2022 in paleobotany

Overview of the events of 2022 in paleobotany

List of years in paleobotany

* …
* 2019
* 2020
* 2021
* 2022
* 2023
* 2024
* 2025
* …

In paleontology
2019
2020
2021
2022
2023
2024
2025

In arthropod paleontology
2019
2020
2021
2022
2023
2024
2025

In paleoentomology
2019
2020
2021
2022
2023
2024
2025

In paleomalacology
2019
2020
2021
2022
2023
2024
2025

In reptile paleontology
2019
2020
2021
2022
2023
2024
2025

In archosaur paleontology
2019
2020
2021
2022
2023
2024
2025

In mammal paleontology
2019
2020
2021
2022
2023
2024
2025

In paleoichthyology
2019
2020
2021
2022
2023
2024
2025

* Art
* Archaeology
* Architecture
* Literature
* Music
* Philosophy
* Science +...

This paleobotany list records new fossil plant taxa that were described during 2022, as well as notes other significant paleobotany discoveries and events which occurred during 2022.

## Algae

[edit]

### Charophytes

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Unit  | Location  | Synonymized taxa  | Notes  | Images

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Lamprothamnium elongatum[1]

Sp. nov

In press

Feist & Floquet

Late Cretaceous

|  |

Spain

|  |

A charophyte.

Lamprothamnium ovoideum[1]

Sp. nov

In press

Feist & Floquet

Late Cretaceous

|  |

Spain

|  |

A charophyte.

Pseudoharrisichara sedanoensis[1]

Sp. nov

In press

Feist & Floquet

Late Cretaceous

|  |

Spain

|  |

A charophyte.

#### Charophyte research

[edit]

* A study on the Paleocene charophyte flora from the South Gobi area in the Junggar Basin (China) and on the Paleogene fossil record of charophytes is published by Cao et al. (2022), who interpret their findings as evidence of the dispersal of charophyte lineages from Asia to Europe in the middle to late Eocene, possibly facilitated by waterbirds.[2]

### Chlorophytes

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Ardeiporella[3]

Gen. et comb. nov

Valid

Grgasović

Middle Triassic

|  |

Bosnia and Herzegovina

A green alga belonging to the group Dasycladales. Genus includes "Oligoporella" karrerioidea Pia (1935).

Earltonella[4]

Gen. et sp. nov

|  |

LoDuca in LoDuca et al.

Silurian (Llandovery)

Earlton Formation

Canada
( Ontario)

A green alga belonging to the group Bryopsidales. Genus includes new species E. fredricksi.

Milanovicella? canadillana[5]

Sp. nov

In press

Torromé & Schlagintweit

Late Cretaceous (Santonian–Campanian)

|  |

Spain

A green alga belonging to the group Dasycladales.

Neophysoporella[3]

Gen. et comb. nov

Valid

Grgasović

Late Triassic, possibly also Middle Triassic

|  |

France

A green alga belonging to the group Dasycladales. Genus includes "Diplopora" lotharingica Benecke (1898), "Physoporella" jomdaensis Flügel & Mu (1982) and "Physoporella" zamparelliae Parente & Climaco (1999).

Protocodium[6]

Gen. et sp. nov

|  |

Chai, Aria & Hua

Ediacaran

Dengying Formation

China

A green alga belonging to the family Codiaceae. Genus includes new species P. sinense.

Succodium luciae[7]

Sp. nov

Valid

Vachard & Krainer

Permian-Triassic transition

|  |

Italy

A green alga belonging to the group Dasycladales.

## Lycopodiopsida

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Lepacyclotes ordosensis[8]

Sp. nov

Valid

Deng in Deng et al.

Middle Triassic (Ladinian)

Tongchuan Formation

China

A member of the family Isoetaceae.

Lycopodicaulis[9]

Gen. et sp. nov

Valid

Herrera et al.

Early Cretaceous (late Barremian–early Aptian)

Huolinhe Formation

China

A member of the family Lycopodiaceae. Genus includes new species L. oellgaardii.

Multapicifolium[10]

Gen. et sp. nov

Valid

Edwards, Li & Berry

Early Devonian

|  |

China

A member of Protolepidodendrales of uncertain phylogenetic placement. Genus includes new species M. sinense.

Nothostigma sepeensis[11]

Sp. nov

|  |

Spiekermann, Jasper, Guerra-Sommer & Uhl in Spiekermann et al.

Permian (Cisuralian)

|  |

Brazil

A member of Lycopodiopsida of uncertain affinities.

Omprelostrobus[12]

Gen. et sp. nov

|  |

Liu et al.

Devonian (Famennian)

Wutong Formation

China

A member of Isoetales of uncertain affinities. Genus includes new species O. gigas.

Pleuromeia obovata[13]

Sp. nov

|  |

Deng in Deng et al.

Middle Triassic (Ladinian)

Tongchuan Formation

China

A lycopsid.

Porongodendron[14]

Gen. et sp. nov

Valid

Prestianni et al.

Carboniferous (Mississippian)

|  |

Argentina

An isoetalean lycopsid. Genus includes new species P. minitensis.

Selaginella alata[15]

Sp. nov

|  |

Li & Wang in Li et al.

Cretaceous (Albian-Cenomanian)

Burmese amber

Myanmar

A species of Selaginella.

Selaginella cretacea[16]

Sp. nov

In press

Li et al.

Cretaceous

Burmese amber

Myanmar

A species of Selaginella.

### Lycopsid research

[edit]

* Description of new fossil material of Guangdedendron micrum, providing new information on the morphology of this plant, is published by Gao et al. (2022).[17]
* Xu, Liu & Wang (2022) describe new fossil material of Sublepidodendron grabaui from the Devonian (Famennian) Wutong Formation (China), providing new information on the morphology of the female reproductive organs of this plant.[18]

## Marchantiophyta

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Radula patrickmuelleri[19]

Sp. nov

Valid

Feldberg, Schäfer-Verwimp & Renner in Feldberg et al.

Cretaceous (Albian-Cenomanian)

Burmese amber

Myanmar

A liverwort, a species of Radula.

Radula tanaiensis[19]

Sp. nov

Valid

Feldberg, Schäfer-Verwimp & Renner in Feldberg et al.

Cretaceous (Albian-Cenomanian)

Burmese amber

Myanmar

A liverwort, a species of Radula.

Ricciopsis asturicus[20]

Sp. nov

Valid

Santos et al.

Late Jurassic (Kimmeridgian)

Lastres Formation

Spain

A liverwort belonging to the family Ricciaceae.

Ricciopsis cortaderitaensis[21]

Gen. et sp. nov

Valid

Savoretti et al.

Middle Triassic

|  |

Argentina

A liverwort.

Ricciellites[21]

Sp. nov

Valid

Savoretti et al.

Middle Triassic

|  |

Argentina

A liverwort.
Genus includes new species R. unsaltoensis.

### Marchantiophyta research

[edit]

* New specimens of Radula heinrichsii, providing new information on the morphology of this liverwort, are described from the Cretaceous Burmese amber by Wang et al. (2022).[22]

## Ferns and fern allies

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Acrostichopteris alcainensis[23]

Sp. nov

Valid

Skog & Sender

Early Cretaceous (Albian)

Escucha Formation

Spain

A member of the family Hymenophyllaceae.

Coniopteris antarctica[24]

Sp. nov

In press

Trevisan et al.

Late Cretaceous

|  |

Antarctica

|  |

Diodonopteris virgulata[25]

Sp. nov

In press

Zhou et al.

