2020 in paleobotany

Overview of the events of 2020 in paleobotany

List of years in paleobotany

* …
* 2017
* 2018
* 2019
* 2020
* 2021
* 2022
* 2023
* …

In paleontology
2017
2018
2019
2020
2021
2022
2023

In arthropod paleontology
2017
2018
2019
2020
2021
2022
2023

In paleoentomology
2017
2018
2019
2020
2021
2022
2023

In paleomalacology
2017
2018
2019
2020
2021
2022
2023

In reptile paleontology
2017
2018
2019
2020
2021
2022
2023

In archosaur paleontology
2017
2018
2019
2020
2021
2022
2023

In mammal paleontology
2017
2018
2019
2020
2021
2022
2023

In paleoichthyology
2017
2018
2019
2020
2021
2022
2023

* Art
* Archaeology
* Architecture
* Literature
* Music
* Philosophy
* Science +...

This article records new taxa of fossil plants described during the year 2020, as well as other significant discoveries and events related to paleobotany that occurred in 2020.

## Flowering plants

[edit]

### Alismatales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Limnobiophyllum pedunculatum[1]

Sp. nov

Valid

Low, Su & Xing in Low et al.

Late Oligocene

|  |

China

A member of the family Araceae.

### Apiales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Paleopanax puryearensis[2]

Sp. nov

Valid

Na, Blanchard & Wang

Middle Eocene

Cockfield Formation

United States
( Tennessee)

A member of the family Araliaceae.

### Arecales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Echimonocolpites chicxulubensis[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

|  |

Mexico

Pollen of a flowering plant, probably a member of the family Arecaceae.

Palmoxylon ceroxyloides[4]

Sp. nov

In press

Khan, Hazra & Bera in Khan et al.

Late Cretaceous (Maastrichtian)-Paleocene (Danian)

Deccan Intertrappean Beds

India

A petrified palm stem of a member of the subfamily Ceroxyloideae.

Palmoxylon dindoriensis[5]

Sp. nov

Valid

Khan, Roy & Bera in Khan et al.

Late Cretaceous (Maastrichtian)-Paleocene (Danian)

Deccan Intertrappean Beds

India

A petrified palm stem.

Sabalites dawsonii[6]

Sp. nov

Valid

Greenwood & Conran

Eocene

|  |

Canada
( British Columbia)

|  |

Sabalites karondiensis[7]

Sp. nov

In press

Roy, Hazra & Khan in Roy et al.

Late Cretaceous-Paleocene (latest Maastrichtian-earliest Danian)

Deccan Intertrappean Beds

India

A palm frond .

Spinizonocolpites riochiquensis[8]

Sp. nov

Valid

Vallati & De Sosa Tomas in Vallati, De Sosa Tomas & Casal

Late Cretaceous (Maastrichtian)

Lago Colhué Huapí Formation

Argentina

A member of Arecaceae described on the basis of fossil pollen grains. Announced in 2019; the final version of the article naming it was published in 2020.

### Buxales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Pachysandra europaea[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

|  |

Germany

A species of Pachysandra. Announced in 2019; the final version of the article naming it was published in 2020.

### Caryophyllales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Gomphrenipollis garciae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant, possibly produced by members of the family Amaranthaceae. Announced in 2020; the final version of the article naming it was published in 2021.

### Chloranthales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Sarcandraxylon[11]

Gen. et sp. nov

Valid

Pipo, Iglesias & Bodnar

Late Cretaceous (early–middle Campanian)

Santa Marta Formation

Antarctica

A member of the family Chloranthaceae. Genus includes new species S. sanjosense.

### Cornales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Amersinia littletonensis[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

United States
( Colorado)

|  |

Langtonia parva[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

United States
( Colorado)

A member of the family Mastixiaceae.

Mastixicarpum hoodii[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

United States
( Colorado)

A member of the family Mastixiaceae.

Nyssa gergoei[13]

Sp. nov

Valid

Hably

Miocene

|  |

Hungary

A tupelo.

Nyssa gyoergyi[13]

Sp. nov

Valid

Hably

Miocene

|  |

Hungary

A tupelo.

Platycrater iljinskajae[14]

Sp. nov

In press

Denk et al.

Late Oligocene

|  |

Russia

|  |

Portnallia alexanderi[12]

Sp. nov

Valid

Huegele & Manchester

Early Paleocene

Denver Formation

United States
( Colorado)

A member of the family Mastixiaceae.

### Crossosomatales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Staphylea woodworthensis[15] | Sp. nov  | In press  | Zhu & Manchester  | Oligocene | Renova Formation |  United States
( Montana)  | A species of Staphylea.  |

### Cucurbitales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Coriaripites goodii[16]

Sp. nov

Valid

Barreda, Palazzesi & Tellería in Renner et al.

Late Cretaceous (Campanian–early Maastrichtian)

Lopez de Bertodano Formation
Santa Marta Formation
Snow Hill Island Formation

Antarctica

Pollen grains similar to those of extant members of the genus Coriaria.

Echitriporites jolyi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen produced by members of the genus Cayaponia. Announced in 2020; the final version of the article naming it was published in 2021.

### Ericales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Andrewsiocarpon puryearensis[2]

Sp. nov

Valid

Na, Blanchard & Wang

Middle Eocene

Cockfield Formation

United States
( Tennessee)

A member of the family Theaceae.

Anubiscarpon[17]

Gen. et sp. nov

Valid

Smith & Manchester

Middle Eocene

Clarno Formation

United States
( Oregon)

A member of the family Theaceae. Genus includes new species A. andersonae.

### Fabales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Cercis zekuensis[18]

Sp. nov

Valid

Li et al.

Early Miocene

|  |

China

A species of Cercis. Announced in 2020; the final version of the article naming it was published in 2021.

Gleditsia pliocaenica[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

|  |

Germany

A species of Gleditsia. Announced in 2019; the final version of the article naming it was published in 2020.

Menendoxylon lutzi[19] | Sp. nov  | In press  | Baez & Crisafulli  | Miocene | Chiquimil |  Argentina | Fossil wood of a member of the family Fabaceae.  |

Parkiidites marileae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen produced by members of the genus Parkia. Announced in 2020; the final version of the article naming it was published in 2021.

Prioria martineziorum[20]

Sp. nov

Valid

Rodríguez-Reyes & Estrada-Ruiz

Oligocene-Miocene

Santiago Formation

Panama

A species of Prioria.

Psilastephanocolporites deoliverae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen produced by members of the family Polygalaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Psilastephanocolporites endoporatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Striatopollis grahamii[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

|  |

Mexico

Pollen of an eudicot, probably a member of the family Fabaceae.

### Fagales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Alnus chaybulakensis[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

|  |

Kazakhstan

An alder.

Berryophyllum hainanensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

China

A member of the family Fagaceae.

Carpinus asymmetrica[23]

Sp. nov

In press

Xue & Jia in Xue et al.

Early Miocene

| Maguan |

China

A species of Carpinus.

Carpinus symmetrica[23] |

Sp. nov

In press

Xue & Jia in Xue et al.

Early Miocene

Maguan Basin

China

A species of Carpinus

Carya pipecreekensis[24]

Sp. nov

In press

Swinehart & Farlow

Late Neogene

Pipe Creek Sinkhole

United States
( Indiana)

A hickory.

Castaneophyllum hainanensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

China

A member of the family Fagaceae.

Castaneophyllum lanceolata[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

China

A member of the family Fagaceae.

Castanopsis bulgarica[25] | Sp. nov  | Valid  | Mantzouka, Ivanov & Bozukov  | Late Miocene–early Pliocene (late Messinian–early Zanclean)  | Pokrovnik |  Bulgaria | A species of Castanopsis. Announced in 2020; the final version of the article naming it was published in 2021.  |

Lithocarpus changchangensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

China

A species of Lithocarpus.

