2021 in paleoentomology

Overview of the events of 2021 in paleoentomology

|  | List of years in paleoentomology  |

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* …
* 2018
* 2019
* 2020
* 2021
* 2022
* 2023
* 2024
* …

In paleontology
2018
2019
2020
2021
2022
2023
2024

In paleobotany
2018
2019
2020
2021
2022
2023
2024

In arthropod paleontology
2018
2019
2020
2021
2022
2023
2024

In paleoichthyology
2018
2019
2020
2021
2022
2023
2024

In paleomalacology
2018
2019
2020
2021
2022
2023
2024

In paleomammalogy
2018
2019
2020
2021
2022
2023
2024

In reptile paleontology
2018
2019
2020
2021
2022
2023
2024

In archosaur paleontology
2018
2019
2020
2021
2022
2023
2024

* Art
* Archaeology
* Architecture
* Literature
* Music
* Philosophy
* Science +...

Further information: 2021 in arthropod paleontology and 2021 in paleontology

This paleoentomology list records new fossil insect taxa that were to be described during the year 2021, as well as notes other significant paleoentomology discoveries and events which occurred during that year.

## Clade Amphiesmenoptera[edit]

### Lepidoptera[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

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Acanthocorona[1]

Gen. et 7 sp. nov

Valid

Mey, Léger & Lien

Cretaceous
Cenomanian

Burmese amber

Myanmar

A Lophocoronidae moth.
The type species is A. skalskii
The genus also includes A. bowangi, A. kuranishii, A. muelleri, A. sattleri, A. spinifera, & A. wichardi.

|  Acanthocorona skalskii

Agathiphagama[1]

Gen. et sp. nov

Valid

Mey, Léger & Lien

Cretaceous
Cenomanian

Burmese amber

Myanmar

An agathiphagid aglossatan moth.
The type species is A. perdita.

|  Agathiphagama perdita

Phyllocnistis cretacea[2]

Sp. nov

Valid

Fischer

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A species of Phyllocnistis

Phyllonorycter inopinata[2]

Sp. nov

Valid

Fischer

Lutetian

Baltic amber

Russia
( Kaliningrad Oblast)

A species of Phyllonorycter

#### Lepidopteran research[edit]

* The first known Cretaceous caterpillar armoured with spines is described from the Burmese amber by Haug & Haug (2021).[3]
* Fossil leaf mine produced by a lyonetiid-like leaf-mining moth, representing the oldest record of such moths reported to date, is described from the Upper Cretaceous (Campanian) Kaiparowits Formation (Utah, United States) by Maccracken et al. (2021), who name a new ichnotaxon Leucopteropsa spiralae.[4]

### Trichopterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

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Archaeopolycentra achupomotet[5]

Sp. nov

Valid

Melnitsky & Ivanov

Late Cretaceous (Santonian)

Kheta Formation
(Taymyr amber)

Russia
( Krasnoyarsk Krai)

A member of the family Polycentropodidae.

Archaeopolycentra turvalsia[5]

Sp. nov

Valid

Melnitsky & Ivanov

Late Cretaceous (Santonian)

Kheta Formation
(Taymyr amber)

Russia
( Krasnoyarsk Krai)

A member of the family Polycentropodidae.

Cernotina fossilinova[6]

Sp. nov

Valid

Wichard & Neumann

Probably early Miocene

Dominican amber

Dominican Republic

A caddisfly, a species of Cernotina.

Electrotrichia rasnitsyni[7]

Sp. nov

Valid

Melnitsky, Ivanov & Perkovsky

Late Eocene

Rovno amber

Ukraine

A member of the family Hydroptilidae.

Folindusia aenigmatosa[8]

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

|  |

Russia

A larval case of a caddisfly.

Folindusia expansa[8]

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

|  |

Russia

A larval case of a caddisfly.

Folindusia inconspicua[8]

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

|  |

Russia

A larval case of a caddisfly.

Folindusia inventa[8]

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

|  |

Russia

A larval case of a caddisfly.

Folindusia robusta[8]

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

|  |

Russia

A larval case of a caddisfly.

Folindusia tortilis[8]

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

|  |

Russia

A larval case of a caddisfly.

Holocentropus tutkaktut[9]

Sp. nov

Valid

Melnitsky, Ivanov & Perkovsky

Eocene (Priabonian)

Rovno amber

Ukraine

A species of Holocentropus.

Palaeopsilotreta kachini[10]

Sp. nov

Valid

Wichard, Müller & Xu

Cretaceous

Burmese amber

Myanmar

A member of the family Odontoceridae.

Palaeopsilotreta succini[10]

Sp. nov

Valid

Wichard, Müller & Xu

Cretaceous

Burmese amber

Myanmar

A member of the family Odontoceridae.

Plectrocnemia kirmikhia[11]

Sp. nov

Valid

Melnitsky, Ivanov & Perkovsky

Eocene (Priabonian)

Rovno amber

Ukraine

A species of Plectrocnemia.

Psilotreta fossilis[10]

Sp. nov

Valid

Wichard, Müller & Xu

Cretaceous

Burmese amber

Myanmar

A species of Psilotreta.

Purbimodus khramovi[12]

Sp. nov

Valid

Sukatsheva & Aristov

Early Cretaceous

|  |

Russia

A member of the family Vitimotauliidae.

Terrindusia nova[8]

Sp. nov

Valid

Sukacheva & Vasilenko

Early Cretaceous

|  |

Russia

A larval case of a caddisfly.

Wormaldia denticulata[13]

Sp. nov

In press

Wang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A caddisfly, a species of Wormaldia.

Wormaldia diplobifurca[13]

Sp. nov

In press

Wang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A caddisfly, a species of Wormaldia.

## Clade Antliophora[edit]

### Dipterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

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Acartophthalmites crassipes[14]

Sp. nov

Valid

Roháček & Hoffeins

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A member of Opomyzoidea belonging to the new family Clusiomitidae.

Acartophthalmites luridus[14]

Sp. nov

Valid

Roháček & Hoffeins

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A member of Opomyzoidea belonging to the new family Clusiomitidae.

Acartophthalmites rugosus[14]

Sp. nov

Valid

Roháček & Hoffeins

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A member of Opomyzoidea belonging to the new family Clusiomitidae.

Aleksiana[15]

Gen. et sp. et comb. nov

Valid

Krzemiński et al.

Cretaceous

Burmese amber

Myanmar

A member of the crane fly family Limoniidae. The type species is A. rasnitsyni; genus also includes "Antocha" lapra Podenas & Poinar (2009).

Allarete patriciae[16]

Sp. nov

Valid

Nel

Early Eocene

Oise amber

France

A member of the family Cecidomyiidae.

Allodia paleoafricana[17]

Sp. nov

Valid

Bouju et al.

Early Miocene

Ethiopian amber

Ethiopia

A species of Allodia.

Apocephalus chiapanecus[18]

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

Mexican amber

Mexico

A species of Apocephalus.

Apocephalus dominicanus[18]

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

Dominican amber

Dominican Republic

A species of Apocephalus.

Apocephalus miocenus[18]

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

|  |  |

A species of Apocephalus.

Araripogon pulchrus[19]

Sp. nov

In press

Lamas, Sampronha & Ribeiro

Early Cretaceous (Aptian)

Crato Formation

Brazil

A member of the family Asilidae.

Bashkonia[20]

Gen. et sp. nov

Valid

Lukashevich

Middle Triassic

Röt Formation

Germany

A member of the family Nadipteridae. Genus includes new species B. franconica.

Bibio castaneipennis[21]

Sp. nov

Valid

Skartveit & Wedmann

Oligocene

|  |

Germany

A species of Bibio.

Bibio succineus[22]

Sp. nov

Valid

Skartveit

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Bibio.

Burmahesperinus[23]

Gen. et 3 sp. nov

Valid

Ševčík et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Bibionidae. The type species is B. antennatus; genus also includes B. conicus and B. pedicellatus.

Burmasymmerus[24]

Gen. et 2 sp. nov

Valid

Ševčík, Hippa & Burdíková

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Ditomyiidae. The type species is B. korneliae; genus also includes B. wieslawi.

Burmatricha[24]

Gen. et sp. nov

Valid

Ševčík, Hippa & Burdíková

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Sciaroidea of uncertain phylogenetic placement. The type species is B. mesozoica.

Cascomixticus[25]

Gen. et sp. nov

Valid

Poinar & Vega

Cretaceous (Albian/Cenomanian)

Burmese amber

Myanmar

A member of the family Apsilocephalidae. The type species is C. tubuliferous.

Clusiomites[14]

Fam. et gen. et comb. et sp. nov

Valid

Roháček & Hoffeins

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A member of Opomyzoidea, the type genus of the new family Clusiomitidae. The type species is "Acartophthalmites" clusioides Roháček (2016); genus also includes new species C. ornatus.

Crenoptychoptera gaoi[26]

Sp. nov

Valid

Liu & Huang

Middle–Late Jurassic

Jiyuan Basin

China

A member of the family Ptychopteridae.

Cretobibio rasnitsyni[27]

Sp. nov

Valid

Lukashevich, Amorim & Ribeiro

Early Cretaceous (Aptian)

Crato Formation

Brazil

A member of the family Bibionidae.

Cretolimonia dayana[28]

Sp. nov

|  |

Kopeć in Kopeć et al.

Jurassic/Cretaceous boundary

Glushkovo Formation

Russia
( Zabaykalsky Krai)

A member of the subfamily Architipulinae.

Cretolimonia lukashevichae[28]

Sp. nov

|  |

Kopeć & Krzemiński in Kopeć et al.

Jurassic/Cretaceous boundary

Dain Formation

Russia
( Zabaykalsky Krai)

A member of the subfamily Architipulinae.

Cretolimonia mikolajczyki[28]

Sp. nov

|  |

Kopeć, Krzemiński, Soszyńska-Maj in Kopeć et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the subfamily Architipulinae.

Cretolimonia pseudojurassica[28]

Sp. nov

|  |

Krzemiński in Kopeć et al.

Jurassic/Cretaceous boundary

Dain Formation

Russia
( Zabaykalsky Krai)

A member of the subfamily Architipulinae.

Cretothereva[29]

Gen. et sp. nov

In press

Carmo, Lamas & Ribeiro

Early Cretaceous

Crato Formation

Brazil

A member of the family Therevidae. Genus includes new species C. antiqua.

Cretpenthetria[30]

Gen. et sp. nov

Valid

Li, Zhang & Xiao

Cretaceous

Burmese amber

Myanmar

A member of the family Bibionidae. Genus includes new species C. burmensis.

Culex ekaterinae[31]

Sp. nov

Valid

Giłka, Harbach & Perkovsky

Eocene

Rovno amber

Ukraine

A mosquito, a species of Culex.

Culicoides paleopestis[32]

Sp. nov

In press

Peñalver et al.

Eocene

Anglesea amber

Australia

A species of Culicoides.

Cyttaromyia freiwaldi[33]

Sp. nov

Valid

Kania-Kłosok et al.

Eocene

Green River Formation

United States

A member of the family Cylindrotomidae.

Cyttaromyia gelhausi[33]

Sp. nov

Valid

Kania-Kłosok et al.

Eocene

Green River Formation

United States

A member of the family Cylindrotomidae.

Decessia[34]

Gen. et sp. nov

Valid

Krzemiński et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the crane fly family Limoniidae. The type species is D. podenasi.

Elephantomyia christelae[35]

Sp. nov

|  |

Kania-Kłosok & Krzemiński

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Elephantomyia.

Elephantomyia prima[35]

Sp. nov

|  |

Kania-Kłosok & Krzemiński

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Elephantomyia.

Forcipomyia rasnitsyni[36]

Sp. nov

Valid

Szadziewski, Sontag & Pankowski

Miocene

Ethiopian amber

Ethiopia

A species of Forcipomyia.

Helius hispanicus[37]

Sp. nov

|  |

Kania-Kłosok, Krzemiński & Arillo

Early Cretaceous (Albian)

Utrillas Group
(Álava amber)

Spain

A species of Helius.

Helius turolensis[37]

Sp. nov

|  |

Kania-Kłosok, Krzemiński & Arillo

Early Cretaceous (Albian)

Utrillas Group
(San Just amber)

Spain

A species of Helius.

Hirmoneura messelense[38]

Sp. nov

Valid

Wedmann et al.

Eocene (Lutetian)

Messel Formation

Germany

A member of the family Nemestrinidae.

Lebambromyia sacculifera[39]

Sp. nov

Valid

Badano, Zhang, Yeates & Cerretti in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Phoroidea.

Leptotarsus andradei[40]

Sp. nov

Valid

Ribeiro, Santos & Santos

Early Cretaceous

Crato Formation

Brazil

A species of Leptotarsus.

Leptotarsus gelhausi[40]

Sp. nov

Valid

Ribeiro, Santos & Santos

Early Cretaceous

Crato Formation

Brazil

A species of Leptotarsus.

Leptotarsus ivoneae[40]

Sp. nov

Valid

Ribeiro, Santos & Santos

Early Cretaceous

Crato Formation

Brazil

A species of Leptotarsus.

Leptotarsus lemeae[40]

Sp. nov

Valid

Ribeiro, Santos & Santos

Early Cretaceous

Crato Formation

Brazil

A species of Leptotarsus.

Mesoacentron[41]

Gen. et sp. nov

Valid

Giłka, Zakrzewska, Lukashevich & Cranston in Giłka et al.

Late Cretaceous (Santonian)

Kheta Formation
(Taimyr amber)

Russia

A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is M. kaluginae.

Meunierohelea anglesensis[32]

Sp. nov

In press

Peñalver et al.

Eocene

Anglesea amber

Australia

A member of the family Ceratopogonidae.

Minyohelea nexuosa[42]

Sp. nov

In press

Pielowska-Ceranowska, Azar & Szwedo

Early Cretaceous (Barremian)

Lebanese amber

Lebanon

A member of the family Ceratopogonidae.

Myriophora asiatica[18]

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

Zhangpu amber

China

A species of Myriophora.

Neoarisemus groehni[43]

Sp. nov

Valid

Wagner

Eocene

|  |

Ukraine

A member of the family Psychodidae.

Orimarguloides[44]

Gen. et sp. nov

Valid

Men & Podenas

Cretaceous

Burmese amber

Myanmar

A member of the crane fly family Limoniidae. Genus includes new species O. simplex.

Palaeocentron[41]

Gen. et sp. nov

Valid

Giłka, Zakrzewska, Lukashevich & Cranston in Giłka et al.