Early Permian

|  |

China

A botryopteid fern.

Discosoropteris[26]

Gen. et 2 sp. nov

In press

Pšenička et al.

Carboniferous (Pennsylvanian)

Kladno Formation

Czech Republic

A leptosporangiate fern. Genus includes new species D. chlupatum and D. zlatkokvacekii.

Dryopterites beishanensis[27]

Sp. nov

In press

Ren & Sun in Ren et al.

Early Cretaceous

Chijinbao Formation

China

A fern
Announced online in 2022
officially published in 2023

Gleichenia nagalingumiae[28]

Sp. nov

|  |

Cantrill et al.

Miocene

|  |

Australia

A species of Gleichenia.

Hymenophyllites angustus[15]

Sp. nov

|  |

Li & Wang in Li et al.

Cretaceous (Albian-Cenomanian)

Burmese amber

Myanmar

A member of the family Hymenophyllaceae. Originally described as a species of Hymenophyllites, but subsequently moved to the genus Trichomanes sensu lato by Li et al. (2023).[29]

Hymenophyllites kachinensis[15]

Sp. nov

|  |

Li & Wang in Li et al.

Cretaceous (Albian-Cenomanian)

Burmese amber

Myanmar

A member of the family Hymenophyllaceae.

Hymenophyllites setosus[15]

Sp. nov

|  |

Li & Wang in Li et al.

Cretaceous (Albian-Cenomanian)

Burmese amber

Myanmar

A member of the family Hymenophyllaceae.

Microlepia burmasia[30]

Sp. nov

Valid

Long, Wang & Shi in Long et al.

Cretaceous

Burmese amber

Myanmar

A fern of uncertain affinities. Originally described as a dennstaedtiaceous fern belonging to the genus Microlepia, but this classification was contested by Zhang (2024).[31] Published online in 2022, but the issue date of the article naming it is listed as March 2023.

Mikasapteris[32]

Gen. et sp. nov

Valid

Nishida et al.

Late Cretaceous

Yezo Group

Japan

A probable stem polypod leptosporangiate fern. Genus includes new species M. rothwellii.

Paralophosoria[33]

Gen. et sp. nov

Valid

Morales-Toledo, Mendoza-Ruiz & Cevallos-Ferriz

Middle Jurassic

Otlaltepec Formation

Mexico

A member of the family Dicksoniaceae. Genus includes new species P. jurassica.

Phyllotheca douroensis[34]

Sp. nov

|  |

Barbosa et al.

Carboniferous (Gzhelian)

Douro Carboniferous Basin

Portugal

A member of Equisetales.

Scolecopteris zhoui[35]

Sp. nov

In press

Zhang et al.

Early Permian

Taiyuan Formation

China

A member of Marattiales belonging to the family Psaroniaceae.

Wolfeniana[36]

Nom. nov

Valid

Deshmukh

Devonian

Hampshire Group

United States
( West Virginia)

A member of Stauropteridales; a replacement name for Gillespiea Erwin & Rothwell (1989).

### Fern and fern ally research

[edit]

* Pecopteris lativenosa is interpreted as a member of the late Paleozoic marattialean family Psaroniaceae by Li et al. (2022).[37]

## Gnetales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Bassitheca[38]

Gen. et sp. nov

Valid

Manchester et al.

Late Jurassic

Morrison Formation

United States Utah

A gnetale. Genus includes the species B. hoodiorum.

Dichoephedra[39]

Gen. et sp. nov

In press

Ren et al.

Early Cretaceous

Chijinbao Formation

China

A member of the family Ephedraceae. Genus includes new species D. beishanensis.

## Bennettitales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Dictyozamites barnardi[40]

Sp. nov

Valid

Saadatnejad

Late Triassic (Rhaetian)

Kalariz Formation

Iran

A member of Bennettitales.

Dictyozamites fakhri[40]

Sp. nov

Valid

Saadatnejad

Late Triassic (Rhaetian)

Kalariz Formation

Iran

A member of Bennettitales.

Kimuriella[41]

Gen. et sp. nov

Valid

Pott & Takimoto

Late Jurassic (Oxfordian)

Tochikubo Formation

Japan

A member of Bennettitales. Genus includes new species K. densifolia.

Zamites pateri[42]

Sp. nov

Valid

Kvaček

Late Cretaceous (Cenomanian)

Peruc–Korycany Formation

Czech Republic

|  |

## Ginkgophytes

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Glossophyllum lanceolatum[43]

Sp. nov

|  |

Sun & Deng in Sun et al.

Late Triassic

|  |  China |  |

Glossophyllum panii[43]

Sp. nov

|  |

Sun & Deng in Sun et al.

Late Triassic

|  |  China |  |

Pseudotorellia baganuriana[44]

Sp. nov

In press

Nosova & Kostina

Early Cretaceous (Aptian–Albian)

Khuren Dukh Formation

Mongolia

|  |

Pseudotorellia zhoui[45]

Sp. nov

In press

Dong et al.

Middle-Late Jurassic

Daohugou Beds

China

|  |

Umaltolepis zhoui[45]

Sp. nov

In press

Dong et al.

Middle-Late Jurassic

Daohugou Beds

China

|  |

### Ginkgophyte research

[edit]

* Revision of Ginkgo abaniensis, based on data from leaves from the Jurassic Mura Formation (Russia), is published by Frolov & Mashchuk (2022), who emend the diagnosis of this species, and transfer Ginkgo abaniensis, Ginkgo glinkiensis and Ginkgo capillata to the genus Ginkgoites.[46]

## Conifers

[edit]

### Araucariaceae

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Agathoxylon argentinum[47]

Sp. nov

Valid

Bodnar et al.

Late Triassic

Ischigualasto Formation

Argentina

|  |

### Cheirolepidiaceae

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Brachyoxylon yanqingense[48]

Sp. nov

|  |

Cheng et al.

Late Jurassic

Tuchengzi Formation

China

A probable member of the family Cheirolepidiaceae.

Frenelopsis antunesii[49]

Sp. nov

In press

Mendes & Kvaček

Early Cretaceous (Aptian–Albian)

Figueira da Foz Formation

Portugal

A member of the family Cheirolepidiaceae.

Pseudofrenelopsis zlatkoi[50]

Sp. nov

|  |

Kvaček & Mendes

Early Cretaceous (Aptian-Albian)

Figueira da Foz Formation

Portugal

|  |

### Cupressaceae

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Cupressinanthus klebsii[51]

Sp. nov

Valid

Sadowski, Schmidt & Kunzmann

Eocene

|  |

Europe (Baltic Sea region)

Cupressaceous pollen cone.

Patagotaxodia[52]

Gen. et sp. nov

Valid

Andruchow-Colombo et al.

Late Cretaceous (Maastrichtian)

Lefipán Formation

Argentina

A member of the family Cupressaceae. Genus includes new species P. lefipanensis.

### Pinaceae

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Keteleerioxylon changchunense[53]

Sp. nov

|  |

Shi, Sun, Meng & Yu in Shi et al.

Early Cretaceous (Albian)

Yingcheng Formation

China

A Keteleeria-like wood morphogenus.

Nothotsuga mulaensis[54]

Sp. nov

|  |

Li & Dong in Dong et al.

Miocene

Changtai Formation

China

A species of Nothotsuga.

Pinus prehwangshanensis[55]

Sp. nov

|  |

Bazhenova, Wu & Jin in Bazhenova et al.

