Arthropods of the Cambrian Period

Extinct arthropods of the Cambrian period.

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This list contains extinct arthropod genera from the Cambrian Period of the Paleozoic Era.

## Dinocaridida

[edit]

Genus  | Year of Discovery  | Location of Fossils

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Anomalocaris | 1892  | Emu Bay Shale

Maotianshan Shales

Burgess Shale

Cucumericrus | 1995  | Maotianshan Shales
Paranomalocaris | 2013  | Wulongqing Formation
Cambroraster | 2019  | Burgess Shale

Maotianshan Shales

Hurdia | 1992  | Burgess Shale
Peytoia | 1911  | Burgess Shale

Zawiszyn Formation

Ursulinacaris | 2019  | Mount Cap Formation

Carrara Formation

Titanokorys | 2018  | Burgess Shale
Pahvantia | 1981  | Wheeler Shale

Marjum Formation

Amplectobelua | 1995  | Maotianshan Shales

Burgess Shale

Houcaris | 2021  | Maotianshan Shales

Pioche Shale

Pyramid Shale

Lyrarapax | 2014  | Maotianshan Shales
Caryosyntrips | 2010  | Burgess Shale

Wheeler Shale

Marjum Formation

Valdemiedes Formation

Tamisiocaris | 2010  | Sirius Passet
Opabinia | 1912  | Burgess Shale
Utaurora | 2022  | Wheeler Formation
Kerygmachela | 1993  | Sirius Passet
Laminacaris | 2018  | Maotianshan Shales
Pambdelurion | 1997  | Sirius Passet
Mobulavermis | 2023  | Pioche Shale
Echidnacaris | 2023  | Emu Bay Shale
Utahnax | 2022  | Wheeler Shale?
Buccaspinea | 2021  | Marjum Formation
Guanshancaris | 2023  | Guanshan Biota
Omnidens | 2006  | Maotianshan Shales

Xiaoshiba Lagerstatte

Innovatiocaris | 2022  | Maotianshan Shales
Shucaris[1] | 2024  | Maotianshan Shales
Stanleycaris | 2018  | Burgess Shale

Qingjiang biota

## stem-Deuteropoda

[edit]

Genus  | Year of Discovery  | Location of Fossils

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Liangshanella | 1997  | Chengjiang Biota

Burgess Shale

Kunmingella | 1912  | Maotianshan Shales
Comptaluta | 1968  | Maotianshan Shales
Jiucunella | 1991  | Maotianshan Shales
Kunyangella | 1965  | Maotianshan Shales
Fuxianhuia | 1987  | Maotianshan Shales
Alacaris | 2018  | Xiaoshiba Lagerstatte
Chengjiangocaris | 1991  | Maotianshan Shales
Guangweicaris | 2007  | Guanshan Biota
Tauricornicaris | 2017  | Maotianshan Shales
Isoxys | 1890  | Burgess Shale

Chengjiang Biota

Sirius Passet

Surusicaris | 2015  | Burgess Shale
Kylinxia | 2020  | Maotianshan Shales
Erratus | 2022  | Helinpu Formation
Fengzhengia | 2022  | Maotianshan Shales
Xiaocaris | 2020  | Maotianshan Shales
Shankouia | 2005  | Maotianshan Shales
Liangwangshania | 2005  | Maotianshan Shales

## Chelicerata

[edit]

Genus  | Year of Discovery  | Location of Fossils

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|

Sanctacaris | 1988  | Burgess Shale
Habelia | 1912  | Burgess Shale
Mollisonia | 1912  | Maotianshan Shales

Burgess Shale

Langston Formation

Wheeler Shale

Thelxiope | 1975  | Burgess Shale

Wheeler Shale

Fezouata Formation

Linyi Lagerstätte

Cambropycnogon | 2002  | Orsten Fauna

## Hymenocarina

[edit]

Genus  | Year of Discovery  | Location of Fossils

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Branchiocaris | 1929  | Wheeler Shale

Burgess Shale

Tokummia | 2017  | Burgess Shale
Canadaspis | 1978  | Burgess Shale

Pioche Shale

Maotianshan Shales

Waptia | 1912  | Burgess Shale
Pectocaris | 1999  | Maotianshan Shales

Burgess Shale

Clypecaris | 1999  | Maotianshan Shales
Pauloterminus | 2002  | Sirius Passet
Perspicaris | 1977  | Burgess Shale
Nereocaris | 2012  | Burgess Shale
Plenocaris | 1912  | Burgess Shale