Cretaceous (probably Albian–Cenomanian)

Burmese amber

Myanmar

A member of the family Chironomidae belonging to the subfamily Chironominae and the tribe Pseudochironomini. The type species is P. krzeminskii.

Palaeoglaesum pilosus[45]

Sp. nov

Valid

Skibińska, Krzemiński & Zhang in Skibińska et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.

Palaeoglaesum stebneri[45]

Sp. nov

Valid

Skibińska & Krzemiński in Skibińska et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.

Palaeoglaesum teres[45]

Sp. nov

Valid

Skibińska & Albrycht in Skibińska et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Psychodidae belonging to the subfamily Bruchomyiinae.

Paleoplatyura agnieszkae[46]

Sp. nov

Valid

Ševčík, Krzemiński & Skibińska

Cretaceous

Burmese amber

Myanmar

A species of Paleoplatyura.

Paleoplatyura magnifica[46]

Sp. nov

Valid

Ševčík, Krzemiński & Skibińska

Cretaceous

Burmese amber

Myanmar

A species of Paleoplatyura.

Paleoplatyura miae[46]

Sp. nov

Valid

Ševčík, Krzemiński & Skibińska

Cretaceous

Burmese amber

Myanmar

A species of Paleoplatyura.

Paleotelmatoscopus[47]

Gen. et comb. et sp. nov

Valid

Curler & Skibińska

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Psychodidae. The type species is "Pericoma" formosa Meunier (1905); genus also includes new species P. madrizi.

Polymera alexanderi[48]

Sp. nov

Valid

Krzemiński et al.

Miocene

Dominican amber

Dominican Republic

A species of Polymera.

Protopenthetria[49]

Gen. et sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Bibionidae. Genus includes new species P. skartveiti.

Rhabdomastix asiatica[50]

Sp. nov

Valid

Kania-Kłosok et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A species of Rhabdomastix.

Rhabdomastix cretacica[50]

Sp. nov

Valid

Kania-Kłosok et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A species of Rhabdomastix.

Rhabdomastix krzeminskae[50]

Sp. nov

Valid

Kania-Kłosok et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A species of Rhabdomastix.

Rhabdomastix myanmae[50]

Sp. nov

Valid

Kania-Kłosok et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A species of Rhabdomastix.

Rhipidoxylomyia rasnitsyni[16]

Sp. nov

Valid

Nel

Early Eocene

Oise amber

France

A member of the family Cecidomyiidae.

Stempellinella gilkai[51]

Sp. nov

Valid

Zakrzewska & Jankowska

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Stempellinella.

Trichoneura xavieri[52]

Sp. nov

Valid

Kania-Kłosok et al.

Early Cretaceous (Albian)

Utrillas Group

Spain

A species of Trichoneura.

Zhangsolva burmensis[53]

Sp. nov

In press

Zhang & Zhang

Cretaceous

Burmese amber

Myanmar

A member of the family Zhangsolvidae.

#### Dipteran research[edit]

* Baranov et al. (2021) document a high diversity of fly larvae from the Eocene Baltic amber, preserving taxonomically, morphologically, and ecologically important characters, and evaluate the implications of this finding for the knowledge of the functional ecology of the Eocene fly larvae.[54]

### Mecopterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

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Conicholcorpa longa[55]

Sp. nov

|  |

Zhang, Shih & Ren in Zhang et al.

Middle Jurassic

Jiulongshan Formation

China

A scorpionfly belonging to the family Holcorpidae.

Glanomerope[56]

Gen. et sp. nov

Valid

Poschmann & Nel

Permian

Meisenheim Formation

Germany

A scorpionfly belonging to the family Protomeropidae. Genus includes new species G. virgoferroa.

Longiphlebia incompleta[57]

Sp. nov

In press

Lian, Cai & Huang

Late Jurassic

Tiaojishan Formation

China

A scorpionfly belonging to the family Orthophlebiidae.

Mesopsyche jinsuoguanensis[58]

Sp. nov

Valid

Lian, Cai & Huang

Middle Triassic

Tongchuan Formation

China

A member of the family Mesopsychidae.

Mesopsyche liaoi[58]

Sp. nov

Valid

Lian, Cai & Huang

Late Triassic

Huangshanjie Formation

China

A member of the family Mesopsychidae.

## Clade Archaeorthoptera[edit]

### †Caloneurodea[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

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Aviobiella[59]

Gen. et sp. nov

Valid

Nel & Roques

Carboniferous (Moscovian)

|  |

France

A member of Archaeorthoptera belonging to the order Caloneurodea. Genus includes new species A. garroustei.

Oudardgramma[60]

Gen. et sp. nov

|  |

Nel & Roques

Carboniferous (Bashkirian)

|  |

France

The type species is O. bruayensis.

Paraspiculum[61]

Gen. et sp. nov

In press

Dvořák et al.

Carboniferous (Moscovian)

|  |

Germany

A member of Archaeorthoptera belonging to the group Caloneurodea. Genus includes new species P. brauckmannii.

### †Cnemidolestodea[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

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Avionxixia[62]

Gen. et sp. nov

Valid

Nel & Roques

Carboniferous (Moscovian)

|  |

France

A member of Archaeorthoptera belonging to the family Cnemidolestidae. The type species is A. gui.

Ctenoptilus frequens[63]

Sp. nov

Valid

Chen et al.

Carboniferous (Bashkirian to Moscovian)

Yanghugou Formation

China

A member of Archaeorthoptera belonging to the family Ctenoptilidae.

Palatinarkema[64]

Gen. et sp. nov

Valid

Nel & Poschmann

Early Permian (probably Asselian–?Sakmarian)

Meisenheim Formation

Germany

A member of Archaeorthoptera belonging to the family Cnemidolestidae. The type species is P. prokopi.

Piesbergopterum[61]

Gen. et 2 sp. nov

In press

Dvořák et al.

Carboniferous (Moscovian)

|  |

France
Germany

A member of Archaeorthoptera belonging to the group Cnemidolestodea. Genus includes new species P. punctatum Dvořák et al. and P. avionensis Nel & Roques.[65]

### Orthopterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

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Amberotridactylus[66]

Gen. et sp. nov

In press

Du, Xu & Zhang

Cretaceous

Burmese amber

Myanmar

A member of the family Tridactylidae. Genus includes new species A. cheni.

Archaboilus polyneurus[67]

Sp. nov

Valid

Gu, Yue & Ren in Gu et al.

Middle Jurassic (Bathonian–Callovian)

Jiulongshan Formation

China

An ensiferan belonging to the group Hagloidea and the family Haglidae.

Caelielca[68]

Gen. et sp. nov

In press

Uchida

Cretaceous

Burmese amber

Myanmar

A member of the family Elcanidae. Genus includes new species C. spinocrus.

Cascogryllus setosus[66]

Sp. nov

In press

Du, Xu & Zhang

Cretaceous

Burmese amber

Myanmar

A member of the family Tridactylidae.

Cratoelcana rasnitsyni[69]

Sp. nov

In press

Nel & Jouault

Early Cretaceous

Crato Formation

Brazil

A member of the family Elcanidae.

Hukawnelca[68]

Gen. et sp. nov

In press

Uchida

Cretaceous

Burmese amber

Myanmar

A member of the family Elcanidae. Genus includes new species H. gracile.

Panorpidium spica[70]

Sp. nov

In press

Kim et al.

Early Cretaceous (Albian)

Jinju Formation

South Korea

An orthopteran belonging to the family Elcanidae.

Permohagla[71]

Gen. et sp. nov

Valid

Gorochov

Late Permian

|  |

Russia
( Kemerovo Oblast)

A member of Hagloidea. Genus includes new species P. tomica.

Phyllotridactylus[72]

Gen. et sp. nov

In press

Xu et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Tridactylidae. Genus includes new species P. wangi.

Pseudoacrida sennlaubi[69]

Sp. nov

In press

Nel & Jouault

Early Cretaceous

Crato Formation

Brazil

A member of the family Locustopsidae.

#### Orthopteran research[edit]

* Revision of the fossil record of the family Trigonidiidae is published by Desutter-Grandcolas et al. (2021).[73]

### †Titanoptera[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Theiatitan[74]

Fam. et gen. et sp. nov

Valid

Schubnel, Roques & Nel in Schubnel et al.

Carboniferous (Moscovian)

|  |

France

A member of Titanoptera belonging to the new family Theiatitanidae. The type species is T. azari.

### Other archaeorthopterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Archaeogerarus[62]

Fam. et gen. et sp. nov

Valid

Nel & Roques

Carboniferous (Moscovian)

|  |

France

A member of Archaeorthoptera belonging to the new family Archaeogeraridae. The type species is A. schubneli.

Avionugonioneura[75]

Gen. et sp. nov

Valid

Nel & Roques

Carboniferous (Moscovian)

|  |

France

A member of Archaeorthoptera possibly belonging to the family Nugonioneuridae. Genus includes new species A. jouaulti.

Contracladus[61]

Gen. et sp. nov

In press

Dvořák et al.

Carboniferous (Moscovian)

|  |

Germany

A member of Archaeorthoptera of uncertain phylogenetic placement. Genus includes new species C. impar.

Duquesnia[76]

Gen. et sp. nov

Valid

Nel & Roques in Nel, Roques & Duquesne

Carboniferous (Moscovian)

|  |

France

A member of Archaeorthoptera. Genus includes new species D. gallica.

Penanoptera[77]

Gen. et sp. nov

Valid

Nel & Poschmann

Early Permian

Saar–Nahe Basin

Germany

A member of Archaeorthoptera. Genus includes new species P. manseri.

## Clade Coleopterida[edit]

### Coleoptera[edit]

#### Adephaga[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

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|

Antephilorhizus[78]

Gen. et sp. nov

Valid

Kirichenko-Babko, Perkovsky & Vasilenko

Eocene

Rovno amber

Ukraine

A ground beetle belonging to the tribe Lebiini. Genus includes new species A. pripiatiensis.

Balticeler[79]

Gen. et sp. nov

Valid

Schmidt & Maddison in Schmidt, Scholz & Maddison

Eocene

Baltic amber

Europe (Baltic Sea region)

A ground beetle belonging to the subfamily Trechinae. The type species is B. kerneggeri.

Calosoma kimi[80]

Sp. nov

Valid

Lee & Nam

Early Miocene

Geumgwangdong Formation

South Korea

A species of Calosoma.

#### Archostemata[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
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|
|
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|
|
|

Jurodes shef[81]

Sp. nov

Valid

Yan, Strelnikova & Cai

Early Cretaceous (Berriasian-Barremian)

Gusinoe Ozero Group

Russia
( Buryatia)

A Jurodid beetle.

Kirejtomma[82]

Gen. et comb. nov

Valid

Li & Cai

Cretaceous

Burmese amber

Myanmar

An Ommatid beetle
The type species is "Clessidromma" zengi (2020).

|  Kirejtomma zengi

Limnomma[83]

Gen. et sp. nov

Valid

Li & Cai in Li et al.

Middle Jurassic

Haifanggou Formation

China

An Ommatid beetle.
The type species is L. daohugouense.

|  Limnomma daohugouense

Omma axsmithi[84]

Sp. nov

In press

Jarzembowski, Zheng & Zhao

Cretaceous

Burmese amber

Myanmar

An Omma species Ommatid beetle.

Omma forte[85]

Sp. nov

Valid

Li & Cai
in Li, Huang & Cai

Cretaceous
(Albian to Cenomanian)

Burmese amber

Myanmar

An Omma species Ommatid beetle.

Paraodontomma leptocristatum[86]

Sp. nov

Valid

Li, Yamamoto & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

Myanmar

An Ommatid beetle.

|  Paraodontomma leptocristatum

Permocatinus[87]

Gen. et sp. nov

Valid

Ponomarenko

Late Permian

|  |

Russia

A Coleocatiniid beetle.
The type species is P. tomiensis.

Tomiaplus[87]

Gen. et comb. et sp. nov

Valid

Ponomarenko

Late Permian

|  |

Russia

A Triaplid beetle.
The type species is "Triaplus" sibiricus (2013),
as well as new species T. minimus.

Tunguskagyrus уani[87]

Sp. nov

Valid

Ponomarenko

Late Permian

|  |

Russia

A Triaplid beetle.

Zygadenia alexrasnitsyni[88]

Sp. nov

Valid

Strelnikova & Yan

Early Cretaceous

Zaza Formation

Russia

An Ommatid beetle.

#### Myxophaga[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Triamyxa[89]

Fam., Gen. et sp. nov

Valid

Qvarnström et al.

Late Triassic (Carnian)

Keuper succession (Drawno Beds)

Poland

A Triamyxid beetle in the suborder Myxophaga.
The type species is T. coprolithica.

|  Triamyxa coprolithica

#### Polyphaga[edit]

##### Bostrichiformia[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
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|
|
|
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|
|

Archaenosodendron[90]

Gen. comb. et 3 sp. nov

Valid

Li & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

Myanmar

A nosodendrid beetle.
The type species is "Nosodendron" cretaceum;
genus also includes new species A. angulare, A. explanatum and A. remotidens.

Cretomegatoma[91]

Gen. et comb. nov

Valid

Háva

Late Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A carpet beetle.
The type species is "Megatoma" atypica (2017).

Mesonosa[92]

Gen. et sp. nov

Valid

Tihelka et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A wounded-tree beetle.
The type species is M. scandens.

Petalium bruteno[93]

Sp. nov

Valid

Alekseev & Bukejs

Late Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

A Petalium species death watch beetle.

Petalium widewuto[93]

Sp. nov

Valid

Alekseev & Bukejs

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A Petalium species death watch beetle.

Trogoderma ainu[94]

Sp. nov

Valid

Perkovsky, Háva & Zaitsev

Middle Eocene

Sakhalin amber

Russia

A Trogoderma species carpet beetle.

Tuberphradonoma[91]

Gen. et sp. nov

Valid

Háva

Late Cretaceous
Cenomanian)

Burmese amber

Myanmar

A carpet beetle.
The type species is T. burmitica.

Xyletinus lobanovi[95]

Sp. nov

Valid

Bukejs, Alekseev & Háva

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A Xyletinusspecies death watch beetle.

##### Cucujiformia[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Aberrokorynetes oceanojubilaei[96]

Sp. nov

In press

Kolibáč et al.

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A Korynetinae checkered beetle.

Aliattalus[97]

Gen. et sp. nov

Valid

Tshernyshev

Late Eocene

Baltic amber

Europe

A Malachiinae malachite beetle.
The type species is A. intercalaris.

Archaeosciaphilus gyrommatus[98]

Sp. nov

In press

Legalov & Poinar

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A Polydrusini Entiminae broad nosed weevil.