Quercus aimeeana[26]

Sp. nov

Valid

Sadowski, Schmidt & Denk

Eocene

Baltic amber

Europe (Baltic Sea region)

An oak.

Quercus borissovii[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

|  |

Kazakhstan

An oak.

Quercus campanulata[26]

Sp. nov

Valid

Sadowski, Schmidt & Denk

Eocene

Baltic amber

Europe (Baltic Sea region)

An oak.

Quercus casparyi[26]

Sp. nov

Valid

Sadowski, Schmidt & Denk

Eocene

Baltic amber

Europe (Baltic Sea region)

An oak.

Quercus changchangensis[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

China

An oak.

Quercus conwentzii[26]

Sp. nov

Valid

Sadowski, Schmidt & Denk

Eocene

Baltic amber

Europe (Baltic Sea region)

An oak.

Quercus longianthera[26]

Sp. nov

Valid

Sadowski, Schmidt & Denk

Eocene

Baltic amber

Europe (Baltic Sea region)

An oak.

Quercus multipilosa[26]

Sp. nov

Valid

Sadowski, Schmidt & Denk

Eocene

Baltic amber

Europe (Baltic Sea region)

An oak.

Quercus paleoargyrotricha[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

China

An oak.

Quercus paleohypargyrea[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

China

An oak.

Quercus paleolamellosa[22]

Sp. nov

Valid

Liu & Jin in Liu, Song & Jin

Eocene

Changchang Formation

China

An oak.

### Garryales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Eucommia szaferi[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

|  |

Germany

A species of Eucommia. Announced in 2019; the final version of the article naming it was published in 2020.

### Gentianales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Asclepiadospermum[27]

Gen. et 2 sp. nov

Valid

Del Rio et al.

Early Eocene

Niubao Formation

China

An asclepiadoid Apocynaceae genus.
Included species A. marginatum and A. ellipticum.

Margocolporites carinae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen produced by members of the genus Rauvolfia. Announced in 2020; the final version of the article naming it was published in 2021.

### Icacinales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Iodes elliptica[28]

Sp. nov

In press

Del Rio et al.

Early Oligocene

Wenshan Basin

China

A member of the family Icacinaceae.

Iodes passiciensis[29]

Sp. nov

Valid

Del Rio & De Franceschi

Early Eocene

|  |

France

A member of the family Icacinaceae.

Manchesteria[30]

Gen. et sp. nov

In press

Stull & Rozefelds in Rozefelds et al.

Cenozoic (mid-Miocene or, more likely, middle Eocene)

|  |

Australia

A member of the family Icacinaceae. Genus includes new species M. australis.

Pyrenacantha simonsii[31]

Sp. nov

Valid

Stull et al.

Early Oligocene

|  |

Egypt

A species of Pyrenacantha.

### Lamiales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Echitricolpites cruziae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen produced by members of the genus Aegiphila. Announced in 2020; the final version of the article naming it was published in 2021.

Multiareolites? reticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen produced by members of the family Acanthaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Retistephanocolpites curvimuratus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Retistephanocolpites pardoi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant, possibly produced by members of the genus Amphilophium. Announced in 2020; the final version of the article naming it was published in 2021.

### Laurales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Actinodaphnoxylon[32]

Gen. et sp. nov

In press

Akkemik et al.

Eocene (Lutetian)

|  |

Turkey

A member of the family Lauraceae. Genus includes new species A. zileensis.

Mezilaurinoxylon oleiferum[33]

Sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

Argentina

A member of the family Lauraceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Patagonoxylon[33]

Gen. et sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

Argentina

A member of Laurales of uncertain phylogenetic placement. Genus includes new species P. scalariforme. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Thymolepis[34]

Gen. et sp. nov

Valid

Chambers & Poinar

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

Possibly an early representative of Monimiaceae. Genus includes new species T. toxandra.

Valviloculus[35]

Gen. et sp. nov

Valid

Poinar et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

Possibly a member of Laurales related to the families Monimiaceae and Atherospermataceae. Genus includes new species V. pleristaminis.

### Liliales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Smilax fujianensis[36]

Sp. nov

Valid

Dong et al.

Middle Miocene

|  |

China

A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.

Smilax zhangpuensis[36]

Sp. nov

Valid

Dong et al.

Middle Miocene

|  |

China

A species of Smilax. Announced in 2020; the final version of the article naming it was published in 2021.

### Magnoliales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Magnolia nanningensis[37]

Sp. nov

Valid

Huang et al.

Late Oligocene

Nanning Basin

China

A species of Magnolia.

Magnolia waltheri[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

|  |

Germany

A species of Magnolia. Announced in 2019; the final version of the article naming it was published in 2020.

Melloniflora[38]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous

Potomac Group

United States
( Virginia)

A relative of extant early-diverging members of the Magnoliales. Genus includes new species M. virginiensis.

### Malpighiales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Salix palaeofutura[39]

Sp. nov

Valid

Narita et al.

Miocene

Bifuka Formation

Japan

A willow.

### Malvales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Bastardioxylon[19] | Gen. et sp. nov  | In press  | Baez & Crisafulli  | Miocene | Chiquimil |  Argentina | Fossil wood of a member of the family Malvaceae. Genus includes new species B. antiqua.  |
Dipterocarpus dindoriensis[40] | Sp. nov  | Valid  | Khan, Spicer & Bera in Khan et al. | Late Cretaceous (Maastrichtian)  | Deccan Intertrappean Beds |  India | A species of Dipterocarpus.  |

Echiperiporites germeraadii[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

Echiperiporites jaramilloi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen produced by members of the genus Hibiscus. Announced in 2020; the final version of the article naming it was published in 2021.

Echiperiporites titanicus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen produced by members of the genus Malachra. Announced in 2020; the final version of the article naming it was published in 2021.

Retistephanocolporites elizabeteae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen produced by members of the genus Ceiba. Announced in 2020; the final version of the article naming it was published in 2021.

Veraguasoxylon[20]

Gen. et sp. nov

Valid

Rodríguez-Reyes & Estrada-Ruiz

Oligocene-Miocene

Santiago Formation

Panama

A member of the family Malvaceae. Genus includes new species V. panamense.

### Myrtales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Eucalyptus xoshemium[41]

Sp. nov

Valid

Gandolfo & Zamaloa in Zamaloa, Gandolfo & Nixon

Eocene (Ypresian)

Huitrera Formation

Argentina

A species of Eucalyptus.

Mangroveoxylon[42]

Gen. et comb. nov

In press

Moya & Brea

Late Miocene?

Ituzaingó Formation

Argentina

A member of the family Combretaceae; a new genus for "Menendoxylon" areniensis Lutz (1979).

Myrceugenellites grandiporosum[33]

Sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

Argentina

A member of the family Myrtaceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Primotrapa[43]

Gen. et sp. nov

Valid

Li & Li in Li et al.

Late Eocene to early Miocene

Hannuoba

China
Czech Republic
France
Germany

A member of Trapoideae. Genus includes new species P. weichangensis, as well as "Carpolithus" pomelii Saporta (1878) and "Hemitrapa" alpina Su & Zhou in Su et al. (2018).

### Nymphaeales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Praenymphaeapollenites[44]

Gen. et sp. nov

Valid

Barrón, Peris & Labandeira in Peris et al.

Cenomanian

Burmese amber

Myanmar

Pollen of a member of Nymphaeaceae.
Genus includes new species P. cenomaniensis.