Late Pleistocene

Maoming Basin

China

A pine.

Pinus shengxianica[56]

Sp. nov

|  |

Li, Hu & Xiao in Li et al.

Miocene

Shengxian Formation

China

A pine.

### Podocarpaceae

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Phyllocladoxylon antarcticum[57]

Sp. nov

Announced

Pujana et al.

Oligocene

San José Formation

Chile

A podocarpaceous wood morphospecies
Announced in 2022
Officially published in 2023

Podocarpoxylon resinosum[57]

Sp. nov

Announced

Pujana et al.

Oligocene

San José Formation

Chile

A podocarpaceous wood morphospecies
Announced in 2022
Officially published in 2023

Podocarpus mexicanoxylon[58]

Sp. nov

|  |

Castañeda-Posadas

Miocene

|  |

Mexico

A species of Podocarpus.

### Sciadopityaceae

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Zhangoxylon[59]

Gen. et sp. nov

In press

Jiang et al.

Middle to Late Jurassic (Callovian to Kimmeridgian)

|  |

China

A member of the family Sciadopityaceae. Genus includes new species Z. yanliaoense.

### Voltziales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Hexicladia[60]

Gen. et sp. nov

Announced

Wang et al.

Permian (Cisuralian)

Shanxi Formation

China

A voltzialean conifer.
The type species is H. yongchangensis.
Announced in 2022
Officially published in 2023

### Other conifers

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Aciphyllum[61]

Gen. et sp. nov

|  |

Barbacka & Górecki in Barbacka et al.

Early Jurassic (Hettangian)

Zagaje Formation

Poland

A needle leaf similar to the leaves of Pinus. Genus includes new species A. triangulatum.

Ductoagathoxylon tsaaganensis[62]

Sp. nov

In press

Cai, Zhang & Feng in Cai et al.

Late Permian

|  |

Mongolia

|  |

Sidashia[63]

Gen. et sp. nov

In press

Forte, Kustatscher & Van Konijnenburg-van Cittert in Forte et al.

Middle Triassic (Anisian)

|  |

Italy

Genus includes new species S. tridentata.

Ullrichia[64]

Gen. et comb. nov

Valid

Kerp et al.

Permian

|  |

Germany

The type species is "Lebachia" laxifolia (1939);
genus also includes "L." intermedia (1939) and "L." mucronata (1939).

### Conifer research

[edit]

* Bodnar et al. (2022) reassess the anatomy and systematics of the permineralized conifer-like woods from the Triassic strata from Argentina, confirm the assignment of the logs related to the families Cupressaceae and Cheirolepidiaceae, as well as three taxa related to Araucariaceae (Agathoxylon cozzoi, Agathoxylon protoaraucana and Agathoxylon argentinum), and argue that the fossil woods previously assigned to the families Podocarpaceae and Taxaceae do not have enough preserved characters to support such assignment.[65]
* A study on the pattern of conifer turnover across the Cretaceous-Paleogene boundary in the Raton and Denver basins (Colorado, United States) is published by Berry (2022).[66]
* Mantzouka, Akkemik & Güngör (2022) describe fossil woods of Cupressinoxylon matromnense from the middle Miocene Eşelek volcanic deposits (Gökçeada, Turkey), preserved with feeding damage produced by members of the agromyzid genus Protophytobia, and supporting the existence of an eastern Mediterranean Miocene Climatic Optimum hotspot which additionally included Greek islands of Lemnos and Lesbos.[67]

## Flowering plants

[edit]

### Chloranthales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Canrightia foveolata[68]

Sp. nov

Valid

Friis et al.

Early Cretaceous (Aptian-Albian)

Almargem Formation

Portugal

|  |

Proencistemon[68]

Gen. et sp. nov

Valid

Friis et al.

Early Cretaceous (Aptian-Albian)

Almargem Formation

Portugal

Genus includes new species P. portugallicus.

### Magnoliids

[edit]

#### Laurales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Araliaephyllum silvapinedae[69]

Sp. nov

In press

Rubalcava-Knoth & Cevallos-Ferriz

Cretaceous (Albian–Cenomanian)

La Cintura Formation

Mexico

|  |

Argapaloxylon salvadorensis[70]

Sp. nov

Valid

Vasquez-Loranca & Cevallos-Ferriz

Miocene

|  |

El Salvador

A member of the family Lauraceae.

Catula[71][72]

Gen. et sp. nov

Valid

Maccracken et al.

Late Cretaceous (Campanian)

Kaiparowits Formation

United States
( Utah)

A member of the family Lauraceae. Genus includes new species C. gettyi.

Cryptocaryoxylon irregularis[73]

Sp. nov

Valid

Akkemik, Iamandei & Çelik

Early Miocene

Hançili Formation

Turkey

Fossil wood of a member of the family Lauraceae.

Laurinoxylon scalariforme[70]

Sp. nov

Valid

Vasquez-Loranca & Cevallos-Ferriz

Miocene

|  |

El Salvador

A member of the family Lauraceae.

Mezilaurinoxylon americana[70]

Sp. nov

Valid

Vasquez-Loranca & Cevallos-Ferriz

Miocene

|  |

El Salvador

A member of the family Lauraceae.

Mezilaurinoxylon draconis[70]

Sp. nov

Valid

Vasquez-Loranca & Cevallos-Ferriz

Miocene

|  |

El Salvador

A member of the family Lauraceae.

Mezilaurinoxylon miocenica[70]

Sp. nov

Valid

Vasquez-Loranca & Cevallos-Ferriz

Miocene

|  |

El Salvador

A member of the family Lauraceae.

#### Magnoliales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Fissistigma himachalensis[74]

Sp. nov

Valid

Singh et al.

Miocene

|  |

India

A species of Fissistigma.

Magnolia allasoniae[75]

Sp. nov

Valid

Martinetto in Niccolini et al.

Miocene (Messinian)

Piedmont Basin

Italy

A species of Magnolia.
First named in 1995 but did not meet ICBN requirements; subsequently validated in 2022.

#### Piperales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Aristospermum[76]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Aptian–Albian)

|  |

Portugal
United States
( Virginia)

A member of the family Aristolochiaceae. Genus includes new species A. huberi.

Siratospermum[76]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Late Cretaceous (Cenomanian)

|  |

United States
( Maryland)

A member of the family Aristolochiaceae. Genus includes new species S. mauldinense.

### Monocots

[edit]

#### Lilioid monocots

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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Pandanus estellae[77]

Sp. nov

Valid

Rozefelds et al.

Oligocene

|  |

Australia

A species of Pandanus.

#### Commelinid monocots

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Sabalites ghughuaensis[78]

Sp. nov

In press

Kumar, Hazra & Khan in Kumar et al.

Late Cretaceous-Paleocene (Maastrichtian-Danian)

Deccan Intertrappean Beds

India

A member of the family Arecaceae belonging to the subfamily Coryphoideae.

Sabalites umariaensis[78]

Sp. nov

In press

Kumar, Hazra & Khan in Kumar et al.

Late Cretaceous-Paleocene (Maastrichtian-Danian)

Deccan Intertrappean Beds

India

A member of the family Arecaceae belonging to the subfamily Coryphoideae.