Arkagalarhinus[99]

Gen. et sp. nov

In press

Legalov

Late Cretaceous
(Santonian-Campanian)

Arkagala Formation

Russia

A Chilecarinae ithycerid weevil.
The type species is A. zherikhini.

Asiomira dubrovinae[100]

Sp. nov

Valid

Nabozhenko & Bukejs

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

An Alleculinae darkling beetle.

Aulacoscelis santanensis[101]

Sp. nov

In press

Legalov

Early Cretaceous
(Aptian-Albian)

Santana Group

Brazil

An orsodacnid leaf beetle.

Baltocar sontagae[102]

Sp. nov

Valid

Bukejs & Legalov

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A Sayrevilleinae tooth-nosed snout weevil.

Baltosidis[103]

Gen. et 3 sp. nov

Valid

Szawaryn

Eocene

Baltic amber

Europe

A Microweiseini Microweiseinae lady beetle.
The type species is B. damzeni;
genus also includes B. damgaardi and B. szadziewskii.

Bronchotibia[104]

Gen. et sp. nov

In press

Poinar & Legalov

Miocene
(Burdigalian)

Dominican amber

Dominican Republic

A Lymantini Molytinae weevil.
The type species is B. adunatus.

Cathartosilvanus perkovskyi[105]

Sp. nov

Valid

Alekseev & Bukejs

Late Eocene

Rovno amber

Ukraine

A silvanid beetle.

Ceutorhynchus zerovae[106]

Sp. nov

Valid

Legalov et al.

Late Eocene

Rovno amber

Ukraine

A weevil.

Coslonatus[107]

Gen. et sp. nov

Valid

Li & Cai

Cenomanian

Burmese amber

Myanmar

A Colydiinae? Zopheridae tenebrionoid beetle.
The type species is C. rasnitsyni.

Cretaidgia[108]

Gen. et sp. nov

In press

Zhao, Liu & Yu

Late Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A prionocerid beetle.
The type species is C. burmensis.

Diapus ethiopicus[109]

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

Ethiopian amber

Ethiopia

A Platypodinae pinhole borer ambrosia weevil.

Diapus resinae[109]

Sp. nov

In press

Solórzano-Kraemer et al.

Miocene

Zhangpu amber

China

A Platypodinae pinhole borer ambrosia weevil.

Dorytomus mikhailovi[110]

Sp. nov

Valid

Legalov, Nazarenko & Perkovsky

Late Eocene

Rovno amber

Ukraine

A species of Dorytomus Curculioninae weevil.

Electroxylita[111]

Gen. et sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

A false darkling beetle.
The type species is E. chronographica.

Eurocrossotus[112]

Gen. et sp. nov

Valid

Vitali

Eocene

Baltic amber

Europe (Baltic Sea region)

A longhorn beetle belonging to the tribe Crossotini. The type species is E. alekseevi.

Glaesotropis rohdendorfi[113]

Sp. nov

Valid

Legalov, Nazarenko & Perkovsky

Eocene
(Priabonian)

Rovno amber

Ukraine

A zygaenodinin anthribine anthribid fungus weevil.

Glesoconomorphus ekaterinae[114]

Sp. nov

Valid

Telnov et al.

Late Eocene
Priabonian

Rovno amber

Ukraine

A mycterid palm beetle.

Gracilenticrus[115]

Gen. et sp. nov

Valid

Yu, Kolibáč & Ślipiński
in Yu et al.

Late Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A lophocaterid beetle.
The type species is G. burmiticus.

Heterhelus buzina[116]

Sp. nov

Valid

Kupryjanowicz, Lyubarsky & Perkovsky

Eocene

Rovno amber

Ukraine

A Kateretid short-winged flower beetle. Originally described as a species of Heterhelus, but Kirejtshuk, Jenkins Shaw & Smirnov (2023) transferred it to the genus Antirhelus.[117]

Isomira lobanovi[118]

Sp. nov

Valid

Nabozhenko & Bukejs

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

An Isomira species claw comb darkling beetle.

Kaszubia[119]

Gen. et sp. nov

Valid

Legalov, Kupryjanowicz & Perkovsky

Late Eocene
Priabonian

Baltic amber

Poland

A cossonine weevil.
The type species is K. wanati.

Kulindrobor[120]

Gen. et 2 sp. nov

Valid

Tihelka et al.

Late Cretaceous
Cenomanian

Burmese amber

Myanmar

A tenebrionoid beetle of uncertain phylogenetic placement.
Genus includes new species K. enigmaticus and K. magnus.

Lobanoviella[121]

Gen. et sp. nov

In press

Kirejtshuk & Reid

Eocene

Baltic amber

Europe

A lobanovielline palophagine megalopodid leaf beetle.
The type species is L. andreyi.

Madelinia capillata[111]

Sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

A false darkling beetle.

Melanosiagon[122]

Gen. et sp. nov

Valid

Batelka & Prokop

Late Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A wedge-shaped beetle.
The type species is M. serraticornis.

Metaclisa ottoi[123]

Sp. nov

Valid

Nabozhenko, McKellar & Bukejs

Late Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

AMetaclisa species metaclisine darkling beetle.

Microscapha kugelanni[111]

Sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

A false darkling beetle.

Microtrogossita[124]

Gen. et sp. nov

Valid

Li & Cai
in Li et al.

Cretaceous
Cenomanian

Burmese amber

Myanmar

A Trogossitid bark-gnawing beetle.
The type species is M. qizhihaoi.

Monolepta rappsilberi[125]

Sp. nov

Valid

Bukejs et al.

Late Eocene
Priabonian

Bitterfeld amber

Germany

A Monolepta species skeletonizing leaf beetle.

Neolatitergum[126]

Nom. nov

Valid

Zhang

Jurassic

Jiulongshan Formation

China

A Trogossitid bark-gnawing beetle.
A replacement name for "Latitergum" Yu et al. (2014).

Oedemera girulskii[127]

Sp. nov

Valid

Szawaryn, Sontag & Kubisz

Late Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

An Oedemera species false blister beetle.

Pacyclaxyra[128]

Gen. et sp. nov

Valid

Tihelka, Huang & Cai

Late Cretaceous
Cenomanian

Burmese amber

Myanmar

A sooty mould beetle.
The type species is P. azari.

Palaeatalasis[129]

Gen. et sp. nov

Valid

Legalov

Eocene
(Ypresian)

Green River Formation

United States
( Utah)

A Megamerini Sagrinae leaf beetle.
The type species is P. monrosi.

Palaeoanoplus[130]

Trib., Gen., et sp. nov

Valid

Legalov

Late Eocene
Priabonian

Baltic amber

Europe

A Palaeoanoplini Curculioninae weevil.
The type species is P. horridus.

Paleopegorhinus[131]

Gen. et sp. nov

Valid

Poinar, Brown & Legalov

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A Chilecarinae new york weevil.
The type species is P. micrommatus.

Palaeopeplus[132]

Gen. et sp. nov

Valid

Powell & Cline

Miocene
(Burdigalina)

Dominican amber

Dominican Republic

A Cillaeinae sap beetle.
The type species is P. cascus.

Pelretes[133][134][135]

Gen. et sp. nov

Valid

Tihelka et al.

Cretaceous

Burmese amber

Myanmar

A short-winged flower beetle.
The type species is P. vivificus .

Phloiotrya inmarinata[111]

Sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

A Phloiotrya species false darkling beetle.

Protodasytes[136]

Gen. et sp. nov

Valid

Tihelka et al.

Late Cretaceous
(Cenomanian)

Charentese amber

France

A soft-wing flower beetle.
The type species is P. cretaceus.

Protomauroania[137]

Gen. et 2 sp. nov

Valid

Tshernyshev in Tshernyshev & Shcherbakov

Late Eocene

Baltic amber
Rovno amber

Baltic Sea region
Ukraine

A Dasytinae malachite beetle.
The type species P. chaetophorata
Genus also includes P. mikhailovi.[138]

Protostomopsis[139]

Gen. et sp. nov

Valid

Bukejs, Ślipiński & Alekseev
in Bukejs et al.

Eocene

Baltic amber

Poland
Russia
( Kaliningrad Oblast)

A cerylonid beetle. The type species is P. pandema.

Pseudomesauletes lobanovi[102]

Sp. nov

Valid

Bukejs & Legalov

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A Rhynchitinae tooth-nosed snout weevil.

Pulchritudo[140]

Gen. et sp. nov

Valid

Krell & Vitali

Eocene
(Ypresian)

Green River Formation

United States
( Colorado)

A Sagrinae leaf beetle.
The type species is P. attenboroughi.

Rhamphophorus[141]

Trib., gen. et sp. nov

Valid

Poinar & Brown

Cretaceous
Cenomanian

Burmese amber

Myanmar

A Rhamphophorini Cimberidinae pine flower weevil.
The type species is R. legalovii.

Salignacicola[122]

Gen. et comb. nov

Valid

Batelka & Prokop

Late Cretaceous
(Cenomanian)

|  |

France

A wedge shaped beetle.
The type species is "Macrosiagon" ebboi (2004).

Symphora glaesonauta[111]

Sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

A false darkling beetle.

Symphora pollocki[111]

Sp. nov

Valid

Alekseev & Bukejs

Middle Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

A false darkling beetle.

Teredolaemus primigenius[142]

Sp. nov

Valid

Alekseev et al.

Late Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

A teredid beetle.

Toxorhynchus michalskii[143]

Sp. nov

Valid

Legalov

Late Eocene
Priabonian

Baltic amber

Europe

An Apioninae straight-snouted weevil.

Trematosphindus[144]

Gen. et sp. nov

Valid

Li & Cai
in Li et al.

Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A slime mold beetle.
The type species is T. newtoni

Zaiwa[145]

Gen. et sp. nov

Valid

Lyubarsky et al.

Cretaceous
Cenomanian

Burmese amber

Myanmar

A lophocaterid beetle.
The type species is Z. pankowskiorum.

Zherikhiniellus[143]

Gen. et sp. nov

Valid

Legalov

late Eocene
Priabonian

Baltic amber

Europe

A Nanophyinae straight-snouted weevil.
The type species is Z. rarus.

##### Cucujiformia research[edit]

* A piece of Burmese amber preserving the complete series of free-living stages (except of the last larval instar) of a Cretaceous parasitoid beetle Paleoripiphorus is described by Batelka, Engel & Prokop (2021).[146]

##### Elateriformia[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Amberophytum maculatum[147]

Sp. nov

Valid

Poinar, Vega & Legalov

Late Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A cerophytid rare click beetle.

Anoeuma[148]

Gen. et sp. nov

Valid

Li, Kundrata & Cai in Li et al.

Cretaceous (Albian to Cenomanian)

Burmese amber

Myanmar

An elateroid click beetle of uncertain phylogenetic placement.
The type species is A. lawrencei.

Arturmiles[149]

Gen. et sp. nov

Valid

Fanti

Eocene

Prussian Formation
(Baltic amber)

Poland

A soldier beetle belonging to the tribe Cantharini. The type species is A. pankowskiarum.

Balistica vetai[150]

Sp. nov

Valid

Muona

Eocene

Baltic amber

Europe

A Dirhagini Melasinae false click beetle.

Baltodascillus[151]

Gen. et sp. nov

Valid

Kundrata et al.

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A dascillid beetle.
The type species is B. serraticornis.

Burmagrilus[152]

Gen. et sp. nov

Valid

Jiang, Song & Wang in Jiang et al.

Cretaceous

Burmese amber

Myanmar

A metallic wood-boring beetle.
The type species is B. cretacus.

Burmomiles bilineatimaculatus[153]

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

Myanmar

A soldier beetle. Fanti & Müller (2022) considered it to be a species belonging to the genus Elektrokleinia.[154]

Burmomiles laticollis[153]

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

Myanmar

A soldier beetle.

Burmomiles oblongoculus[153]

Sp. nov

In press

Yang et al.

Cretaceous

Burmese amber

Myanmar

A soldier beetle. Fanti & Müller (2022) considered it to be a species belonging to the genus Elektrokleinia.[154]

Captopus[155]

Gen. et sp. nov

Valid

Li & Cai in Li, Huang & Cai

Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A throscid beetle.
The type species is C. depressiceps.

Chelonarium andabata[156]

Sp. nov

Valid

Alekseev & Bukejs in Alekseev et al.

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A turtle beetle.

Chelonarium dingansich[156]

Sp. nov

Valid

Alekseev & Bukejs in Alekseev et al.

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A turtle beetle.

Chelonarium dominicanum[157]

Sp. nov

Valid

Poinar & Háva

Burdigalian

Dominican amber

Dominican Republic

A turtle beetle.

Coomanius balticus[150]

Sp. nov

Valid

Muona

Eocene

Baltic amber

Europe

A Nematodini Macraulacinae false click beetle.

Cretocantharis[158]

Gen. et sp. nov

Valid

Hsiao et al.

Late Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A soldier beetle.
The type species is C. veda.

Cretopachyderes[159]

Gen. et sp. nov

Valid

Kundrata et al.

Cretaceous
Cenomanian

Burmese amber

Myanmar

An Agrypninae click beetle.
The type species is C. burmitinus.

Cretophengodes[160]

Fam. nov., Gen. et sp. nov

Valid

Li, Kundrata, Tihelka & Cai in Li et al.

Cretaceous (Albian or Cenomanian)

Burmese amber

Myanmar

A cretophengodid elateroid beetle.
The type species is C. azari.

Dyscharachthis frickae[150]

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

Europe (Baltic Sea region)

A Dyscharachthini Eucneminae false click beetle.

Electrothroscus[155]

Gen. et sp. nov

Valid

Li & Cai

Cretaceous
Cenomanian

Burmese amber

Myanmar

A throscid false click beetle.
The type species is E. yanpingae.

Entomophthalmus antennalis[150]

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe (Baltic Sea region)

A Dirhagini Melasinae false click beetle.

Erdaia bispinulosa[150]

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

An Orodotini? Macraulacinae false click beetle.

Erdaia longicornis[150]

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

An Orodotini? Macraulacinae false click beetle.

Erdaia longitarsis[150]

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

An Orodotini? Macraulacinae false click beetle.

Euryptychoides[150]

Gen. et sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

An Euryptychini Macraulacinae false click beetle.
The type species is E. magnus.

Euryptychus decoratus[150]

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

An Euryptychini Macraulacinae false click beetle.

Euryptychus excavatus[150]

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

An Euryptychini Macraulacinae false click beetle.

Euryptychus helmwigae[150]

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

An Euryptychini Macraulacinae false click beetle.

Euryptychus sieglindae[150]

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

An Euryptychini Macraulacinae false click beetle.

