropheodonta? wiltzensis[36]

Sp. nov

Valid

Jansen

Early Devonian

Wiltz Formation

Germany
Luxembourg

A member of the family Strophodontidae.

Felsithyris[37]

Gen. et sp. nov

Valid

Mottequin & Weyer

Carboniferous (Mississippian)

|  |

Germany

A member of Spiriferida. The type species is F. hercynica.

Garciaalcaldia[15]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

United States

A member of Rhynchonellida belonging to the subfamily Glossinunilinae. The type species is G. alaskensis.

Ghorispirifer[38]

Gen. et comb. nov

Valid

Mottequin & Brice

Devonian (Famennian)

|  |

Afghanistan
Iran
Iraq

A Cyrtiopsinae. The type species is "Cyrtiopsis graciosa" chakhaensis Brice (1971) (raised to the rank of a separate species G. chakhaensis),
genus also includes "Cyrtiopsis" lapparenti Brice (1971).

Hedeinopsis africana[26]

Sp. nov

Valid

Gourvennec

Paleozoic

Tindouf Basin

Algeria

A member of Spiriferida.

Howellella minsterleyensis[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Ingentistrophia[36]

Gen. et comb. nov

Valid

Jansen

Early Devonian

Katzenelnbogen Formation Seifen Formation

Belgium
Germany

A member of Orthotetida belonging to the family Areostrophiidae. The type species is "Orthothethes" ingens Drevermann (1904).

Jonesea oepiki[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Jordanithyris[39]

Gen. et sp. nov

Valid

Feldman et al.

Middle Jurassic (Bathonian and Callovian)

Hamam
Mughanniyya

Jordan

A member of Terebratulida. Genus includes new species J. ardainensis. Announced in 2019; the final version of the article naming it was published in 2021.

Joviatrypa nakremi[20]

Sp. nov

Valid

Baarli

Silurian (Aeronian)

Solvik

Norway

A member of the family Atrypidae. Announced in 2019; the final version of the article naming was published in 2021.

Judinica[40]

Gen. et 2 sp. et comb. nov

Valid

Oleneva

Devonian

|  |

Belgium
Russia

A member of Spiriferida belonging to the family Adolfiidae. The type species is J. siratchoica; genus also includes new species J. pseudodeflexa, as well as J. pseudomultifida (Vandercammen, 1955), J. biverrucosa (Vandercammen, 1955) and J. solita (Ljaschenko, 1959).

Koeveskallina bifurcata[16]

Sp. nov

Valid

Guo, Chen & Harper

Middle Triassic (Anisian)

Qingyan Formation

China

|  |

Kosoidea australis[41]

Sp. nov

Valid

Zabini et al.

Ordovician (Hirnantian)

Iapó Formation
Vila Maria Formation

Brazil

A member of Discinoidea.

Kutchithyris simoni[42]

Sp. nov

Valid

Feldman et al.

Middle Jurassic (Callovian)

Mughanniyya

Jordan

Announced in 2019; the final version of the article naming it was published in 2021.

Lingulipora budili[43]

Sp. nov

Valid

Mergl

Middle Devonian

Srbsko Formation

Czech Republic

A member of the family Lingulidae.

Linnarssonia ulakhani[35]

Sp. nov

Valid

Ushatinskaya & Korovnikov

Cambrian

|  |

Russia

A member of Acrotretida.

Linoporella maxima[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Loboidothyris fordycei[21]

Sp. nov

Valid

MacFarlan

Early Jurassic (Lower Aratauran to Upper Ururoan)

|  |

New Caledonia
New Zealand

A member of Terebratulida belonging to the family Loboidothyrididae.

Lobothyris simesi[21]

Sp. nov

Valid

MacFarlan

Early Jurassic (Aratauran to Ururoan)

|  |

New Zealand

A member of Terebratulida belonging to the family Lobothyrididae.

Martinia georgei[44]

Sp. nov

Valid

Tazawa in Tazawa, Osawa & Nagura

Carboniferous (Viséan–Bashkirian)

Karaumedate Formation

Belgium
China
Japan
Kazakhstan
Russia
United Kingdom

A member of Spiriferida belonging to the family Martiniidae.

Martinia liuqiaoiensis[17]

Sp. nov

Valid

He, Shi & Shen in He et al.