### Oxalidales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Cunoniantha[45]

Gen. et sp. nov

Valid

Jud & Gandolfo

Paleocene (early Danian)

Salamanca Formation

Argentina

A member of the family Cunoniaceae. Genus includes new species C. bicarpellata. Announced in 2020; the final version of the article naming it was published in 2021.

Elaeocarpus nanningensis[46]

Sp. nov

In press

Liu et al.

Late Oligocene

Yongning Formation

China

A species of Elaeocarpus.

Elaeocarpus prelacunosus[46]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

China

A species of Elaeocarpus.

Elaeocarpus preprunifolioides[46]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

China

A species of Elaeocarpus.

Elaeocarpus prerugosus[46]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

China

A species of Elaeocarpus.

Elaeocarpus preserratus[46]

Sp. nov

In press

Liu et al.

Late Miocene

Foluo Formation

China

A species of Elaeocarpus.

Elaeocarpus presikkimensis[46]

Sp. nov

In press

Liu et al.

Miocene

Erzitang Formation

China

A species of Elaeocarpus.

### Poales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Monoporopollenites scabratus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen produced by members of the family Poaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Rhizomatites[47]

Gen. et sp. nov

In press

Robledo & Anzótegui in Robledo et al.

Miocene-Pliocene

|  |

Argentina

A member of Cyperaceae. Genus includes new species R. cyperoides.

### Proteales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Banksia microphylla[48]

Sp. nov

Valid

Carpenter in Carpenter & Milne

Late Eocene

|  |

Australia

A species of Banksia.

Banksieaeidites zanthus[48]

Sp. nov

Valid

Milne in Carpenter & Milne

Late Eocene

|  |

Australia

A Banksia-like pollen.

Platanus emryi[49]

Sp. nov

Valid

Huegele, Spielbauer & Manchester

Miocene

|  |

United States

A species of Platanus.

### Ranunculales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Cissampelos defranceschii[50]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

China

A species of Cissampelos. Announced in 2020; the final version of the article naming it was published in 2021.

Clematis csabae[13]

Sp. nov

Valid

Hably

Miocene

|  |

Hungary

A species of Clematis.

Diploclisia praeaffinis[51]

Sp. nov

Valid

Jia et al.

Late Miocene

|  |

China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Menispermites bangorensis[50]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Menispermites haominae[50]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Menispermites tibetica[50]

Sp. nov

Valid

Huang in Del Rio et al.

Middle Eocene

Niubao Formation

China

A member of the family Menispermaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Paleoorbicarpum[52]

Gen. et sp. nov

Valid

Han et al.

Paleocene

Sanshui Basin

China

A member of the family Menispermaceae. Genus includes new species P. parvum.

Stephania bangorensis[50]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

China

A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.

Stephania geniculata[52]

Sp. nov

Valid

Han et al.

Paleocene

Sanshui Basin

China

A species of Stephania

Stephania ornamenta[52]

Sp. nov

Valid

Han et al.

Paleocene

Sanshui Basin

China

A species of Stephania

Stephania shuangxingii[50]

Sp. nov

Valid

Del Rio & Su in Del Rio et al.

Middle Eocene

Niubao Formation

China

A species of Stephania. Announced in 2020; the final version of the article naming it was published in 2021.

### Rosales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Berhamniphyllum junrongii[53]

Sp. nov

Valid

Zhou, Wang & Huang in Zhou et al.

Late Eocene

Markam Basin

China

A member of the family Rhamnaceae

Crataegus pentagynoides[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

|  |

Germany

A species of Crataegus. Announced in 2019; the final version of the article naming it was published in 2020.

Hemiptelea kryshtofovichii[21]

Sp. nov

Valid

Averyanova & Xing

Paleogene

|  |

Kazakhstan

A member of the family Ulmaceae.

Scabrastephanoporites[3]

Gen. et sp. nov

Valid

Smith et al.

Eocene (Ypresian)

|  |

Mexico

Pollen of an eudicot, probably a member of the family Ulmaceae or Cannabaceae. Genus includes new species S. variabilis.

### Sapindales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Acer dombeyopsis[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

|  |

Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

Acer viburnoides[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

|  |

Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

Acer vitiforme[9]

Sp. nov

Valid

Kvaček, Teodoridis & Denk

Pliocene

|  |

Germany

A maple. Announced in 2019; the final version of the article naming it was published in 2020.

Brosipollis reticulatus[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

|  |

Mexico

Pollen of a flowering plant, probably a member of the family Burseraceae.

Choerospondias fujianensis[54]

Sp. nov

In press

Wang et al.

Miocene

|  |

China

A species of Choerospondias.

Llanodelacruzoxylon[55] | Gen. et sp. nov  |  | Rodríguez-Reyes, Estrada-Ruiz & Gasson  | Oligocene–Miocene  | Santiago Formation |  Panama | A member of the family Anacardiaceae. Genus includes new species L. sandovalii.  |

Manchestercarpa[56]

Gen. et sp. nov

Valid

Atkinson

Late Cretaceous (Campanian)

|  |

Canada
( British Columbia)

A member of the family Meliaceae described on the basis of a fossil fruit. Genus includes new species M. vancouverensis.

Parametopioxylon[57]

Gen. et sp. nov

Valid

Franco et al.

Miocene

Ituzaingó Formation

Argentina

A member of the family Anacardiaceae described on the basis of fossil wood. Genus includes new species P. crystalliferum.

Psilastephanocolporites hammenii[3]

Sp. nov

Valid

Smith et al.

Eocene (Ypresian)

|  |

Mexico

Pollen of a flowering plant, probably a member of the family Meliaceae

Quinquala[58]

Gen. et sp. nov

Valid

Manchester & Disney in Manchester, Disney & Pham

Eocene

Clarno Formation
Tepee Trail Formation

United States
( Oregon
Wyoming)

A fossil fruit with affinities with the Rutaceae. Genus includes new species Q. obovata.

Rousea cavitata[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Announced in 2020; the final version of the article naming it was published in 2021.

### Saxifragales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Corylopsis grisea[59]

Sp. nov

Valid

Quirk & Hermsen

Early Pliocene

Gray Fossil Site

United States
( Tennessee)

A species of Corylopsis. Announced in 2020; the final version of the article naming it was published in 2021.

Itea polyneura[60]

Sp. nov

In press

Huang & Tian in Tian et al.

Oligocene

Huazhige Formation

China

A species of Itea.

Protoaltingia[61]

Gen. et sp. nov

Valid

Scharfstein, Stockey & Rothwell

Late Cretaceous (Coniacian)

|  |

Canada
( British Columbia)

A member of the family Altingiaceae. Genus includes new species P. comoxense.

### Solanales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Physalis hunickenii[62]

Sp. nov

Valid

Deanna, Wilf & Gandolfo

Early Eocene

Laguna del Hunco Formation

Argentina

A species of Physalis.

### Trochodendrales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
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|

Eotrochion[63]

Gen. et sp. nov

Valid

Manchester, Kvaček & Judd

Paleocene

|  |

United States
( Wyoming)

A member of the family Trochodendraceae. Genus includes new species E. polystylum. Announced in 2020; the final version of the article naming it was published in 2021.

Paraconcavistylon[63]

Gen. et comb. nov

Valid

Manchester, Kvaček & Judd

Ypresian

Klondike Mountain Formation

United States
( Washington)

A Trochodendraceae; a new genus for "Concavistylon" wehrii Manchester et al. (2018). Announced in 2020; the final version of the article naming it was published in 2021.

Trochodendron infernense[63]

Sp. nov

Valid

Manchester, Kvaček & Judd

Paleocene

|  |

United States
( Wyoming)

A species of Trochodendron. Announced in 2020; the final version of the article naming it was published in 2021.