Euryptychus waltrautae[150]

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

An Euryptychini Macraulacinae false click beetle.

Hukawngichthyurus maha[158]

Sp. nov

Valid

Hsiao et al.

Late Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A soldier beetle.

Isorhipis balticus[150]

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

Europe

An Isorhipis species melasine false click beetle

Lycocerus crassepunctatus[161]

Sp. nov

Valid

Kazantsev

Late Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

A soldier beetle.

Malthodes aliger[161]

Sp. nov

Valid

Kazantsev

Late Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

A soldier beetle

Malthodes headsi[162]

Sp. nov

Valid

Fanti

Late Eocene
Priabonian

Baltic amber

Poland

A soldier beetle.

Malthodes jaredi[162]

Sp. nov

Valid

Fanti

Late Eocene
Priabonian

Baltic amber

Poland

A soldier beetle, a species of Malthodes.

Malthodes vladiceniensis[163]

Sp. nov

Valid

Pintilioaie, Fanti & Ionesi

Middle Miocene

Bârnova-Muntele Formation

Romania

A soldier beetle.

Microrhagus (Emyirhagus) vetai[150]

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

Europe

A Microrhagusspecies melasine false click beetle

Murcybolus[164]

Gen. et sp. nov

Valid

Li & Cai
in Li et al.

Cretaceous
Cenomanian

Burmese amber

Myanmar

A Burmolycini dexorine net-winged beetle.
The type species is M. longiantennus.

Nematodes schwertleitae[150]

Sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

A Nematodini Macraulacinae false click beetle.

Neocharoides[150]

Gen. et sp. nov

Valid

Muona

Middle Eocene
Priabonian

Baltic amber

Europe

A Nematodini? Macraulacinae false click beetle.
The type species is N. balticus.

Palaeocantharis[158]

Gen. et sp. nov

Disputed

Hsiao et al.

Late Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A soldier beetle. The type species is P. panna.
Fanti & Müller (2022) considered the genus Palaeocantharis to a be a junior synonym of the genus Elektrokleinia, though the authors maintained P. panna as a distinct species within the latter genus.[154]

Podistra guthriei[162]

Sp. nov

Valid

Fanti

Late Eocene
Priabonian

Baltic amber

Poland

A soldier beetle.

Poecilochrus balticus[150]

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

Europe

A Eucnemini Eucneminae false click beetle.

Potergites brittoni[150]

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

Europe

A Dendrocharini Eucneminae false click beetle.

Pseudomataeopsephus[165]

Gen. et sp. nov

In press

Li, Huang & Cai

Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A water-penny beetle.
The type species is P. burmensis.

Pseudopactopus[155]

Gen. et sp. nov

Valid

Li & Cai
in Li, Huang & Cai

Cretaceous
( Cenomanian)

Burmese amber

Myanmar

A throscid false click beetle.
The type species is P. robustus.

Ptilodactyla eocenica[166]

Sp. nov

Valid

Kundrata, Bukejs & Blank
in Kundrata et al.

Late Eocene
Priabonian

Baltic amber

Russia
( Kaliningrad Oblast)

APtilodactyla species toe winged beetle.

Rhacopus balticus[150]

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

Europe

A Rhacopus species eucnemid false click beetle

Rhagomicrus balticus[150]

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

Europe

A Dirhagini Melasinae false click beetle.

Rheanischia[167]

Gen. et sp. nov

|  |

Li, Chang & Muona

Early Cretaceous
(Hauterivian to Aptian)

Yixian Formation

China

An Anischiinae false click beetle.
The type species is R. brevicornis.

Silis curleri[168]

Sp. nov

Valid

Fanti & Pankowski

Burdigalian

Dominican amber

Dominican Republic

A species of Silis.

Silis hegnai[168]

Sp. nov

Valid

Fanti & Pankowski

Burdigalian

Dominican amber

Dominican Republic

A species of Silis.

Xylophilus balticus[150]

Sp. nov

Valid

Muona

Late Eocene
Priabonian

Baltic amber

Europe

A Xylophilus species eucnemid false click beetle.

###### Elateriformia research[edit]

* A study on the anatomy and phylogenetic relationships of Archaeolus funestus is published by Li et al. (2021).[169]
* A study on the morphological diversity of larvae of extant and fossil members of the family Brachypsectridae is published by Haug et al. (2021).[170]

##### Scarabaeiformia[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Goriresina[171]

Gen. et sp. nov

Valid

Matalin, Perkovsky & Vasilenko

Late Eocene
Priabonian

Rovno amber

Ukraine

A tiger beetle.
The type species is G. fungifora.

Onthophagus pilauco[172]

Sp. nov

Valid

Tello et al.

Late Pleistocene

Pilauco Bajo

Chile

A scarab beetle.

##### Staphyliniformia[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Acritus sutirca[173]

Sp. nov

Valid

Alekseev & Bukejs

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A species of Acritus.

Bacanius gorskii[174]

Sp. nov

Valid

Alekseev & Bukejs

Eocene

Baltic amber

Poland

A species of Bacanius.

Burmagluta[175]

Gen. et sp. nov

Valid

Yin & Cai

Cretaceous

Burmese amber

Myanmar

A Brachyglutini Pselaphinae rove beetle.
The type species is B. rougemonti.

Carinumerus[176]

Gen. et sp. nov

Valid

Caterino

Cretaceous

Burmese amber

Myanmar

An Onthophilinae clown beetle.
The type species is C. maddisoni.

Charhyphus serratus[177]

Sp. nov

In press

Yamamoto & Shavrin

Eocene

Baltic amber

Europe

A Phloeocharinae rove beetle.

Cretosaja[178]

Gen. et sp. nov

In press

Sohn & Nam

Early Cretaceous

Jinju Formation

South Korea

A carrion beetle.
The type species is C. jinjuensis.

Druantia[176]

Gen. et sp. nov

Valid

Caterino

Cenomianian

Burmese amber

Myanmar

A Dendrophilinae clown beetle.
The type species is D. aeterna.

Euconnus nathani[179]

Sp. nov

Valid

Jałoszyński & Perkovsky

Eocene

Rovno amber

Ukraine

A Euconnus species Scydmaeninae rove beetle.

Eusphalerum bukejsi[180]

Sp. nov

Valid

Shavrin in Shavrin & Kairišs

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A Eusphalerum species Omaliinae rove beetle.

Megabythinus[181]

Gen. et sp. nov

Valid

Yin, Zhao & Cai

Cretaceous
Cenomanian

Burmese amber

Myanmar

A Bythinini Pselaphinae rove beetle.
The type species is M. spinitibialis.

Paroxyporus[182]

Gen. et sp. nov

Valid

Cai

Early Cretaceous

Yixian Formation

China

A rove beetle.
The type species is P. fossilis.

Phasmister[176]

Gen. et sp. nov

Valid

Caterino

Cretaceous

Burmese amber

Myanmar

An Onthophilinae clown beetle.
The type species is P. cristatus.

Ponohydrochus[183]

Gen. et sp. nov

Valid

Prokin & Strelnikova

Early Cretaceous
(Hauterivian)

|  |

Russia

A water scavenger beetle.
The type species is P. buryaticus.

Proptomaphaginus alleni[184]

Sp. nov

Valid

Perreau & Haelewaters
in Perreau, Haelewaters & Tafforeau

Miocene
Burdigalian

Dominican amber

Dominican Republic

A Ptomaphagini round fungus beetle.

Trichophya minor[185]

Sp. nov

Valid

Yamamoto & Newton

Cretaceous
(Cenomanian)

Burmese amber

Myanmar

A Trichophya species Trichophyinae rove beetle.

Yethiha[176]

Gen. et sp. nov

Valid

Caterino

Cretaceous

Burmese amber

Myanmar

A Dendrophilinae clown beetle.
The type species is Y. peregrina.

#### Coleopteran research[edit]

* A study on the evolution of beetles from the Early Permian to Middle Triassic, evaluating the impact of the Permian–Triassic extinction event on beetle diversity, is published by Zhao et al. (2021).[186]
* Revision of Notocupes undatabdominus and N. ? multituberatus from the Lower Cretaceous of South China is published by Li, Huang & Cai (2021), who argue that both species do not belong to the genus Notocupes, transfer N. ? multituberatus to the genus Lasiosyne, and consider "N." undatabdominus to be likely a member of the family Artematopodidae.[187]
* The oldest solid wood-borer larvae reported to date (belonging to the families Buprestidae and possibly Cerambycidae) are described from the Cretaceous amber from Myanmar by Haug et al. (2021).[188]
* Mitochondrial genomes and nuclear ribosomal DNA of beetles, in some cases identifiable to species, are recovered from pack rat middens from California (United States) and Baja California (Mexico) dating up to ~ 34,355 years before present by Smith et al. (2021).[189]

## Clade Dictyoptera[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Afrophaga[190]

Gen. et sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

|  |

Algeria

A cockroach belonging to the family Corydiidae. Genus includes new species A. extincta.

Alienopterix mlynskyi[191]

Sp. nov

In press

Sendi in Vršanský et al.

Cretaceous

Burmese amber

Myanmar

An alienopterid.

Alienopterix smidovae[191]

Sp. nov

In press

Hinkelman in Vršanský et al.

Cretaceous

Burmese amber

Myanmar

An alienopterid.

Arceotermes[192]

Fam. et gen. et sp. nov

Valid

Engel & Jiang in Jiang et al.

Cretaceous

Burmese amber

Myanmar

A termite belonging to the new family Arceotermitidae. Genus includes new species A. hospitis.

Caligoptera[193]

Gen. et sp. nov

In press

Sendi

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A cockroach belonging to the family Ectobiidae. Genus includes new species C. hinkelmani.

Chuanblatta[194]

Gen. et 2 sp. nov

In press

Liang et al.

Middle Jurassic

Jiulongshan Formation

China

A cockroach belonging to the family Raphidiomimidae. Genus includes new species C. huntianlingensis and C. lata.

Cuniculoblatta[195]

Gen. et sp. nov

In press

Hinkelman

Cretaceous

Burmese amber

Myanmar

A cockroach. Genus includes new species C. brevialata .

Ectoovia[193]

Gen. et sp. nov

In press

Sendi

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A cockroach belonging to the family Ectobiidae. Genus includes new species E. protecta.

? Elisama algeriaensis[190]

Sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

|  |

Algeria

A member of the family Blattulidae.

Eminespina[196]

Gen. et sp. nov

Valid

Chen, Zhang Shi

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

An alienopterid. Genus includes new species E. burma.

Fractalia[197]

Fam. et gen. et 2 sp. nov

Valid

Hinkelman & Vršanský in Vršanský et al.

Late Jurassic–Late Cretaceous (Kimmeridgian–Cenomanian)

Burmese amber

Germany
Kazakhstan
Myanmar

A member of Corydioidea, the type genus of the new family Fractaliidae Vršanský & Hinkelman. The type species is F. articulata Hinkelman; genus also includes new species F. aristovi Vršanský.

Fragosublatta[198]

Gen. et sp. nov

Valid

Chen, Shih & Ren in Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A cockroach belonging to the family Corydiidae. The type species is F. pectinata.

Glyptotermes abyssinicus[199]

Sp. nov

Valid

Bouju, Jouault & Perrichot

Early Miocene

Ethiopian amber

Ethiopia

A termite belonging to the family Kalotermitidae.

Kinneyblatta[200]

Gen. et sp. nov

Valid

Schneider et al.

Carboniferous (Kasimovian)

Atrasado Formation

United States
( New Mexico)

A member of Blattodea belonging to the family Spiloblattinidae. The type species is K. huberi.

Kriedoblatta[190]

Gen. et sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

|  |

Algeria

A member of the family Liberiblattinidae. Genus includes new species K. gondwanensis.

? Kurablattina samsonovi[190]

Sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

|  |

Algeria

A member of the family Liberiblattinidae.

Labradormantis[201]

Gen. et sp. nov

Valid

Demers‐Potvin et al.

Late Cretaceous (Cenomanian)

Redmond Formation

Canada

A praying mantis belonging to the family Baissomantidae. Genus includes new species L. guilbaulti.

Latiblattella basaltica[202]

Sp. nov

In press

Vršanský, Poschmann & Vidlička

Late Oligocene

|  |

Germany

A species of Latiblattella.

Latiblattella karlgruberi[193]

Sp. nov

In press

Sendi

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

|  |

Laticephalana[203]

Gen. et sp. nov

Valid

Luo et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Umenocoleidae. Genus includes new species L. liuyani.

Lukotermes[204]

Gen. et sp. nov

Valid

Perkovsky & Nel

Eocene (Priabonian)

Rovno amber

Ukraine

A termite belonging to the family Rhinotermitidae. Genus includes new species L. milescaput.

Meiatermes cretacicus[205]

Sp. nov

In press

Bezerra et al.

Early Cretaceous

Crato Formation

Brazil

A termite.

Meloblatta[190]

Gen. et sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

|  |

Algeria

A member of the family Mesoblattinidae. Genus includes new species M. brezinica.

Milesitermes[206]

Gen. et sp. nov

Valid

Jouault et al.

Cretaceous

Burmese amber

Myanmar

A termite belonging to the family Mastotermitidae. Genus includes new species M. engeli.

Miniblattina[207]

Gen. et sp. nov

In press

Sendi

Early Cretaceous (Barremian)

Lebanese amber

Lebanon

A cockroach belonging to the family Liberiblattinidae. Genus includes new species M. libera.

Mongolblatta sendii[208]

Sp. nov

In press

Hinkelman

Cretaceous

Burmese amber

Myanmar

A member of the family Mesoblattinidae.

Morphna cenozoica[209]

Sp. nov

In press

Šmídová, Vidlička & Wedmann

Eocene

Messel pit

Germany

A cockroach belonging to the family Blaberidae.

Morphna cretacica[209]

Sp. nov

In press

Šmídová, Vidlička & Wedmann

Cretaceous

Burmese amber

Myanmar

A cockroach belonging to the family Blaberidae.

Nadveruzenie[191]

Gen. et sp. nov

In press

Vršanský, Hinkelman & Sendi in Vršanský et al.

Cretaceous

Burmese amber

Myanmar

An alienopterid. Genus includes new species N. postava.

Nodosigalea simplivena[210]

Sp. nov

In press

Šmídová

Cretaceous

Burmese amber

Myanmar

A cockroach belonging to the family Corydiidae.

Ocelloblattula margarita[197]

Sp. nov

Valid

Koubová & Vršanský in Vršanský et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Corydioidea.