#### Monocot research

[edit]

* Leaf fossils of costapalmate-palms belonging to the genus Sabalites are described from the ?Santonian–Campanian Belly River Group, Campanian Foremost Formation (Alberta, Canada) and Maastrichtian Frenchman Formation (Saskatchewan, Canada) by Greenwood, Conran & West (2022), who interpret the studied fossils as constraining climate reconstructions for the Late Cretaceous high mid-latitudes of North America (c. 55° N) to exclude significant freezing episodes; the authors also transfer the Late Cretaceous species "Geonomites" imperialis to the genus Phoenicites, and reassess Sabalites carolinensis as more likely to be Campanian than Coniacian–Santonian in age.[79]
* A study on the impact of the absence of megaherbivores in the aftermath of the Cretaceous–Paleogene extinction event on the evolution of palms is published by Onstein, Kissling & Linder (2022).[80]
* A study on the evolutionary history of palms belonging to the group Mauritiinae, as inferred from a phylogenetic analysis incorporating fossil data, is published by Bacon et al. (2022).[81]

### Basal eudicots

[edit]

#### Proteales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Distefananthus[82]

Gen. et sp. nov

|  |

Huegele & Wang

Early Cretaceous (Albian)

Dakota Formation

United States
( Kansas)

A platanaceous inflorescence. Genus includes new species D. hoisingtonensis.

Langeranthus[83]

Gen et sp nov

Valid

Huegele & Manchester

Eocene
Ypresian

Klondike Mountain Formation

USA
Washington

A platanaceous flowering head.
The type species is L. dillhoffiorum

Meliosma eosinica[84]

Sp. nov

|  |

Moiseeva, Kodrul & Jin
in Moiseeva et al.

Late Eocene

Huangniuling Formation

China

A species of Meliosma.

Nelumbo delinghaensis[85]

Sp. nov

|  |

Luo & Jia in Luo et al.

Miocene

Upper Youshashan Formation

China

A species of Nelumbo.

Nelumbo fujianensis[86]

Sp. nov

In press

Dong et al.

Miocene

Fotan Group

China

A species of Nelumbo.

Platimeliphyllum durhamensis[87]

Comb nov

in press

(Wolfe)

Late Eocene

Puget Group
"Upper Fultonian" Loc 9832

USA
Washington

A platanaceous leaf.
Moved from "Fothergilla" durhamensis (1968).

Platimeliphyllum fushunensis[87]

Comb nov

in press

(Chen)

Eocene

Fushun Formation

China

A platanaceous leaf.
Moved from "Betula" fushunensis.

Sapindopsis chinensis[88]

Sp. nov

|  |

Golovneva et al.

Early Cretaceous (Albian)

Dalazi Formation

China

A member of the family Platanaceae.

Sapindopsis orientalis[88]

Sp. nov

|  |

Golovneva et al.

Early Cretaceous (Albian)

Frentsevka Formation

Russia
( Primorsky Krai)

A member of the family Platanaceae.

##### Protealean research

[edit]

* Redescription of the Okanagan Highlands genus Langeria with description of associated stipules and reproductive structures plus formal reassignment of the genus to Platanaceae by Huegele & Manchester is published.[83]

#### Ranunculales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Berberis auriolensis[89]

Sp. nov

Valid

Denk & Sami in Denk et al.

Pleistocene (Calabrian)

|  |

Italy

A species of Berberis.

Berberis siwalica[74]

Sp. nov

Valid

Singh et al.

Miocene

|  |

India

A species of Berberis.

Mahonia mangbangensis[90]

Sp. nov

|  |

Tang et al.

Pliocene

Mangbang Formation

China

A species of Mahonia.

Palaeosinomenium hengduanensis[91]

Sp. nov

|  |

Wu & Zhou in Wu et al.

Eocene

Shuanghe Formation

China

A member of the family Menispermaceae.

### Superasterids

[edit]

#### Aquifoliales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Ilex antiquorum[92]

Nom. nov

Valid

Doweld

Late Cretaceous (Maastrichtian)

|  |

Germany

A holly; a replacement name for Ilex antiqua Knobloch & Mai (1986).

Ilex myricina[92]

Nom. nov

Valid

Doweld

Miocene (Messinian)

|  |

Italy

A holly; a replacement name for Ilex myricoides Massalongo (1858).

#### Caryophyllales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Podopterus mijangosae[93]

Sp. nov

In press

Estrada-Ruiz

Miocene

Mexican amber

Mexico

A species of Podopterus.

#### Cornales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Blackwelloxylon[94]

Nom. nov

Valid

Deshmukh

Pleistocene

|  |

United States
( Mississippi)

A member of the family Cornaceae; a replacement name for Cornoxylon Blackwell (1982).

Exbeckettia[95]

Gen. et comb. nov

Valid

Manchester & Collinson

Early Eocene

London Clay

United Kingdom

A mastixioid fruit; a new genus for Beckettia mastixioides Reid & Chandler (1933).

#### Dipsacales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Sambucus heqingensis[96]

Sp. nov

In press

Huang & Zhou in Huang et al.

Late Pliocene

Heqing Basin

China

A species of Sambucus.

#### Ericales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Paradiospyroxylon[97]

Gen. et sp. nov

In press

Koutecký & Sakala in Koutecký, Sakala & Chytrý

Oligocene

Ústí Formation

Czech Republic

A member of the family Ebenaceae. Genus includes new species P. kvacekii.

#### Icacinales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Palaeophytocrene chicoensis[98]

Sp. nov

|  |

Atkinson

Late Cretaceous (Campanian)

Chico Formation

United States
( California)

A member of the family Icacinaceae.

#### Metteniusales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Calatola verae[99]

Sp. nov

|  |

Estrada-Ruiz et al.

Miocene

Mexican amber

Mexico

A species of Calatola.

### Superrosids

[edit]

#### Cucurbitales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Libasperma[100]

Gen. et sp. nov

In press

Huegele & Manchester

Paleocene

Fort Union Formation

United States
( Montana)

A member of the family Cucurbitaceae. Genus includes new species L. potamoglossensis.

#### Fabales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Albizia yenbaiensis[101]

Sp. nov

Announced

Nguyen, Su & J. Huang in Nguyen et al.

Miocene

Yen Bai Basin

Vietnam

An Albizia species.
Announced in 2022
Officially published January 2023

Anadenantheroxylon kurupaum[102]

Sp. nov

Valid

Ramos et al.

Late Pleistocene

El Palmar Formation

Argentina

A member of the family Fabaceae.

Cedrelinga paleocatenaeformis[102]

Sp. nov

Valid

Ramos et al.

Late Pleistocene

El Palmar Formation

Argentina

A species of Cedrelinga.

Cercioxylon mediterraneum[73]

Sp. nov

Valid

Akkemik, Iamandei & Çelik

Early Miocene

Hançili Formation

Turkey

Fossil wood of a member of the family Fabaceae.

Chloroleucoxylon[102]

Gen. et sp. nov

Valid

Ramos et al.

Late Pleistocene

El Palmar Formation

Argentina

A member of the family Fabaceae. Genus includes new species C. yukeriense.

Enterolobiumoxylon vassalloae[102]

Sp. nov

Valid

Ramos et al.

Late Pleistocene

El Palmar Formation

Argentina

A member of the family Fabaceae.

Leguminocarpum meghalayensis[103]

Sp. nov

Announced

Bhatia, Srivastava & Mehrotra

Late Paleocene

Tura Formation

India

A fabaceous seed pod morphospecies.
Announced in 2022
Officially published in 2023

Microlobiusxylon parafoetidus[102]

Sp. nov

Valid

Ramos et al.