### Vitales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
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|
|

Yua jiangxiensis[64]

Sp. nov

Valid

He & Wang

Miocene

Toupi Formation

China

A species of Yua. Announced in 2020; the final version of the article naming it was published in 2021.

### Other angiosperms

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
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|
|

Aextoxicoxylon kawasianus[65]

Sp. nov

Valid

Vera et al.

Late Cretaceous

Puntudo Chico Formation

Argentina

A fossil dicot wood

Atlantocarpus[66]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

Portugal
United States
( Virginia)

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species A. virginiensis.

Carpolithes gergoei[13]

Sp. nov

Valid

Hably & Erdei in Hably

Miocene

|  |

Hungary

A fossil fruit of a flowering plant of uncertain phylogenetic placement.

Catanthus[67]

Gen. et sp. nov

In press

Friis, Crane & Pedersen

Early Cretaceous

|  |

Portugal

An early flowering plant. Genus includes new species C. dolichostemon.

Cavilignum[68]

Gen. et sp. nov

Valid

Siegert & Hermsen

Early Pliocene

Gray Fossil Site

United States
( Tennessee)

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil endocarps. Genus includes new species C. pratchettii.

Chainandra[69]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

Genus includes new species C. zeugostylus.

Cichoreacidites? flammulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Crotonoidaepollenites echinatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Cyathitepala[70]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

Genus includes new species C. papillosa.

Dasykothon[71]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A flowering plant of uncertain phylogenetic placement, possibly a member of Laurales. Genus includes new species D. leptomiscus.

Dinganthus[72]

Gen. et sp. nov

Valid

Liu et al.

Miocene

Dominican amber

Dominican Republic

A eudicot of uncertain phylogenetic placement. Genus includes new species D. pentamera.

Echistephanoporites annulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Eofructus[73]

Gen. et sp. nov

Valid

Han & Wang

Early Cretaceous

Yixian Formation

China

An infructescence including a central axis and five fruits resembling Liaoningfructus. Genus includes new species E. liutiaogouensis.

Foveotricolporites crassus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Inaperturopollenites microechinatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Ladakhipollenites carmoi[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Lambertiflora[66]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

United States
( Virginia)

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species L. elegans.

Malvacipolloides diversus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Malvacipolloides echibaculatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Malvacipolloides romeroae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Menatanthus[74]

Gen. et sp. nov

Valid

Uhl, Paudayal & El Atfy in Uhl et al.

Paleocene (Thanetian)

|  |

France

A eudicot of uncertain phylogenetic placement. Genus includes new species M mosbruggeri. Announced in 2020; the final version of the article naming it was published in 2021.

Mugideiriflora[66]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Aptian-early Albian)

Almargem Formation

Portugal

An early flowering plant, possibly related to the group Austrobaileyales. Genus includes new species M. portugallica.

Phantophlebia[75]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A flowering plant of uncertain phylogenetic placement, possibly related to myrsinoid members of the family Primulaceae. Genus includes new species P. dicycla.

Platanites willigeri[76]

Sp. nov

Valid

Halamski & Kvaček in Halamski et al.

Late Cretaceous (Santonian)

Czerna Formation

Poland

Trifoliolate platanoid leaves.

Psilaperiporites delicatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Psilaperiporites lunaris[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Ranunculacidites reticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Rasenganus[77]

Gen. et sp. nov

Valid

Xing & Gu

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A possible epizoochorous fruit. Genus includes new species R. auricularus.

Retibrevitricolpites microreticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retibrevitricolporites costaporus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retibrevitricolporites? toigoae[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retipollenites solimoensis[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Retitriporites crassoreticulatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Rhoipites alfredii[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pollen of a flowering plant. Announced in 2020; the final version of the article naming it was published in 2021.

Singpuria[78]

Gen. et sp. nov

Valid

Ramteke, Manchester & Nagrale in Ramteke et al.

Late Cretaceous (Maastrichtian)

Deccan Intertrappean Beds

India

A member of Pentapetalae of uncertain phylogenetic placement. Genus includes new species S. kapgatei.

Sinoherba[79]

Gen. et sp. nov

Valid

Liu & Wang in Liu, Chen & Wang

Early Cretaceous (Barremian–Aptian)

Yixian Formation

China

An early monocot. Genus includes new species S. ningchengensis. Announced in 2020; the final version of the article naming it was published in 2021.

Varifructus[80]

Gen. et sp. nov

In press

Liu et al.

Early Cretaceous

Yixian Formation

China

An early flowering plant. Genus includes new species V. lingyuanensis.

Wireroadia[81]

Gen. et sp. et comb. nov

Valid

Zhang et al.

Late Cretaceous (Cenomanian to Santonian)

|  |

United States
( Alabama
Massachusetts
New Jersey
New York)

A winged fruit of a eudicot of uncertain phylogenetic placement. Genus includes new species W. viccallii, as well as W. major (Hollick).

## Pinales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Agathis ledongensis[82]

Sp. nov

Valid

Oskolski et al.

Late Oligocene–early Miocene

Qiutangling Formation

China

A species of Agathis

Agathoxylon cozzoi[83]

Sp. nov

Valid

Gnaedinger & Zavattieri

Late Triassic

Chihuido Formation

Argentina

|  |

Araucaria cuneoi[84]

Sp. nov

Valid

Noll & Kunzmann

Middle Jurassic

|  |

Argentina

A species of Araucaria.

Araucaria famii[85]

Sp. nov

Valid

Stockey & Rothwell

Late Cretaceous (Campanian)

|  |

Canada
( British Columbia)

A species of Araucaria.

Araucaria fildesensis[86]

Sp. nov

Valid

Shi et al.

Eocene

Fossil Hill Formation

Antarctica
(King George Island)

A species of Araucaria

Araucaria huncoensis[87]

Sp. nov

Valid

Rossetto-Harris in Rossetto-Harris et al.

Early Eocene

Laguna del Hunco Formation

Argentina

A species of Araucaria.

Araucaria stockeyana[84]

Sp. nov

Valid

Noll & Kunzmann

Middle Jurassic

|  |

Argentina

A species of Araucaria.

Araucarites pachacuteci[88] | Sp. nov  |

In press

Martínez in Martínez et al.

Early Cretaceous (Berriasian–Valanginian)

Huancané Formation

Peru

A member of the family Araucariaceae

Brachyoxylon zhouii[89] |

Sp. nov

Valid

Jiang et al.

Early Cretaceous

Guantou Formation

China

A conifer wood. Announced in 2020; the final version of the article naming it was published in 2021.

Brachyphyllum sattlerae[90] | Sp. nov  | Valid  | Batista et al. | Early Cretaceous (Aptian)  | Crato Formation |  Brazil | A member of the family Pinidae.  |

Callialastrobus[91]

Gen. et sp. nov

In press

Kvaček & Mendes

Early Cretaceous (late Aptian–early Albian)

Lusitanian Basin

Portugal

A pollen cone of a member of Araucariaceae. Genus includes new species C. sousai.

Cedrus anatolica[92]

Sp. nov

Valid

Akkemik

Early Miocene

Hançili Formation

Turkey

A species of Cedrus. Announced in 2020; the final version of the article naming it was published in 2021.

Circoporoxylon tibetense[93]

Sp. nov

In press

Xia et al.

Middle Jurassic (Callovian)

Xiali Formation

China

Possibly a member of the family Podocarpaceae.

Comoxostrobus[94]

Gen. et sp. nov

Valid

Stockey, Rothwell & Atkinson

Late Cretaceous (early Coniacian)

|  |

Canada
( British Columbia)

A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species C. rossii.

Cupressinoxylon klimovii[95]

Nom. nov

Valid

Blokhina

Miocene

|  |

Russia

A member of the family Cupressaceae; a replacement name for Cupressinoxylon biotoides Blokhina (1989).