Olenablatta[210]

Gen. et sp. nov

In press

Šmídová

Cretaceous

Burmese amber

Myanmar

A cockroach belonging to the family Olidae. Genus includes new species O. vrsanskyi.

Otazka[190]

Gen. et sp. nov

In press

Vršanský et al.

Cretaceous (Hauterivian?)

|  |

Algeria

A member of the family Mesoblattinidae. Genus includes new species O. systematicka.

Pabuonqed magna[211]

Sp. nov

In press

Song et al.

Cretaceous

Burmese amber

Myanmar

A member of Mastotermitoidea belonging to the family Pabuonqedidae.

Paekthoblatta[212]

Gen. et sp. nov

Valid

So et al.

Early Cretaceous

Sinuiju Formation

North Korea

A member of the family Raphidiomimidae. Genus includes new species P. coreanica.

Periplaneta (?) perialla[213]

Sp. nov

Valid

Anisyutkin & Perkovsky

Eocene

Rovno amber

Ukraine

A cockroach, possibly a species of Periplaneta.

Petropterix fukuiensis[214]

Sp. nov

In press

Oyama, Yukawa & Imai

Early Cretaceous

Kitadani Formation

Japan

A member of the family Umenocoleidae.

Praeblattella arcuata[214]

Sp. nov

In press

Oyama, Yukawa & Imai

Early Cretaceous

Kitadani Formation

Japan

A member of the family Mesoblattinidae.

Praeblattella inexpecta[214]

Sp. nov

In press

Oyama, Yukawa & Imai

Early Cretaceous

Kitadani Formation

Japan

A member of the family Mesoblattinidae.

Pseudomantina[197]

Gen. et 2 sp. nov

Valid

Sendi & Vršanský in Vršanský et al.

Late Jurassic–Early Cretaceous (Kimmeridgian–Barremian)

Lebanese amber

Kazakhstan
Lebanon

A member of the family Liberiblattinidae. The type species is P. occisor Sendi; genus also includes new species P. nigroalba Vršanský.

Reticulitermes grimaldii[206]

Sp. nov

Valid

Jouault et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A termite, a species of Reticulitermes.

Sajda[190]

Gen. et sp. nov

Junior homonym

Vršanský et al.

Cretaceous (Hauterivian?)

|  |

Algeria

A member of Holocompsinae. Genus includes new species S. equatorialis. The generic name is preoccupied by Sajda Dworakowska (1981); Jiang, Xing & Li (2023) coined a replacement name Vrsanskysajda.[215]

Spinaeblattina baekthoensis[216]

Sp. nov

Valid

So et al.

Early Cretaceous

Sinuiju Formation

North Korea

A member of the family Mesoblattinidae.

Stavba delicata[207]

Sp. nov

In press

Sendi

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A cockroach belonging to the family Liberiblattinidae.

Stavba magnoculara[207]

Sp. nov

In press

Sendi

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A cockroach belonging to the family Liberiblattinidae.

Tanytermitalis[192]

Gen. et sp. nov

Valid

Engel & Cai in Jiang et al.

Cretaceous

Burmese amber

Myanmar

A termite belonging to the family Tanytermitidae. Genus includes new species T. philetaerus.

### Dictyopteran research[edit]

* A study on the phylogenetic placement of the family Umenocoleidae, based on data from a new specimen of Umenocoleus sinuatus from the Lower Cretaceous Zhonggou Formation (China), is published by Liuo et al. (2021).[217]
* Description of new alienopterid nymphs from the Cretaceous Burmese amber, and a study on life history and affinities of alienopterids, is published by Luo et al. (2021).[218]
* A study on the sensory organs of Huablattula hui, and on their implications for the knowledge of the ecology of this cockroach, is published by Taniguchi et al. (2021).[219]

## Clade †Glosselytrodea[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

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|
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|
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|
|

Moltenojurina[220]

Gen. et sp. nov

Valid

Béthoux & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A polycytellid Glosselytrodean.
The type species is M. parva.

Polycytella rasnitsyni[220]

Sp. nov

Valid

Béthoux & Anderson

Late Triassic (Carnian)

Molteno Formation

South Africa

A polycytellid Glosselytrodea.

## Hymenopterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Aethemelikertes[221]

Gen. et sp. nov

Valid

Engel in Engel & Davis

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A bee belonging to the tribe Melikertini. The type species is A. emunctorii.

Alivespa gracilenta[222]

Sp. nov

In press

Wu et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Vespidae.

Allommation[223]

Fam. nov., Gen. et 3 sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Apoidea in the new family Allommationidae. Genus includes new species A. procerum, A. elongatum and A. grande.

Amelikertotes[221]

Gen. et comb. nov

Valid

Engel in Engel & Davis

Eocene

Baltic amber
Bitterfeld amber

Germany
Russia
( Kaliningrad Oblast)

A bee belonging to the tribe Melikertini. The type species is "Melikertes" clypeatus Engel (2001).

Ammosphecium[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Crabroninae. Genus includes new species A. diabolicum.

Anochetus miosumatrensis[224]

Sp. nov

Valid

Ngô-Muller, Garrouste & Nel in Ngô-Muller et al.

Miocene

|  |

Indonesia

An ant, a species of Anochetus.

Archaeovespa incompleta[222]

Sp. nov

In press

Wu et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Vespidae.

Archeonesia[225]

Gen. et sp. nov

Valid

Tribull, Pankowski & Colombo

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A Pristocerinae Bethylid.
The type species is A. eocena.

|  Archeonesia eocena

Arkadiserphus[226]

Gen. et sp. nov

Valid

Rasnitsyn

Late Jurassic

Karabastau Formation

Kazakhstan

A member of Proctotrupoidea belonging to the family Peleserphidae. The type species is A. leleji.

Ascogaster (Syntaphus) latitibialis[227]

Sp. nov

Valid

Belokobylskij et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Braconidae belonging to the subfamily Cheloninae.

Aulacicosmus[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Heterogynaidae. Genus includes new species A. setosus.

Austroplebeia fujianica[228]

Sp. nov

Valid

Engel in Engel et al.

Miocene (Langhian)

Zhangpu amber

China

A stingless bee, a species of Austroplebeia.

Burmabracon[229]

Gen. et 2 sp. nov

Valid

Li et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Braconidae belonging to the subfamily Protorhyssalinae. Genus includes new species B. gracilens and B. grossus.

Burmapyris[230]

Gen. et sp. nov

Valid

Jouault, Perrichot & Nel

Cretaceous

|  |

Myanmar

A member of the family Bethylidae belonging to the subfamily Lancepyrinae. Genus includes new species B. azevedoi.

Burmoxyela[231]

Gen. et sp. nov

Valid

Zheng et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A sawfly belonging to the family Syspastoxyelidae. Genus includes new species B. lii.

Burmusculus angustus[232]

Sp. nov

In press

Li et al.

Cretaceous

Burmese amber

Myanmar

A member of Apocrita belonging to the family Burmusculidae.

Buserphites[233]

Gen. et 2 sp. nov

In press

Herbert & McKellar

Cretaceous

Burmese amber

Myanmar

A member of the family Serphitidae. Genus includes new species B. applanatus and B. myanmarensis.

Chainochora[234]

Gen. et sp. nov

Valid

Chen, van Achterberg & Hong

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Braconidae belonging to the subfamily Protobraconinae. The type species is C. syntoma.

Chlorepyris deploegi[235]

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Rovno amber

Ukraine

A member of the family Bethylidae.

Chlorepyris engeli[235]

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

Russia

A member of the family Bethylidae.

Chlorepyris hopei[235]

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

Russia

A member of the family Bethylidae.

Chlorepyris jouaulti[235]

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Rovno amber

Ukraine

A member of the family Bethylidae.

Chlorepyris mckellari[235]

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Rovno amber

Ukraine

A member of the family Bethylidae.

Chlorepyris meunieri[235]

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

Russia

A member of the family Bethylidae.

Chlorepyris neli[235]

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber
Rovno amber

Russia
Ukraine

A member of the family Bethylidae.

Chlorepyris perrichoti[235]

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

Russia

A member of the family Bethylidae.

Christophus[236]

Gen. et 2 sp. nov

In press

Brazidec et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Diapriidae. Genus includes new species C. myanmarensis and C. ohmkuhnlei.

Cretastatus[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Astatinae. Genus includes new species C. longevus.

Cretorussus[237]

Gen. et sp. nov

In press

Jouault, Perrichot & Nel

Cretaceous

Burmese amber

Myanmar

A member of Orussoidea. Genus includes new species C. vilhelmseni.

Curiosivespa sennlaubi[238]

Sp. nov

Valid

Jouault, Rasnitsyn & Nel in Jouault et al.

Early Cretaceous

Crato Formation

Brazil

A member of the family Vespidae.

Diameneura[239]

Gen. et sp. nov

In press

Santer & Álvarez-Parra in Santer et al.

Early Cretaceous (Albian)

Escucha Formation

Spain

A member of the family Spathiopterygidae. The type species is D. marveni.

Drymomyrmex rasnitsyni[240]

Sp. nov

Valid

Radchenko

Late Eocene

Rovno amber

Ukraine

An ant belonging to the subfamily Formicinae.

Elasmosomites arkadyleleji[241]

Sp. nov

Valid

Belokobylskij in Belokobylskij et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Braconidae belonging to the subfamily Euphorinae.

Electrofoenops exaltatus[242]

Sp. nov

In press

Li et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Aulacidae.

Embolemus brachypterus[243]

Sp. nov

Valid

Olmi et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Embolemidae.

Embolemus micropterus[243]

Sp. nov

Valid

Olmi et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Embolemidae.

Eopelecinus marechali[244]

Sp. nov

Valid

Jouault

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Pelecinidae.

Eoteredon[245]

Gen. et sp. nov

Valid

Archibald, Aase & Nel

Early Eocene

Green River Formation

United States
( Wyoming)

A horntail. Genus includes new species E. lacoi.

Ephedrus rasnitsyni[246]

Sp. nov

Valid

Davidian & Kaliuzhna in Davidian, Kaliuzhna & Perkovsky

Middle Eocene

Sakhalinian amber

Russia

A member of the family Braconidae belonging to the subfamily Aphidiinae.

Exilaulacus advenus[242]

Sp. nov

In press

Li et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Aulacidae.

Exilaulacus eximius[242]

Sp. nov

In press

Li et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Aulacidae.

Formica ribbeckei[247]

Sp. nov

Valid

Radchenko & Perkovsky in Radchenko, Perkovsky & Vasilenko

Late Eocene

Rovno amber

Ukraine

An ant, a species of Formica.

Glenocephalus[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Apoidea in the new family Cirrosphecidae. Genus includes new species G. mandibularis.

Gloxinius[235]

Gen. et comb. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

Russia

A member of the family Bethylidae; a new genus for "Epyris" bifossatus Brues (1939).

Grandixyela[248]

Gen. et sp. nov

Valid

Zheng in Zheng et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Syspastoxyelidae. The type species is G. rasnitsyni.

Haidomelikertes[221]

Gen. et sp. nov

Valid

Engel in Engel & Davis

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A bee belonging to the tribe Melikertini. The type species is H. uraeus.

Haptodioctes[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Apoidea in the new family Cirrosphecidae. Genus includes new species H. apiformis.

Helosericus[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Psenini. Genus includes new species H. verrucosus.

Heteroichneumon[249]

Gen. et sp. nov

Valid

Kopylov, Zhang & Zhang

Cretaceous

Burmese amber

Myanmar

A member of the family Ichneumonidae. The type species is H. rasnitsyni.

Heteropimpla pulverulenta[249]

Sp. nov

Valid

Kopylov, Zhang & Zhang

Cretaceous

Burmese amber

Myanmar

A member of the family Ichneumonidae.

Holepyris rasnitsyni[235]

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

Russia

A member of the family Bethylidae.

Holepyris terayamai[235]

Sp. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Rovno amber

Ukraine

A member of the family Bethylidae.

Holopsenella antiqua[250]

Sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

Myanmar

A member of Aculeata of uncertain phylogenetic placement.

Holopsenella burmitica[250]

Sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

Myanmar

A member of Aculeata of uncertain phylogenetic placement.

Holopsenella gothica[250]

Sp. nov

In press

Lepeco & Melo

Cretaceous

Burmese amber

Myanmar

A member of Aculeata of uncertain phylogenetic placement.

Hybristodryinus castaneus[251]

Sp. nov

Valid

Olmi, Guglielmino & Chen in Olmi et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Dryinidae.

Hybristodryinus viriosus[252]

Sp. nov

Valid

Wang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Dryinidae.

Hybristodryinus zaifui[251]

Sp. nov

Valid

Chen, Olmi & Perkovsky in Olmi et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Dryinidae.

Janzenella theia[253]

Sp. nov

Valid

Bremer & Talamas in Bremer, van de Kamp & Talamas

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A species of Janzenella.

Karataus ryonsangensis[254]

Sp. nov

In press

Won & So

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

North Korea

A wasp belonging to the family Ephialtitidae.

Keratodellitha[255]

Gen. et 3 sp. nov

Valid

Jouault et al.

Cretaceous

Burmese amber

Myanmar

A member of Evanioidea belonging to the family Othniodellithidae. Genus includes new species K. anubis, K. basilisci and K. kirin.

Kleistochora[234]

Gen. et sp. nov

Valid

Chen, van Achterberg & Hong

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Braconidae belonging to the subfamily Protobraconinae. The type species is K. dolichura.

Lelejus[256]

Gen. et sp. nov

Valid

Radchenko & Proshchalykin

Late Eocene

Rovno amber

Ukraine

An ant belonging to the subfamily Myrmicinae. The type species is L. venustus.

Lonchodryinus groehni[257]

Sp. nov

Valid

Olmi et al.

Late Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Dryinidae.

Mael[235]

Gen. et comb. nov

Valid

Colombo & Azevedo in Colombo et al.

Eocene (Priabonian)

Baltic amber

Russia

A member of the family Bethylidae belonging to the subfamily Scleroderminae; a new genus for "Calyoza" longiceps Brues (1923).

Megalyrhyssalus[258]

Gen. et sp. nov

Valid

Belokobylskij & Jouault

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Braconidae. Genus includes new species M. clavicornis.

Megarolium[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Bembicinae. Genus includes new species M. paradoxum.

Merascylla[235]

Gen. et comb. nov

Valid

Colombo & Azevedo in Colombo et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Bethylidae belonging to the subfamily Pristocerinae; a new genus for "Epyris" atavellus Cockerell (1920).

Mesoserphites[233]

Gen. et 5 sp. nov

In press

Herbert & McKellar

Cretaceous

Burmese amber

Myanmar

A member of the family Serphitidae. Genus includes new species M. annulus, M. giganteus, M. engeli, M. scutatus and M. viraneacapitis.