Maotianshan Shales

Ercaicunia | 1999  | Maotianshan Shales
Fibulacaris | 2019  | Burgess Shale
Synophalos | 2009  | Maotianshan Shales
Odaraia | 1912  | Burgess Shale

Greater Phyllopod Bed

Protocaris | 1884  | Parker Formation
Hymenocaris | 1853  | Burgess Shale
Tuzoia | 1912  | Burgess Shale

Emu Bay Shale

Rockslide Formation

Spence Shale

Wheeler Formation

Marjum Formation

Kinzers Formation

## Marrellomorpha

[edit]

Genus  | Year of Discovery  | Location of Fossils

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|

Marrella | 1912  | Burgess Shale

Kaili Formation

Balang Formation

Skania | 1931  | Maotianshan Shales

Burgess Shale

Primicaris | 2003  | Maotianshan Shales

## Mandibulata

[edit]

Genus  | Year of Discovery  | Location of Fossils

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Heymonsicambria[2] | 1994  | Orsten Fauna
Skara[3] | 1983  | Orsten Fauna
Goticaris[4] | 1990  | Orsten Fauna
Wujicaris[5] | 2010  | Maotianshan Shales
Cambropachycope[4] | 1990  | Orsten Fauna
Ercaia[6] | 2001  | Maotianshan Shales
Mosineia[7] | 2010  | Blackberry Hill
Cambrocaris[8] | 1991  | Orsten Fauna
Bredocaris[3] | 1983  | Orsten Fauna
Dala[3] | 1983  | Orsten Fauna
Sandtorpia[9] | 2009  | Orsten Fauna
Walossekia[3] | 1983  | Orsten Fauna
Rehbachiella[3] | 1983  | Orsten Fauna
Martinssonia[10] | 1986  | Orsten Fauna
Musacaris[11] | 2010  | Orsten Fauna
Henningsmoenicaris[4] | 1990  | Orsten Fauna
Oelandocaris[3] | 1983  | Orsten Fauna
Aengapentastomum[12] | 2005  | Orsten Fauna
Boeckelericambria[2] | 1994  | Orsten Fauna
Haffnericambria[2] | 1994  | Orsten Fauna
Klausmuelleria[13] | 2003  | Comley Limestone
Mictomerus[7] | 2010  | Potsdam Group
Apankura[14] | 2004  | Santa Rosita Formation
Arenosicaris[7] | 2010  | Blackberry Hill
Yicaris[15] | 2007  | Maotianshan Shales

## Megacheira

[edit]

Genus  | Year of Discovery  | Location of Fossils

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Actaeus | 1970  | Burgess Shale
Alalcomenaeus | 1970  | Burgess Shale

Chengjiang Biota

Wheeler Shale

Leanchoilia | 1912  | Burgess Shale

Chengjiang Biota

Spence Shale

Fortiforceps | 1997  | Maotianshan Shales
Yawunik | 2015  | Burgess Shale
Parapeytoia | 1995  | Maotianshan Shales
Jianfengia | 1987  | Maotianshan Shales
Haikoucaris | 2004  | Chengjiang Biota
Kanoshoia | 2020  | Wheeler Formation
Yohoia | 1912  | Burgess Shale

## Artiopoda

[edit]

Genus  | Year of Discovery  | Location of Fossils

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Bathyuriscus | 1873  | Burgess Shale
Urokodia | 1989  | Maotianshan Shales
Acanthomeridion | 1989  | Chengjiang Biota
Olenoides | 1877  | Burgess Shale

Marjum Formation

Hanburia | 1916  | Burgess Shale

Stephen Formation

Xiuqiella | 1974  |
Acontheus | 1851  |
Hemirhodon | 1937  |
Thoracocare | 1974  | Spence Shale Formation
Abakania | 1973  | Sanashtykgol Formation
Cyclopites | 1951  |
Zacanthoides | 1888  | Burgess Shale
Changaspis | 1961  |
Ogygopsis | 1889  | Burgess Shale
Oryctocephalus | 1886  | Burgess Shale
Chancia | 1924  | Burgess Shale
Ehmaniella | 1951  | Burgess Shale
Elrathia | 1924  | Wheeler Formation