Cupressinoxylon llantenesense[83]

Sp. nov

Valid

Gnaedinger & Zavattieri

Late Triassic

Llantenes Formation

Argentina

|  |
Cupressinoxylon manuelii[96] | Sp. nov  | In press  | Ríos-Santos, Cevallos-Ferriz & Pujana  | Late Cretaceous (Campanian-Maastrichtian)  | Cabullona Group |  Mexico |  |

Ductoagathoxylon wangii[97]

Sp. nov

In press

Gou & Feng in Gou et al.

Middle Jurassic

Xishanyao Formation

China

A conifer stem.

Friisia[98]

Gen. et sp. nov

In press

Mendes & Kvaček

Early Cretaceous (late Aptian – early Albian)

Lusitanian Basin

Portugal

An ovuliferous cone of a member of the family Podocarpaceae. Genus includes new species F. lusitanica.

Juniperoxylon acarcaea[99]

Sp. nov

In press

Akkemik

Early Miocene

Hançili Formation

Turkey

A member of the family Cupressaceae.

Lesbosoxylon kemaliyensis[100]

Sp. nov

Valid

Akkemik & Mantzouka in Akkemik, Mantzouka & Kıran Yıldırım

Miocene

Divriği Formation

Turkey

A member of the family Pinaceae.

Marskea cuspidata[101]

Sp. nov

Valid

Frolov & Mashchuk

Middle Jurassic

Prisayan Formation

Russia

A member of the family Taxaceae.

Mukawastrobus[102]

Gen. et sp. nov

Valid

Stockey, Nishida & Rothwell

Late Cretaceous (late Campanian—early Maastrichtian)

|  |

Japan

A member of the family Cupressaceae belonging to the subfamily Taiwanioideae. Genus includes new species M. satoi.

Piceoxylon yumeniense[103]

Sp. nov

Valid

Zhou, Peng, Deng, Zhang & Yang in Zhou et al.

Early Cretaceous

Xiagou Formation

China

Fossil wood of a member of the family Pinaceae. Announced in 2020; the final version of the article naming it was published in 2021.

Pinuxylon selmeierianum[104]

Sp. nov

Valid

Dolezych & Reinhardt

Paleogene

Eureka Sound Group

Canada
( Nunavut)

A member of the family Pinaceae described on the basis of fossil wood

Protophyllocladoxylon chijinense[103]

Sp. nov

Valid

Zhou, Peng, Deng, Zhang & Yang in Zhou et al.

Early Cretaceous

Xiagou Formation

China

Announced in 2020; the final version of the article naming it was published in 2021.

Tsuga asiatica[105]

Sp. nov

Valid

Wu & Zhou in Wu et al.

Late Paleogene

|  |

China

A species of Tsuga. Announced in 2019; the final version of the article naming it was published in 2020.

## Other seed plants

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Amyelon turpanense[106]

Sp. nov

In press

Shi, Yu & Sun

Permian (Lopingian)

Wutonggou

China

A root of a member of Cordaitales

Androstrobus obovatus[107]

Sp. nov

In press

Bodnar et al.

Late Triassic

Potrerillos Formation

Argentina

A member of Cycadales.

Araripestrobus[108]

Gen. et sp. nov

Valid

Seyfullah, Roberts, Schmidt & Kunzmann in Seyfullah et al.

Early Cretaceous (Aptian-Albian)

Crato

Brazil

A seed plant belonging to the group Erdtmanithecales. Genus includes new species A. resinosus.

Archaeopetalanthus[109]

Gen. et sp. nov

Valid

Naugolnykh

Carboniferous

Listvjanskaya

Russia

A member of Pinophyta belonging to the group Vojnovskyales. Genus includes new species A. progressus.

Battenispermum[110]

Gen. et sp. nov

In press

Mendes, Pedersen & Friis

Early Cretaceous

|  |

Portugal

A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species B. hirsutum.

Carpolithus volantus[111]

Sp. nov

Valid

Gómez et al.

Early Cretaceous (Aptian)

La Cantera Formation

Argentina

A fossil seed, possibly produced by a member of Gnetales.

Ductolobatopitys[112]

Gen. et sp. nov

Valid

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

Brazil

A gymnosperm described on the basis of fossil wood. Genus includes new species D. mussae.

Filigigantopteris[113]

Gen. et 2 sp. nov

In press

Zhou et al.

Late Permian

Nayixiong Formation

China

A gigantopterid. Genus includes new species F. asymmetrica and F. dahaia.

Ginkgo pediculata[114]

Sp. nov

In press

Deng, Yang & Zhou

Early Cretaceous

|  |

China

A species of Ginkgo.

Jianchangia[115]

Gen. et sp. nov

Valid

Yang, Wang & Ferguson

Early Cretaceous (Aptian)

Jiufotang Formation

China

A member of Ephedraceae. Genus includes new species J. verticillata.

Johniphyllum[116]

| Gen. et sp. nov  |

Valid

Looy & Duijnstee

Permian (Guadalupian)

|  |

United States
( Texas)

A member of Voltziales. Genus includes new species J. multinerve.

Jordaniopteris[117]

Gen. et comb. nov

Valid

Anderson in Anderson et al.

Permian (possibly Lopingian)

Um Irna Formation

Jordan

A seed fern. A new genus for "Dicroidium" irnensis Abu Hamad et al. (2008); genus also includes "Dicroidium" jordanensis Abu Hamad et al. (2008), "Dicroidium" robustum Kerp & Vörding (2008) and "Dicroidium" bandelii Abu Hamad et al. (2017).

Kaokoxylon brasiliensis[112]

Sp. nov

Valid

Conceição, Neregato & Iannuzzi in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

Brazil

A conifer described on the basis of fossil wood.

Nilssonia mirovanae[118]

Sp. nov

Valid

Čepičková & Kvaček

Late Cretaceous (Cenomanian)

Peruc-Korycany Formation

Czech Republic

A cycad.

Novaiorquepitys[119]

Gen. et sp. nov

In press

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

Brazil

A gymnosperm stem. Genus includes new species N. maranhensis Conceição, Neregato & Iannuzzi.

Ovalocarpus butmanii[120]

Sp. nov

Valid

Naugolnykh & Linkevich

Permian (Artinskian)

|  |

Russia
( Sverdlovsk Oblast)

A member of Ginkgoales belonging to the family Cheirocladaceae.

Palaeocupressinoxylon[121]

Gen. et sp. nov

In press

Wan, Yang & Wang

Late Permian

Turpan–Hami Basin

China

A silicified gymnospermous fossil wood. Genus includes new species P. uniseriale.

Phoenicopsis anadyrensis[122]

Sp. nov

Valid

Nosova in Zolina et al.

Late Cretaceous–Paleocene (Maastrichtian–Danian)

Rarytkin

Russia
( Chukotka Autonomous Okrug)

| A member of Czekanowskiales |

Pseudovoltzia sapflorensis[116]

Sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

|  |

United States
( Texas)

A member of Voltziales

Pterostoma neehoffii[123]

Sp. nov

Valid

Conran et al.

Middle Miocene

|  |

New Zealand

A cycad

Pteruchus frenguellii[124]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

Jordan

A pollen organ of a seed fern

Pteruchus lepidus[124]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

Jordan

A pollen organ of a seed fern

Samaropsis jinchangensis[125]

Sp. nov

Valid

Hua & Sun in Hua et al.

Early Permian

|  |

China

A seed fossil. Announced in 2019; the final version of the article naming was published in 2020.