Meteorus arasnitsyni[227]

Sp. nov

Valid

Belokobylskij et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A species of Meteorus.

Microtypus eocenus[227]

Sp. nov

Valid

Belokobylskij et al.

Eocene

Rovno amber

Ukraine

A member of the family Braconidae belonging to the subfamily Microtypinae.

Mintara[236]

Gen. et sp. nov

In press

Brazidec et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Diapriidae. Genus includes new species M. parva.

Odontosericus[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Psenini. Genus includes new species O. burmensis.

Orthosyntexis[259]

Gen. et 2 sp. nov

Valid

Gao, Engel, Shih, Gao in Gao et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Anaxyelidae. The type species is O. elegans; genus also includes O. thanti.

Palaeocolastes[227]

Gen. et sp. nov

Valid

Belokobylskij et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Braconidae belonging to the subfamily Exothecinae. The type species is P. bruesi.

Paleoaulacus[260]

Gen. et sp. nov

Valid

Jouault & Nel

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Aulacidae. Genus includes new species P. minutus.

Paleoxyela[261]

Gen. et sp. nov

Valid

Jouault, Aase & Nel

Eocene (Chadronian)

Florissant Formation

United States
( Colorado)

A member of the family Xyelidae. The type species is P. nearctica.

Peleserphus qufuensis[262]

Sp. nov

In press

Yu et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Peleserphidae.

Pheidole pauchil[263]

Sp. nov

Valid

Varela-Hernández & Riquelme

Miocene

Mexican amber

Mexico

An ant, a species of Pheidole.

Pinguixyela[231]

Gen. et comb. nov

Valid

Zheng et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A sawfly belonging to the family Syspastoxyelidae; a new genus for "Syspastoxyela" pinguis.

Platyxyela tenuis[264]

Sp. nov

Valid

Zheng et al.

Middle Jurassic (Callovian-Bathonian boundary)

Jiulongshan Formation

China

A member of the family Xyelidae.

Pompilopterus ryonsangensis[254]

Sp. nov

In press

Won & So

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

North Korea

A wasp belonging to the family Angarosphecidae.

Pristinopterus[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Bembicinae. Genus includes new species P. spectabilis.

Pristopsen[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Psenini. Genus includes new species P. obscurus.

Prosyntexis sennlaubi[265]

Sp. nov

Valid

Jouault & Nel

Early Cretaceous (Aptian)

Crato Formation

Brazil

A member of the family Sepulcidae.

Protopyris[230]

Gen. et sp. nov

Valid

Jouault & Nel in Jouault, Perrichot & Nel

Early Cretaceous (Albian)

Hkamti amber

Myanmar

A member of the family Bethylidae belonging to the subfamily Lancepyrinae. Genus includes new species P. myanmarensis.

Prototeleia[266]

Gen. et sp. nov

Valid

Talamas et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Platygastridae. The type species is P. kleio.

Proxyelia[261]

Gen. et sp. nov

Valid

Jouault, Aase & Nel

Eocene (Ypresian)

Green River Formation

United States
( Wyoming)

A member of the family Xyelidae. The type species is P. pankowskii.

Ptilocosmus[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Heterogynaidae. Genus includes new species P. corniculatus.

Ramageoptera[267]

Gen. et sp. nov

Valid

Jouault & Brazidec

Cretaceous

Burmese amber

Myanmar

A member of the family Bethylidae belonging to the subfamily Protopristocerinae. Genus includes new species R. platycephala.

Rasnichneumon[249]

Gen. et 2 sp. nov

Valid

Kopylov, Zhang & Zhang

Cretaceous

Burmese amber

Myanmar

A member of the family Ichneumonidae. The type species is R. alexandri; genus also includes R. gracilis.

Rasnitsynum[268]

Gen. et sp. nov

Valid

Olmi et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Dryinidae. Genus includes new species R. burmense.

Rogichneumon[249]

Gen. et sp. nov

Valid

Kopylov, Zhang & Zhang

Cretaceous

Burmese amber

Myanmar

A member of the family Ichneumonidae. The type species is R. braconidicus.

Rovenosa alexrasnitsyni[269]

Sp. nov

Valid

Manukyan in Manukyan & Zhindarev

Late Eocene

Prussian Formation
(Baltic amber)

Russia
( Kaliningrad Oblast)

A member of the family Ichneumonidae.

Rovenosa khalaimi[269]

Sp. nov

Valid

Manukyan in Manukyan & Zhindarev

Late Eocene

Prussian Formation
(Baltic amber)

Russia
( Kaliningrad Oblast)

A member of the family Ichneumonidae.

Sakhalinencyrtus[270]

Gen. et sp. nov

Valid

Simutnik in Simutnik, Perkovsky & Vasilenko

Middle Eocene

Sakhalinian amber

Russia

A member of the family Encyrtidae. The type species is S. leleji.

Sakhalinoctonus[271]

Gen. et sp. nov

Valid

Davidian in Davidian, Manukyan & Belokobylskij

Middle Eocene

Sakhalinian amber

Russia

A member of the family Braconidae belonging to the subfamily Aphidiinae. Genus includes new species S. alexrasnitsyni.

Scleroxyela[264]

Gen. et sp. nov

Valid

Zheng et al.

Middle Jurassic (Callovian-Bathonian boundary)

Jiulongshan Formation

China

A member of the family Xyelidae. The type species is S. daohugouensis.

Spalangiopelta darlingi[272]

Sp. nov

Valid

Moser in Moser et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Pteromalidae.

Spalangiopelta semialba[272]

Sp. nov

Valid

Moser in Moser et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Pteromalidae.

Spheciellus[223]

Fam. nov., Gen. et sp. nov.

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Apoidea in the new family Spheciellidae. Genus includes new species S. aenigmaticus.

Stephanorhyssalus[258]

Gen. et sp. nov

Valid

Belokobylskij & Jouault

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Braconidae. Genus includes new species S. longiscapus.

Tetragonula florilega[228]

Sp. nov

Valid

Engel in Engel et al.

Miocene (Langhian)

Zhangpu amber

China

A stingless bee, a species of Tetragonula.

Thigmocosmus[223]

Gen. et sp. nov

Valid

Rosa & Melo

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Heterogynaidae. Genus includes new species T. longicornis.

Tyrannomecia[273]

Gen. et sp. nov

In press

Jouault & Nel

Paleocene

Menat Formation

France

An ant belonging to the stem group of the subfamily Myrmeciinae. Genus includes new species T. inopinata.

Vespula? hassiaca[274]

Sp. nov

In press

Abels & Wedmann

Eocene

Messel pit

Germany

A member of the family Vespidae.

Xeris dorbnikensis[275]

Sp. nov

Valid

Manukyan in Manukyan & Smirnova

Eocene

Baltic amber

Europe (Baltic Sea region)

A horntail. Originally described as a species of Xeris; Archibald & Rasnitsyn (2022) transferred it to the genus Urocerus.[276]

Zigrasimecia goldingot[277]

Sp. nov

In press

Zhuang et al.

Cretaceous

Burmese amber

Myanmar

An ant in the tribe Sphecomyrmini.

### Hymenopteran Research[edit]

* Guo et al. (2021) report the discovery of two queen ant specimens from Burmese amber, belonging to the species Haidomyrmex cerberus and including the first known alate queen in Haidomyrmex.[278]
* Cooccurrence of ants belonging to extant Holarctic genus Lasius and a tropical weaver ant in the same piece of amber is reported from the Bitterfeld amber (Germany) by Radchenko & Perkovsky (2021).[279]
* Original claims that both Melittosphex burmensis and Discoscapa apicula were primitive bees have been rejected by Rosa & Melo, who instead consider the former to be an aculeate wasp of uncertain affinity, and the latter to belong within the crabronid wasp subfamily Crabroninae, a lineage not directly related to bees.[223]

## Clade Neuropterida[edit]

### Neuropterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Ankyloleon[280]

Gen. et sp. nov

Valid

Badano, Haug & Cerretti in Badano et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

An early representative of the clade Myrmeleontiformia. The type species is A. caudatus.

Astioberotha coutreti[281]

Sp. nov

In press

Jouault

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Rhachiberothidae belonging to the subfamily Paraberothinae.

Burmotachinymphes[282]

Gen. et sp. nov

Valid

Cao, Wang & Liu

Cretaceous

Burmese amber

Myanmar

A member of the family Mesochrysopidae. Genus includes new species M. bilobata.

Calobabinskaia[283]

Gen. et sp. nov

Valid

Lu, Wang & Liu

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Babinskaiidae. The type species is C. xiai.

Fuscopolystoechotes[284]

Gen. et sp. nov

Valid

Xu, Zheng, Shih & Wang in Xu et al.

Middle Jurassic

Jiulongshan Formation

China

A member of the family Ithonidae. Genus includes new species F. reticulatus.

Karenina cuneiformis[285]

Sp. nov

Valid

Nakamine

Early Cretaceous (Aptian)

Crato Formation

Brazil

A member of the family Mesochrysopidae.

Lichenipolystoechotes fenestratus[284]

Sp. nov

Valid

Xu, Fang, Shih & Wang in Xu et al.

Middle Jurassic

Jiulongshan Formation

China

A member of the family Ithonidae.

Osmylidia donnae[286]

Sp. nov

Valid

Makarkin, Archibald, & Mathewes

Ypresian

Quilchena site

Canada
British Columbia

A protosmyline Osmylid lance lacewing

Osmylidia glastrai[286]

Sp. nov

Valid

Makarkin, Archibald, & Mathewes

Ypresian

Klondike Mountain Formation

United States
Washington

A protosmyline Osmylid lance lacewing

Osmylidia picta[286]

Comb. nov

Valid

(Hagen)

Priabonian

Baltic amber

Europe

A protosmyline Osmylid lance lacewing
Moved from "Protosmylus" picta (1913)

Osmylidia taliae[286]

Sp. nov

Valid

Makarkin, Archibald, & Mathewes

Ypresian

Green River Formation
Parachute Creek Member

United States
Colorado

A protosmyline Osmylid lance lacewing

Osmyloberotha[287]

Gen. et sp. nov

Valid

Khramov

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A beaded lacewing. Genus includes new species O. simpla.

Paradoxoberotha[288]

Gen. et sp. nov

In press

Nakamine et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Rhachiberothidae. Genus includes new species P. chimaera.

Paraneliana[289]

Gen. et sp. nov

Valid

Jouault & Nel

Early Cretaceous

Crato Formation

Brazil

A member of the family Babinskaiidae. Genus includes new species P. sennlaubi.

Rasnitsyneura[290]

Gen. et comb. nov

Valid

Makarkin, Ansorge & Khramov

Early Jurassic (Toarcian)

|  |

Germany

A member of the family Ithonidae sensu lato belonging to the subfamily Epigambriinae. The type species is "Prohemerobius" aequabilis Bode (1953).

Sinonothochrysa[291]

Gen. et sp. nov

In press

Huang & Nel in Huang et al.

Paleocene

Buxin Formation

China

A member of the family Chrysopidae. Genus includes new species S. zhangi

Stenobabinskaia[283]

Gen. et sp. nov

Valid

Lu, Wang & Liu

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Babinskaiidae. The type species is S. punctata.

Xiaobabinskaia[283]

Gen. et sp. nov

Valid

Lu, Wang & Liu

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Babinskaiidae. The type species is X. lepidotricha.

#### Neuropteran research[edit]

* Review of the known record of spoon-winged lacewing larvae is published Haug, Haug & Haug (2021), who report two specimens from the Cretaceous amber from Myanmar which might represent the first known larvae of members of the subfamily Nemopterinae in the fossil record.[292]
* New lacewing larvae with elongated mouthparts are described from the Cretaceous amber from Myanmar by Haug et al. (2021), who also review the morphological diversity of fossil lacewing larvae.[293]

### Raphidiopterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Archiinocellia protomaculata[294]

comb nov

valid

(Engel)

Lutetian

Green River Formation

USA
Colorado

A raphidiid snakefly.
Moved from Archiinocellia protomaculata Engel (2011)[295]

|  Archiinocellia protomaculata

Electrobaissoptera liui[296]

Sp. nov

In press

Jouault, Engel & Nel

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A baissopterid snakefly.

Megaraphidia antiquissima[294]

Sp. nov

Valid

Archibald & Makarkin

Ypresian

Tranquille Formation

Canada
British Columbia

A raphidiid snakefly.

Megaraphidia hopkinsi[294]

Sp. nov

Valid

Archibald & Makarkin

Ypresian

Allenby Formation

Canada
British Columbia

A raphidiid snakefly.

Megaraphidia klondika[294]

Sp. nov

Valid

Archibald & Makarkin

Ypresian

Klondike Mountain Formation

United States
Washington

A raphidiid snakefly.

Megaraphidia ootsa[294]

Sp. nov

Valid

Archibald & Makarkin

Ypresian

"Unnamed unit" Ootsa Group

Canada
British Columbia

A raphidiid snakefly.

Mesoraphidia baekthoensis[297]

Sp. nov

In press

Ri et al.

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

North Korea

A member of the family Mesoraphidiidae.

Mesoraphidia ryonsangensis[297]

Sp. nov

In press

Ri et al.

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

North Korea

A member of the family Mesoraphidiidae.

## Clade †Palaeodictyopteroidea[edit]

### †Megasecoptera[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Anhuihymen[298]

Gen. et sp. nov

In press

Huang & Nel in Huang et al.

Middle Permian

Yinping Formation

China

A protohymenid Megasecoptera.
The type species is A. medianelongata.

### †Palaeodictyoptera[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Dictyoneura goujonorum[299]

Sp. nov

In press

Logghe et al.

Carboniferous-Permian (Gzhelian-Asselian)

|  |

France

A member of Palaeodictyoptera.

Sandiella herbigi[300]

Sp. nov

In press

Brauckmann, Herd & Leipner

Carboniferous (Moscovian)

|  |

Germany

A member of Palaeodictyoptera belonging to the family Eugereonidae.

#### Palaeodictyopteran research[edit]

* A study on the anatomy of fossil specimens described as consecutive series of different larval stages of palaeodictyopteran species Tchirkovaea guttata and Paimbia fenestrata, evaluating their implications for the knowledge of wing development in palaeodictyopterans, is published by Rosová, Sinitshenkova & Prokop (2021).[301]

## Clade Palaeoptera[edit]

### Ephemeroptera[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Astraeoptera[302]

Gen. et sp. nov

Valid

Brandão et al.