Burgess Shale

Ellipsocephalus | 1883  | Słowiec Sandstone Formation
Phantaspis | 2020  | Mantou Formation
Lisania | 1911  |
Bailiaspis | 1936  | Whitesands Bay Formation

Jbel Lmgaysmat Formation

Manuels River Formation

Maentwrog Formations

Solenoparia | 1935  |
Westergaardites | 1937  |
Yunnanocephalus | 1936  | Maotianshan Shales
Asaphiscus | 1873  |
Olenus | 1827  |
Quitacetra | 1967  |
Ejinaspis | 1986  |
Asthenopsis | 1939  |
Agraulos | 1847  | Menevia Formation

Jince Formation

Manuels River Formation

Coosella | 1936  |
Norwoodia | 1916  |
Tricrepicephalus | 1935  | Chefu Formation

Deadwood Formation

Little Elk Creek

Meniscopsia | 2011  |
Acrocephalella | 1963  |
Cambroproteus | 2015  |
Cedaria | 1924  | Weeks Formation
Conocoryphe | 1847  |
Galeaspis | 1983  |
Genevievella | 1936  | Mungerbar Formation
Meteoraspis | 1890  | Limestone Formation
Validaspis | 1991  |
Palaeolenus | 1912  |
Ptychoparia | 1847  |
Kootenia | 1939  | Burgess Shale

Emu Bay Shale

Misszhouia | 1985  | Maotianshan Shales
Naraoia | 1912  | Burgess Shale

Stephen Formation

Maotianshan Shales

Bertie Group

Balang Formation

Emu Bay Shale

Buenaspis | 1999  | Sirius Passet
Emucaris | 2010  | Emu Bay Shale
Kangacaris | 2010  | Emu Bay Shale

Maotianshan Shales

Liwia | 1988  | Zawiszyn Formation
Estaingia | 1964  |
Bathynotus | 1860  |
Bajanaspis | 1978  | Edreisk Formation
Kleptothule | 1995  | Sirius Passet
Acadoparadoxides | 1957  |
Paradoxides | 1822  | Duda Formation
Eoredlichia | 1950  | Maotianshan Shales
Profallotaspis | 2002  |
Lochmanolenellus | 1952  | Poleta Formation
Olenellus | 1861  | Pioche Shale

Carrara Formation

Kinzers Formation

Archaeaspis | 1965  | Campito Formation
Parapaokannia | 1980  |
Plutonides | 1985  | Purley Shale Formation
Redlichia | 1902  |
Repinaella | 1965  | Pestrotsvet Formation
Ushbaspis | 1965  | Shabakty Formation
Balcoracania | 1970  | Balcoracana Creek
Emuella | 1970  | Emu Bay Shale
Megapharanaspis | 2006  |
Wutingaspis | 1944  |
Mesolenellus | 1993  | Sofiekammen Formation
Olenelloides | 1894  |
Biceratops | 1971  | Pioche Shale
Bradyfallotaspis | 1972  |
Eopeachella | 2009  | Delamar Mountains
Laudonia | 1956  |
Mummaspis | 1992  | Mural Formation
Nephrolenellus | 1993  | Pyramid Shale Member

Carrara Formation

Nevadella | 1936  | Poleta Formation

Sekwi Formation

Paranephrolenellus | 2007  |
Wanneria | 1910  | Kinzers Formation
Bristolia | 1956  | Carrara Formation
Acadoparadoxides | 1957  |
Emigrantia | 1910  |
Bolbolenellus | 1993  |
Buenellus | 1988  | Sirius Passet
Callavia | 1890  | Brigus Formation
Chengjiangaspis | 1980  | Maotianshan Shales
Cirquella | 1993  |
Daguinaspis | 1950  | Amouslek Formation
Fallotaspis | 1953  |
Holmia | 1890  | Sparagmite Formation
Hydrocephalus | 1846  |
Fritzolenellus | 1998  |
Mesonacis | 1859  | Forteau Formation
Xystridura | 1936  |
Peachella | 1910  |
Richterops | 1954  |
Archaeagnostus | 1939  |
Agnostus | 1822  | Marjum Biota