Permian (Changhsingian)

Talung Formation

China

A member of Spiriferida belonging to the family Martiniidae.

Menarhynchus[15]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

United States

A member of Rhynchonellida belonging to the subfamily Sphaerirhynchinae. The type species is M. kuskokwimensis.

Micromartinia[45]

Gen. et sp. nov

Junior homonym

He & Weldon in He et al.

Permian (Changhsingian)

Talung Formation

China

A member of Spiriferida belonging to the superfamily Martinioidea. The type species is M. kwangsiella. The generic name is preoccupied by Micromartinia Amsel (1957).

Minutomarginifera[45]

Nom. nov

Valid

He et al.

Carboniferous and Permian

|  |

China
Thailand

A member of Strophalosiidina belonging to the family Chonopectidae; a replacement name for Eileenella Racheboeuf in Wongwanich et al. (2004).

Minutomena missa[46]

Sp. nov

Valid

Huang et al.

Ordovician (Hirnantian)

Kuanyinchiao Beds

China

|  |

Multispirifer kropotkini[26]

Sp. nov

Valid

Gourvennec

Paleozoic

Tindouf Basin

Algeria

A member of Spiriferida.

Neocyrtina xui[16]

Sp. nov

Valid

Guo, Chen & Harper

Middle Triassic (Anisian)

Qingyan Formation

China

|  |

Neoliothyrina nakremi[47]

Sp. nov

Valid

Bitner in Hryniewicz et al.

Late Paleocene

Basilika Formation

Norway

A member of Terebratulida belonging to the family Sellithyridae.

Neoplatystrophia paludis[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Neospirifer borealis[33]

Sp. nov

Valid

Manankov

Permian

|  |

Mongolia

A member of Spiriferida.

Nisusia multicostata[25]

Sp. nov

Valid

Holmer et al.

Cambrian (Drumian)

Mila Formation

Iran

A member of Kutorginida.

Nottina[20]

Gen. et sp. nov

Valid

Baarli

Silurian (Rhuddanian and Aeronian)

Solvik

Norway

A member of the family Atrypidae. The type species is N. phalerata. Announced in 2019; the final version of the article naming was published in 2021.

Nucleospira elgari[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Nudirostralina minuta[16]

Sp. nov

Valid

Guo, Chen & Harper

Middle Triassic (Anisian)

Qingyan Formation

China

|  |

Opsiconidion nanus[43]

Sp. nov

Valid

Mergl

Devonian (Eifelian)

Acanthopyge Limestone

Czech Republic

A member of Acrotretoidea belonging to the family Biernatidae.

Oretanomena[48]

Gen. et sp. nov.

Valid

Reyes-Abril, Villas & Gutierrez-Marco

Ordovician (Darriwilian)

|  |

Spain

A member of Strophomenoidea. Genus includes new species O. meloui.

Ortarhynchia[24]

Gen. et sp. nov

Valid

Gaetani in Grădinaru & Gaetani

Triassic

|  |

Romania

A member of Rhynchonellida belonging to the family Norellidae and the subfamily Paranorellininae. The type species is O. petersi.

Pachyschizophoria amygdalina[36]

Sp. nov

Valid

Jansen

Early Devonian

Berlé Formation
Emsquarzit Formation
Hohenrhein Formation

Germany
Luxembourg

A member of Enteletoidea belonging to the family Schizophoriidae.

Parabrekia[16]

Gen. et sp. nov

Valid

Guo, Chen & Harper

Middle Triassic (Anisian)

Qingyan Formation

China

Genus includes new species P. yangi.

Paurorthis? llangynogensis[18]

Sp. nov

Valid

Cocks & Popov

Early Ordovician

|  |

United Kingdom

|  |

Pentameroides johnsoni[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Permorhipidomella[17]

Gen. et sp. nov

Valid

He, Shi & Shen in He et al.

Permian (Changhsingian)

|  |

China

A member of Orthida belonging to the family Rhipidomellidae. Genus includes new species P. ovatus.

Piarorhynchella kittli[24]

Sp. nov

Valid

Gaetani in Grădinaru & Gaetani

Triassic

|  |

Romania

A member of Rhynchonellida belonging to the family Norellidae and the subfamily Holcorhynchellinae.