Late Pleistocene

El Palmar Formation

Argentina

A member of the family Fabaceae.

Paleobowdichia[104]

Gen. et comb. nov

Valid

Herendeen et al.

Latest Paleocene to late early Eocene

Lamar River Formation

United States
( Colorado
Wyoming)

A member of Papilionoideae; a new genus for "Acacia" lamarensis Knowlton (1899).

Parapiptadenioxylon[102]

Gen. et sp. nov

Valid

Ramos et al.

Late Pleistocene

El Palmar Formation

Argentina

A Fabaceae genus.
The type species is P. pararigida.

Parvileguminophyllum damalgiriensis[103]

Sp. nov

Announced

Bhatia, Srivastava & Mehrotra

Late Paleocene

Tura Formation

India

A fabaceous leaf morphospecies.
Announced in 2022
Officially published in 2023

Podocarpium tibeticum[105]

Sp. nov

In press

Li, Huang & Su in Li et al.

Late Eocene

Lunpola Basin

China

A member of the family Fabaceae.

Pseudopiptadenioxylon[102]

Gen. et sp. nov

Valid

Ramos et al.

Late Pleistocene

El Palmar Formation

Argentina

A member of the family Fabaceae. Genus includes new species P. uniseriatum.

Tobya[104]

Gen. et comb. nov

Valid

Herendeen et al.

Eocene

Cockfield Formation

United States
( Kentucky
Tennessee)

A member of Papilionoideae; a new genus for "Diplotropis" claibornensis Herendeen & Dilcher (1990).

##### Fabalean research

[edit]

* New fossil material of members of the genus Bauhinia is described from the Eocene of the Puyang Basin (China) by Jia et al. (2022), who interpret their findings as the earliest reliable fossil records of Bauhinia in Asia.[106]
* Moya et al. (2022) study the affinities of fossil legumes Entrerrioxylon victoriensis, Gossweilerodendroxylon palmariensis, Paraoxystigma concordiensis and Cylicodiscuxylon paragabunensis from the Cenozoic Paraná, Arroyo Feliciano and El Palmar formations (Argentina) with extant West African legumes, and discuss the possible migration routes by which these plants may have arrived in South America from Africa.[107]

#### Fagales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

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|
|

Castanopsis zhoui[108]

Sp. nov

In press

Wang et al.

Miocene

Fotan Group

China

A species of Castanopsis.

Comptonia hirsuta[109]

Sp. nov

|  |

Xiao & Ji in Ji et al.

Miocene

Hannuoba Formation

China

A species of Comptonia.

Myricoxylon doganyurtensis[73]

Sp. nov

Valid

Akkemik, Iamandei & Çelik

Early Miocene

|  |

Turkey

Fossil wood of a member of the family Myricaceae.

Nothofagoxylon ruei[57]

Sp. nov

Announced

Pujana et al.

Oligocene

San José Formation

Chile

A nothofagaceous wood morphospecies
Announced in 2022
Officially published in 2023

Pterocarya magnifructa[110]

Sp. nov

Valid

Stults, Tiffney & Axsmith

Pliocene

|  |

United States
( Alabama)

A species of Pterocarya.

Quercus nanningensis[111]

Sp. nov

In press

Liu & Jin in Liu et al.

Late Oligocene

Yongning Formation

China

An oak.

Quercus paleodisciformis[111]

Sp. nov

In press

Liu & Jin in Liu et al.

Late Oligocene

Yongning Formation

China

An oak.

Quercus paleohui[111]

Sp. nov

In press

Liu & Jin in Liu et al.

Late Oligocene

Yongning Formation

China

An oak.

Quercus yongningensis[111]

Sp. nov

In press

Liu & Jin in Liu et al.

Late Oligocene

Yongning Formation

China

An oak.

#### Malpighiales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Belenocarpa[112]

Gen. et comb. nov

Valid

Hamersma et al.

Early Oligocene

|  |

Peru

A member of the family Euphorbiaceae; a new genus for "Jatropha" tertiara Berry.

Elatine odgaardii[113]

Sp. nov

Valid

Bennike in Bennike et al.

Probably early Pleistocene

|  |

Greenland

A species of Elatine. Announced in 2022; the final article version was published in 2023.

Mammeoxylon beylikduezuense[114]

Sp. nov

In press

Akkemik et al.

Late Oligocene-Early Miocene

İstanbul Formation

Turkey

A Mammea relative wood morphospecies

Mammeoxylon paramericana[114]

Comb. nov

In press

(Nelson & Jud) Akkemik & D. Mantzouka

Miocene

|  |

Panama

A Mammea relative wood morphospecies
Moved from Mammea paramericana (2017)

Parinari hilliana[115]

Sp. nov

Valid

Grote in Grote, Duangkrayom & Jintasakul

Late Miocene

Tha Chang beds

Thailand

A species of Parinari.

Parinari khoratensis[115]

Sp. nov

Valid

Grote in Grote, Duangkrayom & Jintasakul

Late Miocene

Tha Chang beds

Thailand

A species of Parinari.

Plukenetia minima[116]

Sp. nov

In press

Poinar

Miocene

Dominican amber

Dominican Republic

A species of Plukenetia.

#### Malvales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Malvacipolloides deccanensis[117]

Sp. nov

|  |

Manchester et al.

Late Cretaceous-Paleocene (Maastrichtian–Danian)

Deccan Intertrappean Beds

India

A member of the family Malvaceae.

Malvacipolloides intertrappea[117]

Sp. nov

|  |

Manchester et al.

Late Cretaceous-Paleocene (Maastrichtian–Danian)

Deccan Intertrappean Beds

India

A member of the family Malvaceae.

Thespesia neopopulnea[118]

Sp. nov

Valid

Hazra, Mahato & Khan in Hazra et al.

Pliocene

Rajdanda Formation

India

A species of Thespesia.

##### Malvalean research

[edit]

* A study on the evolutionary history of Dipterocarpaceae, as indicated by biogeography of pollen fossils from Africa and India, molecular data and fossil amber records, is published by Bansal et al. (2022).[119]

#### Myrtales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Hemitrapa zhangpuensis[120]

Sp. nov

|  |

Dong et al.

Miocene

|  |

China

A member of the family Lythraceae belonging to the subfamily Trapoideae.

Lagerstroemioxylon compactum[121]

Sp. nov

Valid

Harsh & Shekhawat

Eocene

|  |

India

A member of the family Lythraceae.

Lagerstroemioxylon delicatum[121]

Sp. nov

Valid

Harsh & Shekhawat

Eocene

|  |

India

A member of the family Lythraceae.

Myrtineoxylon hoffmannae[57]

Sp. nov

Announced

Pujana et al.

Oligocene

San José Formation

Chile

A myrtaceous wood morphospecies.
Announced in 2022
Officially published in 2023

Trapa natanifolia[122]

Sp. nov

|  |

Han & Jia in Han et al.

Late Eocene

Bailuyuan Formation

China

A water caltrop.

Trapa qaidamensis[123]

Sp. nov

|  |

Cai et al.

Miocene

Shangyoushashan Formation

China

A water caltrop.

#### Oxalidales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Ceratopetalum suciensis[124]

Sp nov

In press

Tang, Smith, & Atkinson

Late Cretaceous
Campanian

Cedar District Formation

United States
( Washington)

A Cunoniaceous species.

Connaroxylon[125]

Gen. et sp. nov

Valid

Baas et al.