Linyi Lagerstätte

Orsten Fauna

Itagnostus | 1979  |
Peronopsis | 1847  |
Acmarhachis | 1938  |
Acadagnostus | 1939  |
Agnostotes | 1963  |
Lejopyge | 1828  |
Glyptagnostus | 1936  |
Egyngolia | 1980  |
Jinghediscus | 1985  |
Pagetia | 1916  | Burgess Shale
Acimetopus | 1966  |
Pentagnostus | 1940  |
Calodiscus | 1935  | Schodack Formation
Mallagnostus | 1935  |
Runcinodiscus | 1976  |
Serrodiscus | 1941  | Brigus Formation
Weymouthia | 1913  |
Analox | 1966  |
Litometopus | 1966  |
Meniscuchus | 1975  |
Ninadiscus | 1980  | Egyngolskaya Suite
Oodiscus | 1966  |
Semadiscus | 1978  |
Stigmadiscus | 1966  |
Parapagetia | 1964  |
Tannudiscus | 1959  | Purley Shale Formation
Chelediscus | 1966  |
Bathydiscus | 1966  |
Cobboldites | 1943  |
Acidiscus | 1966  |
Buenaspis | 1999  | Sirius Passet
Condylopyge | 1847  |
Delgadella | 1908  | Tyuser Formation
Dicerodiscus | 1954  |
Pleuroctenium | 1847  |
Ptychagnostus | 1909  |
Bolboparia | 1966  |
Tchernyshevioides | 1975  |
Toragnostus | 1988  |
Tsunyidiscus | 1966  |
Birmanites | 1934  |
Kendallina | 1953  | Bison Creek Formation
Dikelocephalus | 1852  |
Orygmaspis | 1938  | Morgan Creek Member
Cindarella | 1996  | Maotianshan Shales
Helmetia | 1912  | Burgess Shale

Jince Formation

Tegopelte | 1975  | Burgess Shale

Kicking Horse Shale

Retifacies | 1989  | Maotianshan Shales
Schmalenseeia | 1903  |
Burlingia | 1908  |
Entomaspis | 1931  |
Neodrepanura | 2006  | Maotianshan Shales
Emeraldella | 1912  | Burgess Shale

Wheeler Shale

Molaria | 1912  | Burgess Shale

Sirius Passet

Sidneyia | 1911  | Burgess Shale

Maotianshan Shales

Bailongia | 2021  | Maotianshan Shales
Luohuilinella | 2012  | Maotianshan Shales
Panlongia | 2006  | Guanshan fauna
Pygmaclypeatus | 2000  | Maotianshan Shales
Sinoburius | 1991  | Maotianshan Shales
Retifacies | 1989  | Maotianshan Shales
Kodymirus | 1965  | Paseky Shale
Glypharthrus | 1939  | McKay Group

Clay Creek

Aaveqaspis | 2009  | Sirius Passet
Thulaspis | 2023  | Sirius Passet
Saperion | 1991  | Maotianshan Shales
Squamacula | 1997  | Maotianshan Shales

Emu Bay Shale

Falcatamacaris | 2014  | Weeks Formation
Zhiwenia | 2019  | Xiaoshiba Lagerstatte
Australimicola | 2012  | Emu Bay Shale
Tonglaiia | 2023  | Maotianshan Shales
Arthroaspis | 2013  | Sirius Passet
Kwanyinaspis | 2005  | Maotianshan Shales
Campanamuta | 2011  | Sirius Passet
Rhombicalvaria | 1987  | Maotianshan Shales
Haifengella | 2014  | Maotianshan Shales
Skioldia | 1997  | Maotianshan Shales
Kuamaia | 1987  | Maotianshan Shales
Cindarella | 1996  | Maotianshan Shales
Xandarella | 1991  | Maotianshan Shales
Zhugeia | 2023  | Maotianshan Shales
Eozetetes | 2016  | Emu Bay Shale

## Incertae sedis

[edit]

Genus  | Year of Discovery  | Location of Fossils

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|

Kiisortoqia | 2010  | Sirius Passet
Carnarvonia | 1912  | Burgess Shale
Vladicaris | 1995  |
Burgessia | 1912  | Burgess Shale
Mongolitubulus | 1997  |
Khankaspis | 1969  | Snegurovka Formation
Strabops | 1901  |
Paleomerus | 1956  |
Parapaleomerus | 1999  | Maotianshan Shales
Worthenella | 1911  | Burgess Shale
Bushizheia | 2020  | Maotianshan Shales
Combinivalvula | 1987  | Maotianshan Shales
Diplopyge | 1999  | Maotianshan Shales
Dongshanocaris | 1999  | Maotianshan Shales
Jianshania | 1999  | Maotianshan Shales
Pseudoiulia | 1998  | Maotianshan Shales
Priscansermarinus | 1981  | Burgess Shale
Kootenichela | 2013  |
Forfexicaris | 1999  | Maotianshan Shales
Kockurus | 1995  |
Occacaris | 1999  | Maotianshan Shales
Sarotrocercus | 1981  | Burgess Shale