Platytrochalos (Salopicoelia) morrellswoodensis[31]

Subgen. et sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Productina saalfeldensis[30]

Sp. nov

Valid

Mottequin et al.

Carboniferous (Tournaisian)

Gleitsch Formation
Hangenberg Limestone

Germany

A member of Productida belonging to the family Productellidae.

Ptychomentzelia dobrogeana[24]

Sp. nov

Valid

Gaetani in Grădinaru & Gaetani

Triassic

|  |

Romania

A member of Spiriferida belonging to the family Mentzeliidae.

Ptychomentzelia simionescui[24]

Sp. nov

Valid

Gaetani in Grădinaru & Gaetani

Triassic

|  |

Romania

A member of Spiriferida belonging to the family Mentzeliidae.

Quiringites bitami[26]

Sp. nov

Valid

Gourvennec

Paleozoic

Tindouf Basin

Algeria

A member of Spiriferida.

Rhipidomella parvula[17]

Sp. nov

Valid

He, Shi & Shen in He et al.

Permian (Changhsingian)

|  |

China

A member of Orthida belonging to the family Rhipidomellidae.

Roemerithyris[37]

Gen. et comb. nov

Valid

Mottequin & Weyer

Carboniferous (Mississippian)

|  |

Germany

A member of Spiriferida. The type species is "Spirifer" macrogaster Roemer (1852).

Rugithyris hasibuani[21]

Sp. nov

Valid

MacFarlan

Early Jurassic (Upper Ururoan)

|  |

New Zealand

A member of Terebratulida belonging to the superfamily Loboidothyridoidea.

Rutorhynchia? trigonalis[16]

Sp. nov

Valid

Guo, Chen & Harper

Middle Triassic (Anisian)

Qingyan Formation

China

|  |

Schachriomonia spiraensis[20]

Sp. nov

Valid

Baarli

Ordovician-Silurian

Solvik

Norway

A member of the family Atrypidae. Announced in 2019; the final version of the article naming was published in 2021.

Schellwienella justinianoi[49]

Sp. nov

Valid

Rezende, Ponciano & Brett

Late Devonian

Longá Formation

Brazil

|  |

Sellithyris elizabetha[50]

Sp. nov

Valid

Rojas & Sandy

Early Cretaceous (Valanginian)

Rosablanca Formation

Colombia

|  |

Septatrypa salopiensis[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Shelvothyris[31]

Gen. et comb. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

Genus includes S. furcata (Sowerby, 1839)

Sifella[20]

Gen. et sp. nov

Valid

Baarli

Silurian (Aeronian)

Solvik

Norway

A member of the family Atrypidae. The type species is S. patera. Announced in 2019; the final version of the article naming was published in 2021.

Skenidoides pontyfennensis[18]

Sp. nov

Valid

Cocks & Popov

Early Ordovician

|  |

United Kingdom

|  |

Smirnovina ferraria[34]

Sp. nov

Valid

Bujtor & Vörös

Early Cretaceous

|  |

Hungary

|  |

Spiriferella protodraschei[19]

Sp. nov

Valid

Lee & Shi in Lee et al.

Permian (late Artinskian–early Kungurian)

Kapp Starostin Formation

Norway

A member of Spiriferida belonging to the family Spiriferellidae.

Tamarirhynchia[15]

Gen. et comb. nov

Valid

Baranov & Blodgett

Devonian (Pragian)

Soda Creek Limestone

United States

A member of Rhynchonellida belonging to the subfamily Hebetoechiinae. The type species is "Lancemyonia" varia Tcherkesova (1969).

Tegulithyris? plencnerae[21]

Sp. nov

Valid

MacFarlan

Early Jurassic (Upper Aratauran and Ururoan)

|  |

New Zealand

A member of Terebratulida belonging to the family Tegulithyrididae.

Thebesia shelvensis[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Thuringorhynchus[30]

Gen. et sp. nov

Valid

Mottequin et al.

Carboniferous (Tournaisian)

Gleitsch Formation
Hangenberg Limestone

Germany

A member of Rhynchonellida belonging to the family Pugnacidae. The type species is T. pseudoequitans.

Tornquistia changhsingia[17]

Sp. nov

Valid

He, Shi & Shen in He et al.