Early Cretaceous

Crato Formation

Brazil

A mayfly of uncertain phylogenetic placement.
Genus includes new species A. cretacica.

Burmella[303]

Gen. et 2 sp. nov

Valid

Godunko, Martynov & Staniczek

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A vietnamellid mayfly.
The type species is B. paucivenosa; genus also includes B. clypeata.

Protoligoneuria heloisae[304]

Sp. nov

In press

Storari et al.

Early Cretaceous (Aptian)

Crato Formation

Brazil

A hexagenitid mayfly.

Triassodotes rasnitsyni[305]

Sp. nov

Valid

Sinitshenkova

Triassic

|  |

Russia

A misthodotid mayfly.

#### Ephemeropteran research[edit]

* Sroka et al. (2021) provide new information on the morphology of Permian stem-mayflies Misthodotes sharovi and M. zalesskyi from the Tshekarda locality (Kungurian Koshelevka Formation; Perm Krai, Russia), and infer the life history traits of both the adult and larval stages of these insects.[306]
* Possible underwater mayfly burrows are described from the Turonian Ferron Sandstone (Utah, United States) by King et al. (2021), who interpret this finding as evidence that burrowing mayflies adapted to domicile filter-feeding during or prior to the Turonian, and name a new ichnospecies Glossifungites gingrasi.[307]

### Odonatoptera[edit]

#### Odonata[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Chalcolestes tibetensis[308]

Sp. nov

In press

Xia et al.

Late Eocene

Nima Basin

China

A species of Chalcolestes.

Cretaphylolestes[309]

Gen. et sp. nov

In press

Huang et al.

Early Cretaceous (Aptian)

Shouchang Formation

China

A damselfly belonging to the family Synlestidae. Genus includes new species C. cretacicus.

Dakotaeschnidium[310]

Gen. et sp. nov

Valid

Nel

Late Cretaceous (Maastrichtian)

Fox Hills Formation

United States
( South Dakota)

A dragonfly belonging to the family Aeschnidiidae. Genus includes new species D. spedeni.

Dysagrion pruettae[311]

Sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Klondike Mountain Formation

United States
( Washington)

A member of the family Dysagrionidae.

|  Dysagrion pruettae

Dysagrionites[311]

Gen. et sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Klondike Mountain Formation

United States
( Washington)

A collective group name for any fossil species referable to the Dysagrioninae whose orthotaxon generic affinity is unclear. Includes new species D. delinei.

Epilestes rasnitsyni[312]

Sp. nov

Valid

Felker & Vasilenko

Permian

|  |

Russia

A damselfly belonging to the family Permagrionidae.

Jingguaeshna[313]

Gen. et sp. nov

Valid

Zheng et al.

Miocene

|  |

China

A dragonfly belonging to the group Aeshnoidea, possibly a member of the family Telephlebiidae. Genus includes new species J. taoae.

Marado[314][315]

Gen. et sp. nov

Valid

Petrulevičius

Paleocene

Maíz Gordo Formation

Argentina

A damselfly belonging to the family Coenagrionidae. The type species is M. marado.

Moltenophlebia[316]

Gen. et sp. nov

Valid

Deregnaucourt et al.

Triassic

Molteno Formation

South Africa

A member of the stem group of Odonata, belonging to the group Zygophlebiida within the Triadophlebiomorpha. Genus includes new species M. lindae.

Okanagrion[311]

Gen. et 8 sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Klondike Mountain Formation

United States
( Washington)

A member of the family Dysagrionidae. The type species is O. threadgillae; genus also includes new species O. hobani, O. beardi, O. lochmum, O. angustum, O. dorrellae, O. liquetoalatum and O. worleyae.

|  Okanagrion threadgillae

Okanopteryx[311]

Gen. et 3 sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Tranquille Formation

Canada
( British Columbia)

A member of the family Dysagrionidae. The type species is O. macabeensis; genus also includes new species O. fraseri and O. jeppesenorum.

Phyonganpodagrion[317]

Gen. et sp. nov

In press

So & Won

Early Cretaceous (Barremian–Aptian)

Sinuiju Formation

North Korea

A damselfly in the family Megapodagrionidae. Genus includes new species P. ryonsangae.

Protohemiphlebia[318]

Fam. et gen. et 2 sp. nov

In press

Zheng, Jarzembowski & Nel in Zheng et al.

Cretaceous

Burmese amber

Myanmar

A stem hemiphlebioid damselfly belonging to the new family Protohemiphlebiidae. Genus includes new species P. zhangi and P. meiyingae.

Pulchrairina[319]

Gen. et sp. nov

In press

Martynov, Vasilenko & Perkovsky

Late Eocene

Rovno amber

Ukraine

A damselfly belonging to the group Coenagrionoidea.
Genus includes new species P. electra.

Republica[320]

Gen. et sp. nov

Valid

Archibald & Cannings

Eocene (Ypresian)

Klondike Mountain Formation

United States
( Washington)

A euphaeid damselfly.
Type species is R. weatbrooki.
Not to be confused with the plant genus Republica also from the formation.

|  Republica weatbrooki

Stenodiafanus[311]

Gen. et sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Klondike Mountain Formation

United States
( Washington)

A dysagrionid odonate.
Type species is S. westersidei.

|  Stenodiafanus westersidei

Stenophlebia ryonsangensis[321]

Sp. nov

In press

Won, So & Kim

Early Cretaceous

Sinuiju Formation

North Korea

A stenophlebiid odonate.

Whetwhetaksa[311]

Fam. Nov., Gen. et sp. nov

Valid

Archibald & Cannings in Archibald et al.

Eocene (Ypresian)

Klondike Mountain Formation

United States

The type genus of Whetwhetaksidae, a new family of uncertain placement.
The type species is W. millerae.

|  Whetwhetaksa millerae

##### Odonate research[edit]

* Archibald et al. (2021) establish a new, extinct suborder of Odonata named Cephalozygoptera, close to but distinct from damselflies, composed of the families Dysagrionidae, Sieblosiidae and possibly Whetwhetaksidae;[311] their study is subsequently criticized by Nel & Zheng (2021), who consider evidence presented by Archibald et al. to be insufficient to support a new clade, and reject the suborder Cephalozygoptera as unfounded.[322][323]

#### †Protozygoptera[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Kennedya kedrovkensis[324]

Sp. nov

Valid

Felker

Early Triassic

|  |

Russia

A member of Protozygoptera belonging to the family Kennedyidae.

Kennedya suchonensis[325]

Sp. nov

Valid

Felker

Late Permian

|  |

Russia

|  |

Progoneura kemerovensis[324]

Sp. nov

Valid

Felker

Early Triassic

|  |

Russia

A member of Protozygoptera belonging to the family Kennedyidae.

#### Other odonatopterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Brunellopteron[326]

Gen. et sp. nov

Valid

Béthoux, Deregnaucourt & Norrad in Béthoux et al.

Carboniferous (Moscovian)

Sunbury Creek Formation

Canada
( New Brunswick)

A member of Neodonatoptera of uncertain phylogenetic placement. The type species is B. norradi.

Osnabruggiala[327]

Gen. et sp. nov

Valid

Zessin, Brauckmann & Gröning

Carboniferous (Pennsylvanian)

Osnabrück Formation

Germany

Probably a basal member of Odonatoptera. The type species is O. seppelti.

## Clade †Paoliidea[edit]

### †Paoliida[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Avionblattinopsis[328]

Gen. et sp. nov

In press

Quispe et al.

Carboniferous (Moscovian)

|  |

France

A blattinopsid Paoliidan.
The type species is A. oudardi.

Blattinopsis indefinitus[329]

Sp. nov

Valid

Rasnitsyn & Aristov

Permian (Kungurian)

|  |

Russia
( Krasnoyarsk Krai)

A Blattinopsidae Paoliidan

Glaphyrophlebia borea[330]

Sp. nov

Valid

Aristov, Rasnitsyn & Naugolnykh

Permian

Inta Formation

Russia
( Komi Republic)

A Blattinopsid.

Glaphyrophlebia iva[331]

Sp. nov

Valid

Aristov & Rasnitsyn

Middle Permian

|  |

Russia

A member of the family Blattinopsidae.

Glaphyrophlebia komia[330]

Sp. nov

Valid

Aristov, Rasnitsyn & Naugolnykh

Permian

Inta Formation

Russia
( Komi Republic)

A member of the family Blattinopsidae.

Glaphyrophlebia vorkutensis[330]

Sp. nov

Valid

Aristov, Rasnitsyn & Naugolnykh

Permian

Inta Formation

Russia
( Komi Republic)

A member of the family Blattinopsidae.

## Clade Paraneoptera[edit]

### Hemipterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Acrotiara[332]

Trib. et gen. et sp. nov

In press

Bourgoin & Luo in Luo et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Cixiidae; the type genus of the new tribe Acrotiarini. The type species is A. multigranulata Luo & Bourgoin.

Aneurus damzeni[333]

Sp. nov

Valid

Heiss in Heiss, Kairišs & Bukejs

Eocene

Baltic amber

Russia
( Kaliningrad Oblast)

A species of Aneurus.

Aphelicophontes[334]

Gen. et sp. nov

Valid

Swanson et al.

Eocene

Green River Formation

United States

A member of the family Reduviidae belonging to the subfamily Harpactorinae. Genus includes new species A. danjuddi.

Baltiomiris[335]

Gen. et sp. nov

Valid

Kim, Chérot & Jung

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Miridae. The type species is B. herczeki.

Barremixius[332]

Gen. et comb. nov

In press

Bourgoin & Szwedo in Luo et al.

Early Cretaceous

Weald Clay

United Kingdom

A member of the family Cixiidae; a new genus for "Cixius" petrinus Fennah.

Beaconiella cincta[336]

Sp. nov

Valid

Shcherbakov

Triassic (Ladinian–Carnian)

Madygen Formation

Kyrgyzstan

A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Curvicubitidae.

Beaconiella pulchra[336]

Sp. nov

Valid

Shcherbakov

Triassic (Ladinian–Carnian)

Madygen Formation

Kyrgyzstan

A member of Cicadomorpha belonging to the superfamily Pereborioidea and the family Curvicubitidae.

Burmacader bicoloripennis[337]

Sp. nov

Valid

Souma, Yamamoto & Takahashi

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Tingidae.

Burmacader grandis[338]

Sp. nov

Valid

Heiss & Guilbert

Cretaceous

Burmese amber

Myanmar

A member of the family Tingidae.

Burmala[339]

Gen. et sp. nov

Valid

Liu et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Malmopsyllidae. Genus includes new species B. liaoyaoi.

Burmaprosbole[340]

Gen. et sp. nov

Valid

Qiao et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Tettigarctidae. Genus includes new species B. nervosa.

Burmavianaida[341]

Gen. et sp. nov

Valid

Souma, Yamamoto & Takahashi

Cretaceous (Albian or Cenomanian)

Burmese amber

Myanmar

A member of the family Tingidae. The type species is B. anomalocapitata.

Cretamystilus[342]

Gen. et sp. nov

Valid

Kim, Lim & Jung

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Miridae belonging to the tribe Mecistoscelini. Genus includes new species C. herczeki.

Cretodorus rostellatus[343]

Sp. nov

In press

Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Mimarachnidae.

Cretomultinervis[344]

Gen. et sp. nov

Valid

Fu & Huang

Cretaceous

Burmese amber

Myanmar

A member of the family Sinoalidae. Genus includes new species C. burmensis.

Dachibangus hui[345]

Sp. nov

Valid

Zhang, Yao & Pang in Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Mimarachnidae. Originally described as a species of Dachibangus, but subsequently transferred to the genus Xiaochibangus.[346]

Delphitiara[332]

Gen. et sp. nov

In press

Bourgoin & Luo in Luo et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is D. tibiocoronata Luo & Bourgoin.

Dorytocus jiaxiaoae[347]

Sp. nov

Valid

Song, Szwedo & Bourgoin in Song et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A planthopper.

Duyana[348]

Gen. et sp. nov

Valid

Chen et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A leafhopper belonging to the subfamily Ledrinae. Genus includes new species D. electronica.

Electromyiomma herczeki[349]

Sp. nov

Valid

Kim & Jung

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Miridae.

Eospinosus[350]

Gen. et 2 sp. nov

Valid

Wedmann et al.

Eocene

Messel pit

Germany
United States
( Colorado)

A member of the family Pentatomidae. The type species is E. peterkulkai; genus also includes E. greenriverensis.

Ferriantenna[351]

Gen. et sp. nov

Valid

Cumming & Le Tirant

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Coreidae belonging to the subfamily Coreinae. The type species is F. excalibur.

Hallex martinsnetoi[352]

Sp. nov

In press

Monferran et al.

Early Cretaceous

Crato Formation

Brazil

A leafhopper.

Hypsotsijilia[353]

Gen. et sp. nov

Valid

Cifuentes-Ruiz & Brailovsky in Cifuentes-Ruiz et al.

Early Miocene

La Quinta Formation
(Mexican amber)

Mexico

A member of the family Schizopteridae. The type species is H. bretoni.

Inoderbe[354]

Fam. et gen. et sp. nov

Valid

Shcherbakov & Emeljanov

Cretaceous (Albian–Cenomanian)

Burmese amber

Myanmar

A planthopper belonging to the new family Inoderbidae. The type species is I. rapunzel.

Juroala pulchra[355]

Sp. nov

Valid

Fu & Huang

Late Jurassic

Daohugou Beds

China

A member of the family Sinoalidae.

Karatauella[356]

Nom. nov

Valid

Doweld

Jurassic (Bathonian-Oxfordian)

|  |

Kazakhstan

A member of the family Protopsyllidiidae; a replacement name for Carpenterella Becker-Migdisova (1968).

Latidorsum[357]

Gen. et sp. nov

|  |

Wang, Tang & Yao in Wang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Tingidae. The type species is L. carinbifarium.

Microtingis[358]

Gen. et sp. nov

Junior synonym

Heiss & Golub

Cretaceous

Burmese amber

Myanmar

A member of the family Tingidae. Genus includes new species M. leptosoma. Subsequently Heiss & Golub (2021) considered this species to be a junior synonym of Burmavianaida anomalocapitata.[359]

Multistria[345]

Gen. et sp. nov

Valid

Zhang, Yao & Pang in Zhang et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Mimarachnidae. The type species is M. orthotropa.

Paleocader balticus[360]

Sp. nov

Valid

Heiss & Golub

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Tingidae.

Paleocader pulchellus[360]

Sp. nov

Valid

Heiss & Golub

Eocene

Baltic amber

Europe (Baltic Sea region)

A member of the family Tingidae.