Umaltolepis involuta[126]

Sp. nov

In press

Nosova

Middle Jurassic

Angren Formation

Uzbekistan

|  |

Umaltolepis sogdianica[126]

Sp. nov

In press

Nosova

Middle Jurassic

Angren Formation

Uzbekistan

|  |

Umkomasia aequatorialis[124]

Sp. nov

In press

Blomenkemper et al.

Late Permian

Umm Irna Formation

Jordan

A cupulate structure of a seed fern

Wantus[116]

Gen. et sp. nov

Valid

Looy & Duijnstee

Permian (Guadalupian)

|  |

United States
( Texas)

A member of Voltziales. Genus includes new species W. acaulis.

Wudaeophyton[127]

Gen. et sp. nov

Valid

Pšenička et al.

Early Permian

Taiyuan Formation

China

A small vine, most similar to pteridosperms from the group Callistophytales. Genus includes new species W. wangii.

Yangopteris[128]

Gen. et comb. nov

In press

Zhou et al.

Permian (Asselian)

|  |

China

A seed fern; a new genus for "Alethopteris" ascendens Halle.

Yvyrapitys[119]

Gen. et sp. nov

In press

Conceição & Crisafulli in Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

Brazil

A gymnosperm stem. Genus includes new species Y. novaiorquensis Conceição, Neregato & Iannuzzi.

## Other plants

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Annularia paisii[129]

Sp. nov

Valid

Correia et al.

Carboniferous (Gzhelian)

|  |

Portugal

|  |

Blasiites huolinhensis[130]

Sp. nov

In press

Li et al.

Early Cretaceous

Huolinhe Formation

China

A liverwort belonging to the family Blasiaceae.

Botryopteris multifolia[131]

Sp. nov

In press

He et al.

Permian (Lopingian)

Junlian Formation

China

A fern

Birisia mandshurica[132]

Sp. nov

Valid

Golovneva, Grabovskiy & Zolina

Early Cretaceous (Albian)

Frentsevka Formation

Russia
( Primorsky Krai)

A fern belonging to the family Dicksoniaceae.

Calamites cambrensis[133]

Sp. nov

Valid

Thomas

Carboniferous (Pennsylvanian)

|  |

United Kingdom

|  |

Catenuporella[134]

Gen. et sp. nov

Valid

Zhang et al.

Late Ordovician

Ordos Basin

China

A green alga belonging to the group Dasycladales. Genus includes new species C. gigantia.

Charaxis spicatus[135]

emend. nov.

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Maestrat Basin

Spain

Thallus of Echinochara lazarii. Both taxa has been firstly found anatomically attached

Chlamydomonas hanublikanus[136] |

Sp. nov

|  |

Vršanská & Hinkelman

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A species of Chlamydomonas

Circinites[47]

Gen. et sp. nov

In press

Robledo & Anzótegui in Robledo et al.

Miocene-Pliocene

|  |

Argentina

A fern belonging to the family Pteridaceae. Genus includes new species C. pteridoides.

Clavatisporites cenomaniana[137] |

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

Argentina

Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Clavator calcitrapus var. jiangluoensis[135] |

comb. nov

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Maestrat Basin

Spain

Clavatoracean species.

Collarecodium? nezpercae[138] | Sp. nov  | Valid  | Bucur & Rigaud in Bucur et al. | Late Triassic (Norian)  |  |  United States
( Idaho)  | A green alga belonging to the group Bryopsidales and possibly to the family Udoteaceae.  |

Collarisporites minor[137]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

Argentina

Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Coniopteris sandaolingensis[139]

Sp. nov

Valid

Yuan & Sun in Yuan et al.

Middle Jurassic

Xishanyao Formation

China

|  |

Dimicheleodendron[140]

Gen. et comb. nov

Valid

Thomas & Cleal

Carboniferous

|  |

United Kingdom

A lycophyte; a new genus for "Lepidodendron" hickii.

Drynaria diplosticha[141]

Sp. nov

Valid

Yu & Xie in Yu et al.

Late Miocene

Bangmai Formation

China

A fern belonging to the family Polypodiaceae.

Echinochara lazarii[135]

comb. nov

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Maestrat Basin

Spain

A member of Clavatoraceans.

Equicalastrobus pusillus[142]

Sp. nov

Valid

Zhang & Yan in Zhang et al.

Late Triassic

Baojishan Basin

China

A member of Equisetales. Announced in 2020; the final version of the article naming was published in 2021.

Equisetum yenbaiense[143]

Sp. nov

Valid

Aung et al.

Late Miocene

|  |

Vietnam

A species of Equisetum

Equisetum yongpingense[143]

Sp. nov

Valid

Aung et al.

Late Pliocene

Sanying Formation

Vietnam

A species of Equisetum

Filippoporella[144] | Gen. et sp. nov  | In press  | Sokač & Grgasović  | Early Paleocene |  |  Croatia | A green alga belonging to the group Dasycladales. Genus includes new species F. barattoloi.  |

Frullania partita[145]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A liverwort, a species of Frullania

Frullania vanae[146]

Sp. nov

Valid

Mamontov et al.

Eocene

Rovno amber

Ukraine

A liverwort, a species of Frullania

Gippslandites[147]

Gen. et sp. nov

Valid

McSweeney, Shimeta & Buckeridge

Late Silurian–early Devonian

|  |

Australia

A member of Zosterophyllaceae. Genus includes new species G. minutus.

Gmujij[148] | Gen. et sp. nov  | Valid  | Pfeiler & Tomescu  | Devonian (Emsian)  | Battery Point |  Canada
( Quebec)  | An early euphyllophyte. Genus includes new species G. tetraxylopteroides. Announced in 2020; the final version of the article naming it was published in 2021.  |
Griphoporella minuta[138] | Sp. nov  | Valid  | Bucur & Peybernes in Bucur et al. | Late Triassic  |  |  Japan | A green alga belonging to the group Dasycladales and the family Triploporellaceae.  |
Hansopteris[149] | Gen. et sp. nov  | In press  | Zhou et al. | Permian (Asselian)  |  |  China | An anachoropterid fern. Genus includes new species H. uncinatus.  |
Holosporella magna[138] | Sp. nov  | Valid  | Bucur & Fucelli in Bucur et al. | Late Triassic (Norian)  |  |  United States
( Nevada)  | A green alga belonging to the group Dasycladales and the family Triploporellaceae.  |
Holosporella? rossanae[138] | Sp. nov  | Valid  | Bucur & Del Piero in Bucur et al. | Late Triassic (Norian)  |  |  Canada
( Yukon)  | A green alga belonging to the group Dasycladales and the family Triploporellaceae.  |

Inocladus[150]

Gen. et comb. nov

Valid

LoDuca et al.

Silurian

|  |

United States

An alga related to the group Bryopsidales. Genus includes "Buthotrephis" divaricata White (1901), "B." newlini White (1901), "B." lesquereuxi Grote & Pitt (1876) and "Chondrites" verus Ruedemann (1925). Announced in 2020; the final version of the article naming it was published in 2021.

Intermurella ordosensis[134]

Sp. nov

Valid

Zhang et al.

Late Ordovician

Ordos Basin

China

A green alga belonging to the group Dasycladales.

Jurafructus[151]

Gen. et sp. nov

Valid

Chen et al.

Middle−Late Jurassic

Jiulongshan Formation

China

A plant of uncertain phylogenetic placement, possibly a flowering plant described on the basis of a probable fossil drupe. Genus includes new species J. daohugouensis.

Keraphyton[152]

Gen. et sp. nov

Valid

Champreux, Meyer-Berthaud & Decombeix

Devonian (Famennian)

Mandowa Mudstone Formation

Australia

A member of Iridopteridales of uncertain phylogenetic placement. Genus includes new species K. mawsoniae.

Khasurtya[153]

Gen. et sp. nov

Valid

Mamontov in Kopylov et al.