Permian (Changhsingian)

|  |

China

A member of Chonetidina belonging to the family Anopliidae and the subfamily Anopliinae.

Trigonithyris wilsoni[42]

Sp. nov

Valid

Feldman et al.

Middle Jurassic (Callovian)

Mughanniyya

Jordan

Announced in 2019; the final version of the article naming it was published in 2021.

Triplesia habberleyensis[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Tritoechia bolohaulensis[18]

Sp. nov

Valid

Cocks & Popov

Early Ordovician

|  |

United Kingdom

|  |

Tumarinia uldsiensis[33]

Sp. nov

Valid

Manankov

Permian

|  |

Mongolia

A member of Spiriferinida.

Ukhtomica[40]

Gen. et sp. nov

Valid

Oleneva

Devonian

|  |

Russia

A member of Spiriferida belonging to the family Echinospiriferidae. The type species is U. lata; genus also includes U. apschakensis (Krylova, 1955).

Virgianoides[51]

Gen. et comb. nov

Valid

Jin, Mikulic & Kluessendorf

Silurian (Rhuddanian)

Mayville Dolomite

United States
( Wisconsin)

A member of Pentamerida belonging to the family Virgianidae. The type species is BVirgiana barrandei var. major Savage (1916), raised to the rank of a separate species V. major.

Weiningia ziyunensis[52]

Sp. nov

Valid

Yuan et al.

Carboniferous (Serpukhovian)

Baizuo Formation

China

A member of Spiriferida belonging to the family Martiniidae.

Whitfieldella alvarezensis[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

Xenocrania[53]

Gen. et comb. nov

Valid

Chen & Rong

Ordovician

|  |

China
Myanmar
Poland
United Kingdom
Czech Republic?

The type species is "Palaeocyclus" haimei Reed.

Zeilleria recessa[21]

Sp. nov

Valid

MacFarlan

Early Jurassic (Hettangian)

|  |

New Zealand

A member of Terebratulida belonging to the family Zeilleriidae.

Zeilleria sacciformis[21]

Sp. nov

Valid

MacFarlan

Early Jurassic (Ururoan)

Ururoa Formation

New Zealand

A member of Terebratulida belonging to the family Zeilleriidae.

Zeilleria terezowae[21]

Sp. nov

Valid

MacFarlan

Early Jurassic (Aratauran and Ururoan)

Ururoa Formation

New Zealand

A member of Terebratulida belonging to the family Zeilleriidae.

Zittelina hofmanni[34]

Sp. nov

Valid

Bujtor & Vörös

Early Cretaceous

|  |

Hungary

|  |

Zygospiraella nupera[20]

Sp. nov

Valid

Baarli

Silurian (Aeronian)

Solvik

Norway

A member of the family Atrypidae. Announced in 2019; the final version of the article naming was published in 2021.

Zygospiraella? ventricosa[31]

Sp. nov

Valid

Cocks

Silurian (Llandovery)

|  |

United Kingdom

|  |

## References

[edit]

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25. ^ a b Lars E. Holmer; Mohammad-Reza Kebria-ee Zadeh; Leonid E. Popov; Mansoureh Ghobadi Pour; J. Javier Álvaro; Vachik Hairapetian; Zhifei Zhang (2019). "Cambrian rhynchonelliform nisusioid brachiopods: phylogeny and distribution". Papers in Palaeontology. 5 (3): 559–575\. Bibcode:2019PPal....5..559H. doi:10.1002/spp2.1255. hdl:10261/187807.559-575&rft.date=2019&rft_id=info:hdl/10261/187807&rft_id=info:doi/10.1002/spp2.1255&rft_id=info:bibcode/2019PPal....5..559H&rft.au=Lars+E.+Holmer&rft.au=Mohammad-Reza+Kebria-ee+Zadeh&rft.au=Leonid+E.+Popov&rft.au=Mansoureh+Ghobadi+Pour&rft.au=J.+Javier+Álvaro&rft.au=Vachik+Hairapetian&rft.au=Zhifei+Zhang&rft_id=https://doi.org/10.1002%2Fspp2.1255&rfr_id=info:sid/en.wikipedia.org:2019+in+brachiopod+paleontology" class="Z3988">
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