Paleocader rovnensis[360]

Sp. nov

Valid

Heiss & Golub

Eocene

Rovno amber

Ukraine

A member of the family Tingidae.

Parasinalda sukachevae[361]

Sp. nov

Valid

Golub, Perkovsky & Vasilenko

Late Eocene

Rovno amber

Ukraine

A member of the family Tingidae.

Pentacarinus[332]

Gen. et sp. nov

In press

Bourgoin & Luo in Luo et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of the family Cixiidae belonging to the tribe Acrotiarini. The type species is P. kachinensis Luo & Bourgoin.

Pseudocaulisoculus[362]

Gen. et sp. nov

Valid

Kóbor & Roca-Cusachs

Cretaceous

Burmese amber

Myanmar

A member of the family Yuripopovinidae. Genus includes new species P. longicornis.

Pubivetanthocoris[363]

Gen. et sp. nov

In press

Tang, Wang & Yao in Tang et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Vetanthocoridae. Genus includes new species P. carinalis.

"Pyrrhocoris" rottensis[364]

Nom. nov

Valid

Kment

Late Oligocene

|  |

Germany

A member of Lygaeoidea of uncertain phylogenetic placement; a replacement name for Pyrrhocoris tibialis Statz & Wagner (1950).

Rovnoxestus[365]

Gen. et sp. nov

Valid

Dietrich, Dmitriev & Perkovsky

Late Eocene

Rovno amber

Ukraine

A leafhopper. Genus includes new species R. rasnitsyni.

Saltissus fennahi[366]

Sp. nov

In press

Luo, Liu & Jarzembowski in Luo et al.

Early Cretaceous

Weald Clay

United Kingdom

A member of the family Mimarachnidae.

Scytinoptera tongchuanensis[367]

Sp. nov

In press

Zhang, Zheng & Zhang in Zhang et al.

Middle Triassic

Tongchuan Formation

China

A member of the family Scytinopteridae.

Sinogranulus[368]

Gen. et sp. nov

In press

Zhang, Chen & Zhang

Middle Triassic

Tongchuan Formation

China

A member of Scytinopteroidea belonging to the family Granulidae. Genus includes new species S. qishuiheensis.

Spinonympha[369]

Gen. et sp. nov

Valid

Luo, Wang & Jarzembowski

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A planthopper of uncertain phylogenetic placement. The type species is S. shcherbakovi.

Szeiinia[370]

Fam. nov., Gen. et sp. nov

Valid

Zhang et al.

Late Triassic

Yanchang Formation

China

A planthopper in the new family Szeiiniidae. Genus includes new species S. huanglongensis

Tennentsia orientalis[371]

Sp. nov

In press

Fu, Azar & Huang

Middle Triassic

Yanchang Formation

China

A member of Cicadomorpha belonging to the family Dysmorphoptilidae.

Triknightia[336]

Gen. et sp. nov

Valid

Shcherbakov

Triassic (Ladinian–Carnian)

Madygen Formation

Kyrgyzstan

A member of Cicadomorpha belonging to the superfamily Scytinopteroidea and the family Paraknightiidae. The type species is T. mira.

Tuthillia maculosa[372]

Sp. nov

Valid

Poinar & Brown

|  |

Dominican amber

Dominican Republic

A member of Psylloidea.

Valdicopis[373]

Gen. et sp. nov

Valid

Li, Chen & Jarzembowski

Early Cretaceous

|  |

United Kingdom

A member of the family Procercopidae. Genus includes new species V. tonyi.

Youngeewa simillima[374]

Sp. nov

Valid

Gębicki & Walczak in Gębicki, Walczak & Zmarzły

Eocene

Baltic amber

Europe (Baltic Sea region)

A leafhopper belonging to the subfamily Mileewinae.

#### Hemipteran research[edit]

* Grimaldi & Vea (2021) reinterpret Mesophthirus engeli as a scale insect, and reject the interpretation of this insect as an ectoparasite.[375]
* Redescription of the holotype specimen of Paranoika placida is published by Moura-Júnior et al. (2021), who transfer this species to the genus Lethocerus, and consider this species to be the senior synonym of Lethocerus vetus.[376]
* A fulgorid specimen, representing the oldest occurrence of this family in the fossil record in Asia, is described from the Eocene Lunpola Basin (Qinghai-Tibetan Plateau, China) by Xu et al. (2021).[377]
* Revision of the fossil record of Fulgoromorpha and Cicadomorpha from the Jurassic and Cretaceous of eastern Asia is published by Chen (2021).[378]

### Psocodeans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Burmesopsocus[379]

Gen. et sp. nov

Valid

Yoshizawa in Yoshizawa & Yamamoto

Cretaceous (Albian–Cenomanian)

Burmese amber

Myanmar

A member of Homilopsocidea of uncertain phylogenetic placement. The type species is B. lienhardi.

Cormopsocus baleoi[380]

Sp. nov

Valid

Hakim, Azar & Huang

Cretaceous

Burmese amber

Myanmar

A member of Trogiomorpha belonging to the family Cormopsocidae.

Cormopsocus neli[381]

Sp. nov

Valid

Hakim, Azar & Huang in Hakim et al.

Cretaceous

Burmese amber

Myanmar

A member of Trogiomorpha belonging to the family Cormopsocidae.

Empherium[382]

Gen. et sp. nov

Valid

Hakim, Huang & Azar

Late Cretaceous

|  |

Russia

A member of Trogiomorpha belonging to the family Empheriidae. Genus includes new species E. rasnitsyni.

Eolepinotus zherikhini[382]

Sp. nov

Valid

Hakim, Huang & Azar

Late Cretaceous

|  |

Russia

A member of Trogiomorpha belonging to the family Trogiidae.

Heliadesdakruon[383]

Gen. et sp. nov

Valid

Cumming & Le Tirant

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Trogiomorpha belonging to the family Archaeatropidae. The type species is H. morganae.

Longiglabellus[384]

Gen. et 2 sp. nov

In press

Wang, Li & Yao in Wang et al.

Cretaceous

Burmese amber

Myanmar

A member of the family Cormopsocidae. Genus includes new species L. edentatus and L. pedhyalinus.

Palaeosiamoglaris hammanaensis[385]

Sp. nov

In press

Hakim, Huang & Azar

Early Cretaceous (Barremian)

Lebanese amber

Lebanon

A member of Psocodea belonging to the family Prionoglarididae.

Palaeosiamoglaris hkamtiensis[386]

Sp. nov

In press

Jouault et al.

Early Cretaceous

Hkamti amber

Myanmar

A member of Psocodea belonging to the family Prionoglarididae.

Psyllipsocus myanmarensis[386]

Sp. nov

In press

Jouault et al.

Late Cretaceous (Cenomanian)

Burmese amber

Myanmar

A member of Psocodea belonging to the family Psyllipsocidae, a species of Psyllipsocus.

Psyllipsocus yangi[387]

Sp. nov

Valid

Liang & Liu

Cretaceous

Burmese amber

Myanmar

A member of Psocodea belonging to the family Psyllipsocidae, a species of Psyllipsocus.

Stimulopsocus[388]

Gen. et sp. nov

In press

Liang & Liu

Cretaceous

Burmese amber

Myanmar

A member of the family Cormopsocidae. Genus includes new species S. jiewenae.

#### Psocodean research[edit]

* Four bark lice carrying sand granules and organic material atop their back are described from the Cretaceous amber from Myanmar by Kiesmüller et al. (2021), who interpret this finding as the first fossil evidence of masking behaviour in Cretaceous representatives of Psocodea.[389]

## Clade Perlidea[edit]

### Dermapterans[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Aglyptoderma[390]

Gen. et 2 sp. nov

Valid

Xiong et al.

Middle Jurassic

Jiulongshan Formation

China

A semenoviolid eodermapteran earwig.
Genus includes the new species A. elongatum and A. cylindratum.

Aneuroderma[391]

Gen. et sp. nov

Valid

Xiong, Engel & Ren in Xiong et al.

Middle Jurassic

Jiulongshan Formation

China

A member of the family Protodiplatyidae. The type species is A. oiodes.

Dacryoderma[392]

Gen. et comb. nov

Valid

Engel

Early Jurassic (Sinemurian)

Charmouth Mudstone Formation

United Kingdom

A dermapterid earwig.
The type species is "Brevicula" teres Tihelka (2019).

Eminepygia[393]

Gen. et sp. nov

Valid

Chen & Zhang

Cenomanian

Burmese amber

Myanmar

A pygidicranid earwig.
Genus includes new species E. myanmarensis.

Sinopalaeodermata concavum[391]

Sp. nov

Valid

Xiong, Engel & Ren in Xiong et al.

Middle Jurassic

Jiulongshan Formation

China

A member of the family Dermapteridae.

### Plecoptera[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Balticonemoura[394]

Gen. et sp. nov

In press

Chen

Eocene

Baltic amber

Europe (Baltic Sea region)

A nemourid stonefly.
Genus includes new species B. bulbosus.

Billoperla[395]

Gen. et sp. nov

In press

Chen & Xu

Cretaceous

Burmese amber

Myanmar

A perlid stonefly belonging to the family.
Genus includes new species B. starki.

Isoperla baltica[396]

Sp. nov

Valid

Jouault et al.

Eocene

Baltic amber

Europe (Baltic Sea regon)

A perlodid stonefly.

## Other insects[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Country  | Notes  | Images

|
|
|
|
|
|
|
|

Anakitala[397]

Gen. et sp. nov

Valid

Aristov

Late Permian

|  |

Russia

A mesorthopterid Eoblattidan Polyneopteran.
The type species is A. intermittendia.

Baharellinus elegans[397]

Sp. nov

Valid

Aristov

Late Permian

|  |

Russia

A member of Polyneoptera belonging to the group Eoblattida and the family Blattogryllidae.

Belmophenopterum rasnitsyni[398]

Sp. nov

Valid

Storozhenko & Aristov

Triassic (Ladinian-Carnian)

Madygen Formation

Kyrgyzstan

A member of the family Mesorthopteridae.

Cabarzopterum[399]

Gen. et sp. nov

Valid

Cui et al.

Early Permian

Goldlauter Formation

Germany

A member of Grylloblattida of uncertain phylogenetic placement. Genus includes new species C. magnificus.

Delopterinus alsenzensis[400]

Sp. nov

Valid

Nel & Poschmann

Early Permian

Saar–Nahe Basin

Germany

A miomopteran belonging to the family Palaeomanteidae.

Delopterum novokshonovi[400]

Sp. nov

Valid

Nel & Poschmann

Early Permian

Saar–Nahe Basin

Germany

A miomopteran belonging to the family Palaeomanteidae.

Elmocladus[401]

Gen. et comb. nov

Valid

Rasnitsyn & Aristov

Early Permian

|  |

United States

A member of Polyneoptera belonging to the group Eoblattida and the family Tococladidae. The type species is "Opisthocladus" strictus Carpenter (1976).

Galloembia[402]

Gen. et sp. nov

Valid

Falières, Engel & Nel

Earliest Eocene

Oise amber

France

A webspinner belonging to the family Embiidae. The type species is G. raholai.

Gnethoda odontophora[403]

Sp. nov

In press

Lai et al.

Cretaceous

Burmese amber

Myanmar

A webspinner belonging to the family Clothodidae.

Kamopanorpa rasnitsyni[404]

Sp. nov

Valid

Bashkuev & Sukatsheva

Permian

Minusinsk Coal Basin

Russia

A member of the (stem-trichopteran or stem-amphiesmenopteran) family Microptysmatidae.

Kamopanorpa rotunda[404]

Sp. nov

Valid

Bashkuev & Sukatsheva

Permian

Minusinsk Coal Basin

Russia

A member of the family Microptysmatidae.

Kamopanorpa sivchikovi[404]

Sp. nov

Valid

Bashkuev & Sukatsheva

Permian

Minusinsk Coal Basin

Russia

A member of the family Microptysmatidae.

Liomopterites sperbersbachensis[399]

Sp. nov

Valid

Cui et al.

Early Permian

Goldlauter Formation

Germany

A member of Grylloblattida belonging to the family Liomopteridae.

Liomopterum fuscatum[399]

Sp. nov

Valid

Cui et al.

Early Permian

Goldlauter Formation

Germany

A member of Grylloblattida belonging to the family Liomopteridae.

Madygembia[405]

Gen. et sp. nov

Valid

Aristov & Storozhenko

Triassic (Ladinian or Carnian)

Madygen Formation

Kyrgyzstan

A webspinner belonging to the family Alexarasniidae. The type species is M. rasnitsyni.

Magicivena[406]

Fam. et gen. et 3 sp. nov

Valid

Yang et al.

Early Cretaceous

Yixian Formation

China

A member of Polyneoptera of uncertain phylogenetic placement, belonging to the new family Magicivenidae. The type species is M. sticta; genus also includes M. elegans and M. antennalis.

Multivena[403]

Gen. et sp. nov

In press

Lai et al.

Cretaceous

Burmese amber

Myanmar

A webspinner belonging to the family Sorellembiidae. Genus includes new species M. curvivena.

Pictoborella[399]

Gen. et sp. et comb. nov

Valid

Cui et al.

Early Permian

|  |

Germany

A member of Grylloblattida. Genus includes new species P. clara, as well as "Oborella" germanica Prokop et al. (2021).

Rasnalexia[407]

Fam. et gen. et sp. nov

Valid

Gorochov

Triassic

|  |

Kyrgyzstan

A member of Embioptera belonging to the new family Rasnalexiidae. Genus includes new species R. rasnitsyni.

Uralioma thuringiensis[399]

Sp. nov

Valid

Cui et al.

Early Permian

Goldlauter Formation

Germany

A member of Grylloblattida belonging to the family Liomopteridae.

### Research[edit]

* A well-preserved full-body impression of an elongate winged insect strongly resembling stick insects is described from the Permian (Wordian) Pelitic Formation (France) by Logghe et al. (2021), who name a new ichnotaxon Phasmichnus radagasti.[408]
* A dragonfly-type odonatopteran larva, preserving ancestral morphological characters maintained in mayflies but lost in modern odonatan larvae, is described from the Cretaceous Burmese amber by Haug, Müller & Haug (2021).[409]

## General research[edit]

* Revision of the fossil record of insects from the Triassic of China is published by Zhang et al. (2021).[410]
* A study on the diversity of insect damage types in fossil plants from the Cretaceous (Albian to Cenomanian) Dakota Formation (United States), evaluating their implications for the knowledge of the early evolution of angiosperm florivory and associated pollination, is published by Xiao et al. (2021).[411]

## References[edit]

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