Early Cretaceous

|  |

Russia

A liverwort belonging to the group Marchantiidae. Genus includes new species K. ginkgoides.

Lepidodendron demkinae[154]

Sp. nov

Valid

Mosseichik

Carboniferous (Viséan)

|  |

Russia

|  |

Lobatannularia linjiaensis[155]

Sp. nov

Valid

Xu et al.

Middle Triassic

Linjia Formation

China

A member of Equisetales.

Lygodium sanshuiense[156]

Sp. nov

Valid

Naugolnykh et al.

Paleocene

Buxin Formation

China

A species of Lygodium. Announced in 2019; the final version of the article naming it was published in 2020.

Metzgeriites kujiensis[157]

Sp. nov

Valid

Katagiri in Katagiri & Shinden

Late Cretaceous (Santonian)

Tamagawa Formation

Japan

A liverwort.

Munieria martinclosasi[135] |

comb. nov.

Valid

Pérez-Cano, Bover-Arnal et Martín-Closas in Pérez-Cano et al.

Barremian

Lebanon

Lebanon

Clavatoracean thallus. Formerly known as Charaxis martinclosasi

Neoarthropitys[158] | Gen. et sp. nov  | In press  | Gnaedinger et al. | Middle Triassic | Quebrada de los Fósiles |  Argentina | A member of Equisetales. Genus includes new species N. gondwanaensis.  |

Osmundacaulis asiatica[159]

Sp. nov

Valid

Cheng et al.

Cretaceous

|  |

China

A member of the family Osmundaceae

Osmundacaulis sinica[159]

Sp. nov

Valid

Cheng et al.

Cretaceous

|  |

China

A member of the family Osmundaceae

Ovoidites circumplicatus[160]

Sp. nov

Valid

Zavattieri, Gutiérrez & Monti

Middle Triassic

Quebrada de los Fósiles Formation

Argentina

A green alga belonging to the group Zygnematales.

Ovoidites tripartitus[160]

Sp. nov

Valid

Zavattieri, Gutiérrez & Monti

Middle Triassic

Quebrada de los Fósiles Formation

Argentina

A green alga belonging to the group Zygnematales.

Palaeostachya guanglongii[161]

Sp. nov

In press

Liu et al.

Permian (Asselian)

Taiyuan Formation

China

A member of the family Calamitaceae.

Parazosterophyllum[147]

Gen. et sp. nov

Valid

McSweeney, Shimeta & Buckeridge

Late Silurian–early Devonian

|  |

Australia

A member of Zosterophyllaceae. Genus includes new species P. timsiae.

Patruliuspora oregonica[138] | Sp. nov  | Valid  | Bucur & Rigaud in Bucur et al. | Late Triassic (Norian)  |  |  United States
( Oregon)  | A green alga belonging to the group Dasycladales and the family Polyphysaceae.  |
Patruliuspora pacifica[138] | Sp. nov  | Valid  | Bucur, Del Piero & Peyrotty in Bucur et al. | Late Triassic (Norian)  |  |  Canada
( Yukon)  | A green alga belonging to the group Dasycladales and the family Polyphysaceae.  |
Pellites hamiensis[162] | Sp. nov  | Valid  | Li et al. | Middle Jurassic | Xishanyao |  China | A liverwort belonging to the family Pelliaceae.  |

Plenasium (Aurealcaulis) elegans[163]

Sp. nov

In press

Hiller et al.

Eocene

Na Duong Formation

Vietnam

A member of Osmundaceae

Pleuromeia shaolinii[164]

Sp. nov

Valid

Zhang & Wang in Zhang et al.

Middle Triassic

Linjia

China

|  |

Polycingulatisporites multiverrucata[137]

Sp. nov

Valid

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla

Argentina

Spores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version of the article naming it was published in 2020.

Polypodiisporites minutiverrucatus[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.

Polysporia baetica[165] |

Sp. nov

In press

Álvarez-Vázquez, Bek & Drábková

Carboniferous (Pennsylvanian)

Peñarroya-Belmez-Espiel Coalfield

Spain

A member of Isoetales

Polystichum pacltovae[166]

Sp. nov

Valid

Kvaček in Kvaček & Teodoridis

Oligocene

|  |

Czech Republic

A fern, a species of Polystichum

Proodontosoria[167]

Gen. et sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A fern belonging to the family Lindsaeaceae. Genus includes new species P. myanmarensis.

Proterocladus antiquus[168]

Sp. nov

Valid

Tang et al.

Mesoproterozoic
circa 1 Ga

Nanfen

China

An early siphonocladalean chlorophyte

Psilatriletes cozzuolii[10]

Sp. nov

Valid

Rabelo Leite, Ferreira da Silva-Caminha & D'Apolito

Miocene

Solimões Basin

Brazil

Pteridophyte spore. Announced in 2020; the final version of the article naming it was published in 2021.

Psilochara monevaensis[169]

Sp. nov

Valid

Sanjuan & Soulié-Märsche

Middle Miocene

|  |

Spain

A charophyte.

Qianshouia[170]

Gen. et sp. nov

Valid

Huang et al.

Late Devonian

Wutong Formation

China

A plant of uncertain phylogenetic placement, possibly a lycopsid or a sphenopsid. Genus includes new species Q. mira.

Scolecopteris minuta[171]

Sp. nov

In press

Wan et al.

Early Permian

Taiyuan Formation

China

A fern belonging to the group Marattiales.

Sigillaria pfefferkornii[172]

Sp. nov

In press

D'Antonio, Boyce & Wang

Permian (Asselian)

|  |

China

|  |

Sigillaria wudensis[172]

Sp. nov

In press

D'Antonio, Boyce & Wang

Permian (Asselian)

|  |

China

|  |

Sphaerochara miocenica[169]

Sp. nov

Valid

Sanjuan & Soulié-Märsche

Miocene

|  |

Lebanon
Spain

A charophyte.

Thyrsopteris cretacea[173]

Sp. nov

Valid

Li et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A species of Thyrsopteris

Tomiostrobus sinensis[174]

Sp. nov

Valid

Feng in Feng et al.

Early Triassic (Induan)

Kayitou Formation

China

A member of the family Isoetaceae.

Tumidopteris astra[175]

Sp. nov

Valid

Naugolnykh

Permian (Roadian)

Pechora coal basin

Russia

A fern belonging to the family Gleicheniaceae.

Ufadendron elongatum[176]

Sp. nov

Valid

Tang et al.

Late Permian

Linxi

China

A lycopsid belonging to the family Tomiodendraceae

Uzhurodendron[177]

Gen. et sp. nov

Valid

Mosseichik & Filimonov

Carboniferous (Tournaisian)

Bystrianskaya

Russia
( Krasnoyarsk Krai)

A member of Lycopodiopsida. Genus includes new species U. asiaticum.

Zeilleria fosteri[178]

Sp. nov

Valid

Thomas et al.