Cretaceous
Maastrichtian-earliest Paleocene

Deccan Intertrappean Beds

India

A probable Connaraceous wood morphotaxon.
The type species is C. dimorphum.
First named in 2017 but failed ICBN requirements;[126]
subsequently validated in 2022.[125]

Cunoniocarpa[127]

Gen. et sp. nov

Valid

Matel et al.

Early Eocene

Huitrera Formation

Argentina

A member of Cunoniaceae.
The type species is C. stylosa.

Racemofructus[127]

Gen. et sp. nov

Valid

Matel et al.

Early Eocene

Huitrera Formation

Argentina

A member of Cunoniaceae.
The type species is R. fasciculatus.

Weinmannioxylon trichospermoides[57]

Sp. nov

Announced

Pujana et al.

Oligocene

San José Formation

Chile

A cunoniaceous wood morphospecies.
Announced in 2022
Officially published in 2023

##### Oxalidalean research

[edit]

Tand, Smith, and Atkinson describe the first North American instance of the previously Paleo-Antarctic Rainforest Lineage Cunoniaceae fruits from Sucia Island. Previously considered solely a Gondwanan family, the new species indicate a complex geographic history for the group.[124]

#### Rosales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Eophylica[128]

Gen. et sp. nov

Valid

Shi et al.

Cretaceous

Burmese amber

Myanmar

A rhamnaceous floral morphotaxon.
The type species is E. priscastellata.

Ficoxylon fusiforme[129]

Sp. nov

Valid

El-Noamani

Late Cretaceous

Taref Formation

Egypt

A member of the family Moraceae.

Ficus fujianensis[130]

Sp. nov

In press

Dong et al.

Miocene

|  |

China

A species of Ficus.

Ficus zhangpuensis[130]

Sp. nov

In press

Dong et al.

Miocene

|  |

China

A species of Ficus.

Ventilago pliocenica[131]

Sp. nov

|  |

Hazra et al.

Pliocene

|  |

India

A species of Ventilago.

Ventilago siwalika[131]

Sp. nov

|  |

Hazra et al.

Miocene

|  |

India

A species of Ventilago.

#### Sapindales

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Barkleya[132]

Gen. et comb. nov

Valid

Manchester & Judd

Eocene
Ypresian

Green River Formation

United States
Colorado

An anacardiaceous samara.
The type species is B. schinoloxa.
Moved from Anacardites schinoloxus (1929).
Possibly the fruits of "Rhus" nigricans.

Canarium haominiae[133]

Sp. nov

In press

Yin et al.

Miocene

|  |

China

A species of Canarium.

Canarium maomingense[134]

Sp. nov

|  |

Xiang & Jin in Xiang et al.

Late Pleistocene

Maoming Basin

China

A species of Canarium.

Choerospondias mioaxillaris[135]

Sp. nov

|  |

Xiao & Wu in Xiao et al.

Miocene

Shengxian Formation

China

A species of Choerospondias.

Choerospondias tiantaiensis[135]

Sp. nov

|  |

Xiao & Wu in Xiao et al.

Miocene

Shengxian Formation

China

A species of Choerospondias.

Grimmipollis[136]

Gen. et sp. nov

|  |

Huang, Morley & Hoorn in Huang et al.

Eocene

Yaw Formation

Myanmar

A member of the family Sapindaceae. Genus includes new species G. burmanica.

Koelreuteria kvacekii[137]

Sp. nov

Valid

Chen, Del Rio & Su in Chen et al.

Eocene

Niubao Formation

China

A species of Koelreuteria.

Loxopteroides[138]

Gen. et sp. nov

In press

Manchester & Judd

Eocene

Ione Formation

United States
( California
Oregon)

A member of the family Anacardiaceae. Genus includes new species L. weeksae.

Vaudoisia[139]

Gen. et comb. nov

Valid

Strullu-Derrien et al.

Eocene

|  |

France

A fruit of likely sapindalean affinity; a new genus for "Juglandicarya" gruetii Vaudois-Miéja (1976).

#### Other Eudicots

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Zlatkovia[140]

Gen. et sp. nov

In press

Rothwell & Stockey

Late Cretaceous

St. Mary River Formation

Canada
( Alberta)

An aquatic eudicot.
The type species is Z. crenulata.

### Other angiosperms

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Archaebuda[141]

Gen. et sp. nov

|  |

Chen & Wang

Early Cretaceous (Barremian-Aptian)

Yixian Formation

China

A flower bud of an early angiosperm. Genus includes new species A. lingyuanensis.

Ascarinophyllum[142]

Gen. et sp. nov

Announced 2022

Čepičková & Kvaček

Late Cretaceous
(Cenomanian)

Peruc–Korycany Formation

Czech Republic

A Basal angiosperm leaf morphogenus
Similar to Mesodescolea plicata and Chloranthaceae.
The type species is A. pecinovense.
Officially published in 2023

Covidifructus[143]

Gen. et sp. nov

Valid

Heřmanová et al.

Late Cretaceous (late Turonian-Santonian)

Klikov Formation

Czech Republic

An angiosperm fruit of uncertain affinities, with similarities to the family Dilleniaceae.
The type species is C. multicarpellatus.

Elasmostemon[68]

Gen. et sp. nov

Valid

Friis et al.

Early Cretaceous (Aptian-Albian)

Almargem Formation

Portugal

A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species E. paisii.

Endressistemon[68]

Gen. et sp. nov

Valid

Friis et al.

Early Cretaceous (Aptian-Albian)

Almargem Formation

Portugal

A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species E. cateficensis.

Fairlingtonia microgyna[144]

Sp. nov

|  |

Du et al.

Early Cretaceous

Zhonggou Formation

China

A herbaceous eudicot.

Florigerminis[145]

Gen. et sp. nov

In press

Cui et al.

Middle-Late Jurassic

Jiulongshan Formation

China

A possible flower bud.
The type species is F. jurassica.
First announced online 2021, Final article published in 2022.

|  Florigerminis jurassica

Gansupeltata[146]

Gen. et sp. nov

Valid

Wu et al.

Early Cretaceous (Aptian)

Chijinpu Formation

China

An early flowering plant. Genus includes new species G. beishanensis.

Herbifolia[147]

Gen. et sp. nov

In press

Frolov & Enushchenko

Middle Jurassic (Aalenian)

Irkutsk Coal Basin

Russia

An angiosperm with leaf epidermal structure most similar to those of modern Asparagales and Liliales. Genus includes new species H. antiqua.

Honeytheca[100]

Gen. et sp. nov

In press

Huegele & Manchester

Paleocene

Fort Union Formation

United States
( Montana)

A flowering plant of uncertain affinities. Genus includes new species H. bighornensis.

Ibericarpus[68]

Gen. et sp. nov

Valid

Friis et al.

Early Cretaceous (Aptian-Albian)

Almargem Formation

Portugal

A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species I. cuneiformis.

Lingyuananthus[148]

Gen. et sp. nov

|  |

Wang

Early Cretaceous (Barremian–Aptian)

Yixian Formation

China

An early angiosperm. Genus includes new species L. inexpectus.

Phylica piloburmensis[128]

Sp. nov

Valid

Shi et al.

Cretaceous

Burmese amber

Myanmar

A flowering plant of uncertain affinities. Originally described as a species of Phylica. Oskolski et al. (2024) interpreted it as a flowering plant with an affinity to Rhamnaceae, possibly to an extinct basal lineage;[149] on the other hand Beurel et al. (2024) interpreted it as more likely to have lauralean affinities, and made it the type species of the separate genus Nothophylica.[150]

Santaniella[151]

Gen. et 2 sp. nov

|  |

Gobo et al.

Early Cretaceous (Barremian-Aptian)

Crato Formation

Brazil

Originally described as a member or a relative of the family Ranunculaceae, but subsequently considered to be a mesangiosperm of uncertain affinities, possibly a magnoliid.[152] Genus includes new species S. lobata and S. acuta.

Todziaphyllum saportanum[142]

Comb. nov

Announced

(Velenovský) Čepičková & Kvaček

Late Cretaceous
(Cenomanian)

Peruc–Korycany Formation

Czech Republic

A Basal angiosperm leaf morphogenus
A new combination for Banksites saportanus
Officially published in 2023

Tolmania[153]

Gen. et sp. nov

Valid

Edmonds, Stockey & Rothwell

Late Cretaceous (Maastrichtian)

St. Mary River Formation

Canada
( Alberta)

An aquatic dicot. Genus includes new species T. aquatica.

Valvidistemon[68]

Gen. et sp. nov

Valid

Friis et al.

Early Cretaceous (Aptian-Albian)

Almargem Formation

Portugal

A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species V. globiferus.

### General angiosperm research

[edit]

* Surangea mohgaoensis, originally interpreted as fern megaspores, is reinterpreted as angiosperm fruits by Ramteke et al. (2022).[154]
* Zhang et al. (2022) describe rich assemblages of spiny plant fossils from the Eocene (Bartonian) Niubao Formation (Tibet, China), preserving seven different spine morphologies, and interpret this finding as evidence of the presence of a diversity of spiny plants in Eocene central Tibet, as well as evidence of a rapid diversification of spiny plants in Eurasia around that time.[155]
* A preliminary report on a new fossil angiosperm flora of the Lesvos Petrified Forest at Akrocheiras east of Sigri on Lesbos, Greece is given by Kafetzidou et al. Preliminary taxa identifications are given and commentary on the climactic implications are made.[156]
* A study aiming to determine the relationship between past atmospheric CO2 and temperature fluctuations and the shifts in diversification rates of Poaceae and Asteraceae is published by Palazzesi et al. (2022).[157]

## Other plants

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Aegianthus irkutensis[158]

Sp. nov

In press

Nosova & Tekleva

Middle Jurassic

Prisayan Formation

Russia

Pollen cone with pollen of ginkgoalean or gnetophytalean affinity.

Aysenoxylon[57]

Gen et sp nov

Announced

Pujana et al.

Oligocene

San José Formation

Chile

A wood morphospecies of uncertain affinity.
The type species is A. patorarensis.
Announced in 2022
Officially published in 2023

Bryokhutuliinia ignatovii[159]

Sp. nov

Valid

Frolov, Kazanovsky & Enushchenko

Early Jurassic (Toarcian)

Middle Subformation of Prisayan Formation

Russia
( Irkutsk Oblast)

A member of Bryopsida of uncertain affinities.

Combina[160]

Gen. et sp. nov

|  |

Santos & Wang

Middle Triassic (Anisian)

Calcena Formation

Spain

A cone-like reproductive organ of a seed plant. Genus includes new species C. triassica.

Dioonitocarpidium rossicum[161]

Sp. nov

Valid

Gomankov

Permian

|  |

Russia

A member of Cycadales.

Europoxylon garapensis[162]

Sp. nov

In press

Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

Brazil

A gymnosperm.

Jarudia[163]

Gen. et sp. nov

|  |

Shi et al.

Early Cretaceous

Huolinhe Formation

China

A seed-bearing structure of a corystosperm seed fern. Genus includes new species J. zhoui.

Komlopteris distinctiva[61]

Sp. nov

|  |

Barbacka in Barbacka et al.

Early Jurassic (Hettangian)

Zagaje Formation

Poland

Cuticle of a seed fern.

Lesleya ceriacoi[164]

Sp. nov

In press

Correia et al.

Carboniferous (Gzhelian)

Douro Carboniferous Basin

Portugal

An early gymnosperm.

Palaeodichelyma kiritchkovae[159]

Sp. nov

Valid

Frolov, Kazanovsky & Enushchenko

Early Jurassic (Pliensbachian)

Lower Subformation of Prisayan Formation

Russia
( Irkutsk Oblast)

A member of Bryopsida of uncertain affinities.

Paragigantopteris[165]

Gen. et sp. nov

In press

Ma et al.

Permian (Wuchiapingian)

Lungtan Formation

China

A gigantopterid. Genus includes new species P. qingloongensis.

Pauthecophyton hezhangensis[166]

Sp. nov

|  |

Wang et al.

Devonian (Pragian-Emsian)

Danlin Formation

China

A euphyllophyte of uncertain affinities.

Piterophyton[167]

Gen. et sp. nov

Valid

Naugolnykh

Ordovician

|  |

Russia
( Leningrad Oblast)

A rhyniophyte of uncertain affinities. The type species is P. caudatum.

Polycanaloxylon[162]

Gen. et sp. nov

In press

Conceição et al.

Permian (Cisuralian)

Pedra de Fogo Formation

Brazil

A gymnosperm. Genus includes new species P. merlottii.

Psilophyton diakanthon[168]

Sp. nov

|  |

Colston, Landaw & Tomescu

Devonian (Emsian)

Battery Point Formation

Canada
( Quebec)

A member of the group Trimerophytopsida.

Renbernia[169]

Gen. et sp. nov

In press

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

United States
( Virginia)

A seed plant similar to Brenneria potomacensis. Genus includes new species R. zhoui.

Rhyniotaenium[170]

Gen. et sp. nov

In press

Krings

Early Devonian

Rhynie chert

United Kingdom

An alga, probably a green alga belonging to the family Mesotaeniaceae. Genus includes new species R. velatum.

Sinoglossa[171]

Gen. et sp. nov

Valid

Zhang et al.

Middle Triassic

Linjia Formation

China

A member of Glossopteridales. The type species is S. sunii.

Taimyria[172]

Gen. et sp. nov

Valid

Naugolnykh & Mogutcheva

Early Triassic (Induan)

Keshinskian/Keshinskaya Formation

Russia
( Krasnoyarsk Krai)

A member of Peltaspermales belonging to the family Angaropeltaceae. Genus includes new species T. triassica.

Taungurungia[173]

Gen. et sp. nov

Valid

McSweeney, Shimeta & Buckeridge

Devonian (Pragian–Emsian)

Norton Gully Sandstone Formation

Australia

A plant of uncertain affinities, similar to members of Zosterophyllopsida. Genus includes new species T. garrattii.

Teyoua[174]

Gen. et sp. nov

In press

Huang, Liu & Xue

Devonian (probably Pragian)

Mangshan Group

China

A polysporangiate land plant. Genus includes new species T. antrorsa.

Traskia[175]

Gen. et sp. nov

Valid

Rothwell et al.

Jurassic

|  |

Canada
( British Columbia)

A stem-cycad. Genus includes new species T. maahlae.

Vetiplanaxis obtusus[176]

Sp. nov

In press

Li et al.

Cretaceous

Burmese amber

Myanmar

A moss belonging to the group Hypnodendrales.

Wilhowia[177]

Gen. et sp. nov

Valid

Gensel

Devonian (Emsian)

Battery Point Formation

Canada
( Quebec)

A basal euphyllophyte. Genus includes new species W. phocarum.