Carboniferous (Bashkirian)

|  |

United Kingdom

A fern

## General research

[edit]

* A study on the evolutionary history of green plants is published by Nie et al. (2020).[179]
* Description of new fossil material of Yurtusia uniformis from the Cambrian Yanjiahe Formation (China) and a study on the phylogenetic relationships and possible life cycle of this organism is published by Shang et al. (2020), who consider Y. uniformis to be a likely green microalga.[180]
* A study on the phylogenetic relationships of extant and fossil complex thalloid liverworts (Marchantiidae) is published by Flores et al. (2020).[181]
* Evidence of development of dichotomous roots in euphyllophytes that were extant during the Devonian and Carboniferous periods is presented by Hetherington, Berry & Dolan (2020), who interpret their findings as indicating that dichotomous root branching evolved in both lycophytes and euphyllophytes.[182]
* An early land plant producing multiple spore size classes is described from the Lower Devonian Campbellton Formation (Canada) by Bonacorsi et al. (2020).[183]
* A study on the impact of the appearance and evolution of herbivorous tetrapods on the evolution of land plants from the Carboniferous to the Early Triassic is published by Brocklehurst, Kammerer & Benson (2020).[184]
* A study on the production of periderm in Late Paleozoic arborescent lycopsids is published by D'Antonio & Boyce (2020), who argue that these lycopsids did not grow from sporelings into large trees through the production of a periderm cylinder, because physiological limitations would have prohibited the production of thick periderm.[185]
* A study on the architecture and development of the Carboniferous arborescent lycopsid Paralycopodites is published by DiMichele & Bateman (2020).[186]
* New information on the anatomy of Weichselia reticulata is presented by Blanco-Moreno, Decombeix & Prestianni (2020).[187]
* A study on the phylogenetic placement of the extinct fern genus Coniopteris is published by Li et al. (2020).[188]
* New information on the morphology of Paleoazolla patagonica is presented by Benedetti et al. (2020), who evaluate the implications of this taxon for the knowledge of the evolution of water ferns.[189]
* A study aiming to determine which ferns were most likely to be the producers of Cyathidites spores from earliest Paleocene plant localities across western North America, and were most likely to be among the first plants in western North America to thrive in the immediate aftermath of the Cretaceous–Paleogene extinction event, is published by Berry (2020).[190]
* A study on the morphology and development of Genomosperma, and on its implications for the knowledge of the evolutionary origins of seed development, is published by Meade, Plackett & Hilton (2020).[191]
* A pollen organ resembling seed fern pollen organs Dictyothalamus and Melissiotheca is described from the Lopingian Umm Irna Formation (Jordan) by Zavialova et al. (2020), who interpret this finding as evidence of persistence of lyginopterid seed ferns until the late Permian.[192]
* Evidence of increasing atmospheric CO2 concentration at the onset of the end-Triassic extinction event, based on data from fossil leaves of the seed fern Lepidopteris ottonis from southern Sweden, is presented by Slodownik, Vajda & Steinthorsdottir (2020), who confirm L. ottonis as a valid proxy for pCO2 reconstructions.[193]
* A study on the anatomy of the seed cone scales of Krassilovia mongolica is published by Herrera et al. (2020), who argue that K. mongolica and Podozamites harrisii are the seed cones and leaves of the same extinct plant, and name a new family Krassiloviaceae within the order Voltziales.[194]
* A study on the microscopic wood anatomy of a fossil tree trunk of Agathoxylon arizonicum with the characteristic external features of a fire scar from the Upper Triassic Chinle Formation (Petrified Forest National Park, Arizona, United States) is published by Byers et al. (2020), who evaluate the implications of this specimen for the knowledge of the evolution of fire-adapted plant traits.[195]
* A putative bamboo "Chusquea" oxyphylla from the early Eocene Laguna del Hunco biota (Argentina) is reinterpreted as a conifer by Wilf (2020), who transfers this species to the genus Retrophyllum.[196]
* A study on evolutionary history of conifers as indicated by fossil and molecular data, aiming to determine whether the rise of angiosperms drove the decline of conifers and other gymnosperms, is published by Condamine et al. (2020).[197]
* Presence of secretory tissues is reported in extinct flowers from the Cretaceous amber from Myanmar and Cenozoic Dominican amber (including specimens preserved while in the process of emitting compounds) by Poinar & Poinar (2020).[198]
* Fossil pollen of flowering plants is reported from the Aptian and Albian of Australia by Korasidis & Wagstaff (2020), who evaluate the implications of their findings for the knowledge of the timing of the appearance and diversification of the flowering plants in the high-latitude southern basins of Australia.[199]
* A study on the morphology of palm and palm-like pollen from the Eocene Yaw Formation (Myanmar), and on the implications of these fossils for the knowledge of distribution and diversity of Eocene palms across the globe, is published by Huang et al. (2020).[200]
* Fossils fruits of Illigera eocenica, representing the second fossil occurrence of Illigera worldwide and the first in Asia, are described from the Eocene Niubao Formation (central Tibetan Plateau) by Wang et al. (2020), who evaluate the implications of this finding for the knowledge of the climate in the central Tibetan Plateau during the early middle Eocene, and for the knowledge of the floristic links between Asia and North America during the Paleogene.[201]
* A study on the morphology and phylogenetic relationships of Montsechia vidalii is published by Gomez et al. (2020).[202]
* Eocene leaves of members of the family Urticaceae with stinging trichomes are described from the Okanogan Highlands (British Columbia, Canada) by DeVore et al. (2020).[203]
* A revision of the fossil record of the family Nothofagaceae from South America is published by Pujana et al. (2020).[204]
* A study on the extinction of plants from south polar terrestrial ecosystems during the Permian–Triassic extinction event and on their recovery after this extinction event, based on data from the Sydney Basin (Australia), is published by Mays et al. (2020).[205]
* A study on the impact of ecological disturbances around the Permian–Triassic boundary (from the Wuchiapingian to Ladinian) on land plant communities is published by Nowak, Vérard & Kustatscher (2020).[206]
* A study on the age of the Paleogene Kanaka Creek fossil flora (Huntingdon Formation; British Columbia, Canada) and on its implications for reconstructions of the contemporaneous paleoclimate and paleoenvironment is published by Mathewes, Greenwood & Love (2020).[207]
* Evidence from Eocene plant fossils from the Bangong-Nujiang suture indicating that the Tibetan Plateau area hosted a diverse subtropical ecosystem approximately 47 million years ago and that this area was both low and humid at the time is presented by Su et al. (2020), who also report that the composition of this flora is similar to Early-Middle Eocene floras in both North America and Europe, but shows little affinity to Eocene floras from the Indian Plate.[208]
* A study aiming to estimate leaf dry mass per area in fossil plants from 22 western North American sites spanning the Eocene–Oligocene transition is published online by Butrim & Royer (2020), who evaluate the implications of their findings for the knowledge of the impact of the environmental changes occurring during the Eocene–Oligocene transition on leaf-economic strategies of plants.[209]
* A study on the Neogene paleobotanical record and climate in the northernmost part of the Central Andean Plateau, based on data from the Descanso Formation (Peru), is published by Martínez et al. (2020), who report the earliest evidence of a puna-like ecosystem in the Pliocene and a montane ecosystem without modern analogs in the Miocene.[210]
* Fossil fruits (mericarps) of the neoendemic Apiaceae Melanoselinum (≡ Daucus) decipiens were reported from the lacustrine and fluvial sediments of Porto da Cruz, Madeira, dated 1.3 Ma, by Góis-Marques et al. 2020.[211] This paper not only reports the oldest Daucus s.l. fossil known to date but also the first fossil evidence of a plant with insular woodiness (see Island gigantism).
* The leaf fossil Mesodescolea plicata from the Early Cretaceous of Patagonia, first interpreted as a cycad with affinities with extant Stangeria, is reinterpreted as an angiosperm leaf with affinities with Austrobaileyales or Chloranthales by Coiro et al. 2020,[212] with implications for the evolution of leaf shape in the early radiation of the angiosperms.
* A study on the phylogenetic relationships of 10 Cretaceous flower taxa (Chloranthistemon endressii, Dakotanthus cordiformis, Kajanthus lusitanicus, Mauldinia mirabilis, Microvictoria svitkoana, Paleoclusia chevalieri, Paradinandra suecica, Spanomera mauldiniensis, Tylerianthus crossmanensis and Virginianthus calycanthoides) is published by Schönenberger et al. (2020).[213]

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