Xadzigacalix[178]

Gen. et sp. nov

Valid

Klymiuk, Rothwell & Stockey

Early Cretaceous (Valanginian)

|  |

Canada
( British Columbia)

A gymnosperm of uncertain phylogenetic placement, possibly having affinities with gnetophytes or angiosperms. Genus includes new species X. quatsinoensis.

Xinhangia[179]

Gen. et sp. nov

|  |

Yang & Wang

Devonian (Famennian)

Wutong Formation

China

A fern-like plant of uncertain affinities. Genus includes new species X. spina.

### Other plant research

[edit]

* A study on the xylem development in Leptocentroxyla, and on its implications for the knowledge of the evolution of pith, is published by Tomescu & McQueen (2022).[180]
* Decombeix et al. (2022) report evidence of tylosis formation in permineralized wood of Dameria hueberi from the Tournaisian of Australia.[181]
* The first comprehensive crown reconstruction of Medullosa stellata var. typica, based on data from a specimen from the Chemnitz petrified forest (Germany), is presented by Luthardt et al. (2022).[182]
* Fossil material of Rhabdotaenia is reported from the Permian Umm Irna Formation (Jordan) by Blomenkemper et al. (2022), representing the northernmost occurrence of this Gondwanan leaf type reported to date.[183]

## Palynology

[edit]

Name  | Novelty  | Status  | Authors  | Age  | Type locality  | Location  | Notes  | Images

|
|
|
|
|
|
|
|

Anapiculatisporites radiatus[184]

Sp. nov

Valid

Playford

Carboniferous (Mississippian)

Lyall Formation

Australia

A trilete spore.

Apricasporites[184]

Gen. et sp. nov

Valid

Playford

Carboniferous (Mississippian)

Lyall Formation

Australia

A trilete spore. Genus includes new species A. cancellosus.

Camptotriletes inaequabilis[184]

Sp. nov

Valid

Playford

Carboniferous (Mississippian)

Lyall Formation

Australia

A trilete spore.

Camptotriletes suggrandis[184]

Sp. nov

Valid

Playford

Carboniferous (Mississippian)

Lyall Formation

Australia

A trilete spore.

Convolutispora inreligata[184]

Sp. nov

Valid

Playford

Carboniferous (Mississippian)

Lyall Formation

Australia

A trilete spore.

Endosporites circumsaeptus[184]

Sp. nov

Valid

Playford

Carboniferous (Mississippian)

Lyall Formation

Australia

A trilete spore.

Foveosporites magnus[184]

Sp. nov

Valid

Playford

Carboniferous (Mississippian)

Lyall Formation

Australia

A trilete spore.

Granulatisporites commutabilis[184]

Sp. nov

Valid

Playford

Carboniferous (Mississippian)

Lyall Formation

Australia

A trilete spore.

Paxillitriletes permicus[185]

Sp. nov

In press

Sui, McLoughlin & Feng in Sui et al.

Permian (Wuchiapingian–Changhsingian)

Xuanwei Formation

China

A lycophyte megaspore.

Sergipea multipapillata[186]

Sp. nov

|  |

Hu et al.

Early Cretaceous

Bongor Basin

Chad

A gymnosperm pollen.

Sparganiaceaepollenites reticulatus[187]

Sp. nov

Junior homonym

Samant et al.

Late Cretaceous (Maastrichtian)

Mandla Formation

India

Pollen of a member of the family Typhaceae/Sparganiaceae. The name is preoccupied by Sparganiaceaepollenites reticulatus Doktorowicz-Hrebnicka (1960) ex Krutzsch & Vanhoorne (1977); DeBenedetti et al. (2025) coined a replacement name Sparganiaceaepollenites intertrappeansis.[188]

Volkheimerites[189]

Gen. et sp. nov

In press

Narváez et al.

Paleocene (Danian)

Salamanca Formation

Argentina

Pollen of a flowering plant.
The type species is V. labyrinthus.

Yezopollis[190]

Gen. et sp. nov

In press

Legrand, Yamada & Nishida

Late Cretaceous (Cenomanian–Turonian)

Mikasa Formation

Japan

A Normapolles-type flowering plant pollen. Genus includes new species Y. mikasaensis.

## Research

[edit]

* Review of the studies on the origin of the land flora is published by Bowman (2022).[191]
* A study on the evolution of body plans of members of Viridiplantae, based on a review of the fossil record, molecular data and developmental biology, is published by Niklas & Tiffney (2022).[192]
* A study on the biodiversity of land plants at the equator during their first major diversification in the Late Silurian–Early Devonian is published by Wellman et al. (2022).[193]
* A study on the evolution of heterospory during the Devonian is published by Leslie & Bonacorsi (2022).[194]
* Seven coniferous nurse logs that have been colonized by conifer and equisetalean roots are reported from four Permian intervals in the Ordos Basin (China) by Feng et al. (2022), indicating that conifer tree stems probably functioned as hosts to both conspecific and interspecific seedlings in the Cathaysian Flora.[195]
* A study on the impact of the Intertropical Convergence Zone in the emerging South Atlantic region on Aptian plant communities from eight Brazilian sedimentary basins is published by Carvalho et al. (2022), who report evidence of an overall predominance of xerophytic plants, attesting to more dry conditions, and of a humidification trend towards the end of the late Aptian resulting in the predominance of hydrophytes, hygrophytes, tropical lowland flora and upland flora, indicative of prevalence of lowland and montane rainforests.[196]
* A study on the distribution and relative abundances of major plant groups from the Albian Gates Formation (Alberta, Canada) is published by Kalyniuk et al. (2022).[197]
* A study on the relationship between whole-genome duplication, seed traits and the selectivity of the survival of plants during the Cretaceous–Paleogene extinction event is published by Berry & Jaganathan (2022).[198]
* New Oligocene flora is described from the Dong Ho Formation (Vietnam) by Huang et al. (2022), who interpret the studied fossils as evidence of long-term environmental, floristic and vegetational stability in this region since the Paleogene.[199]
* Gentis et al. (2022) describe fossil wood specimens from the Miocene Natma Formation (Myanmar), representing an assemblage dominated by members of the families Fabaceae and Dipterocarpaceae, interpreted as coming from different types of low altitude forest ecosystems (tropical wet evergreen, tropical dry and deciduous, and tropical littoral), and interpreted as indicative of a monsoonal climate with an alternance of a dry season and a wet season.[200]
* Abundant compression floras dominated by angiosperm leaves are described from two sites of probable Pliocene age in Brunei by Wilf et al. (2022), who interpret these floras as evidence of dipterocarp-dominated lowland rainforests in the Malay Archipelago before the Pleistocene.[201]
* A study on the impact of the extinct Neotropical megafauna on the variability in plant functional traits and biome geography in Central and South America is published by Dantas & Pausas (2022).[202]
* A study on plant material from rock overhangs from mid-late Holocene sites along the Kawarau-Cromwell-Roxburgh Gorges in Central Otago (New Zealand), much of which was likely transported as roosting material or consumed by moa birds, and on its implications for the knowledge of the mid-late Holocene regional vegetation of Central Otago and the knowledge of vegetation changes since mid-late Holocene, is published by Pole (2022).[203]
* A study on the role of hydraulic failure in the evolution of early vascular plants is published by Bouda et al. (2022), suggesting that drought selection played a key role in the diversification of vascular arrangements beginning with the Devonian explosion.[204]

## References

[edit]

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