## References

[edit]

1. ^ Wu, Yu; Pates, Stephen; Liu, Cong; Zhang, Mingjing; Lin, Weiliang; Ma, Jiaxin; Wu, Yuheng; Chai, Shu; Zhang, Xiangliang; Fu, Dongjing (16 July 2024). "A new radiodont from the lower Cambrian (Series 2 Stage 3) Chengjiang Lagerstätte, South China informs the evolution of feeding structures in radiodonts". Journal of Systematic Palaeontology. 22 (1). Bibcode:2024JSPal..2264887W. doi:10.1080/14772019.2024.2364887. hdl:10871/136118.
2. ^ a b c Walossek, Dieter; Müller, Klaus J. (1994). "Pentastomid parasites from the Lower Palaeozoic of Sweden". Transactions of the Royal Society of Edinburgh: Earth Sciences. 85 (1): 1–37. doi:10.1017/S0263593300006295.
3. ^ a b c d e f Müller, Klaus J. (1983). "Crustacea with preserved soft parts from the Upper Cambrian of Sweden". Lethaia. 16 (2): 93–109. doi:10.1111/j.1502-3931.1983.tb01704.x.
4. ^ a b c Waloszek, Dieter; Müller, Klaus (1990-10-01). "Upper Cambrian stem-lineage crustaceans and their bearing upon the monophyly of Crustacea and the position of Agnostus". Lethaia. 23: 409–427. doi:10.1111/j.1502-3931.1990.tb01373.x.
5. ^ Zhang, Xi-guang; Maas, Andreas; Haug, Joachim T.; Siveter, David J.; Waloszek, Dieter (2010-06-22). "A Eucrustacean Metanauplius from the Lower Cambrian". Current Biology. 20 (12): 1075–1079. doi:10.1016/j.cub.2010.04.026. ISSN 0960-9822. PMID 20493703. S2CID 14341240.
6. ^ Chen, Jun-Yuan; Vannier, jean; Huang, Di-Ying (7 November 2001). "The origin of crustaceans: new evidence from the Early Cambrian of China". Proceedings of the Royal Society of London. Series B: Biological Sciences. 268 (1482): 2181–2187. doi:10.1098/rspb.2001.1779.
7. ^ a b c Collette, Joseph H.; Hagadorn, James W. (July 2010). "Three-dimensionally preserved arthropods from Cambrian Lagerstätten of Quebec and Wisconsin". Journal of Paleontology. 84 (4): 646–667. doi:10.1666/09-075.1.
8. ^ Waloszek, Dieter; Szaniawski, Hubert (October 1991). "Cambrocaris baltica n. gen. n. sp., a possible stem-lineage crustacean from the Upper Cambrian of Poland". Lethaia. 24 (4): 363–378. Bibcode:1991Letha..24..363W. doi:10.1111/j.1502-3931.1991.tb01488.x.
9. ^ Haug, Joachim T.; Maas, Andreas; Waloszek, Dieter (September 2009). "† Henningsmoenicaris scutula , † Sandtorpia vestrogothiensis gen. et sp. nov. and heterochronic events in early crustacean evolution". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 100 (3): 311–350. doi:10.1017/S1755691010008145.
10. ^ Waloszek, Dieter (July 1986). "Martinssonia elongata gen. et sp.n., a crustacean-like euarthropod from the Upper Cambrian 'Orsten' of Sweden". Zoologica Scripta. 15: 73–92. doi:10.1111/j.1463-6409.1986.tb00211.x. S2CID 84246646.
11. ^ Haug, Joachim T.; Waloszek, Dieter; Haug, Carolin; Maas, Andreas (March 2010). "High-level phylogenetic analysis using developmental sequences: The Cambrian †Martinssonia elongata, †Musacaris gerdgeyeri gen. et sp. nov. and their position in early crustacean evolution". Arthropod Structure & Development. 39 (2–3): 154–173. doi:10.1016/j.asd.2010.01.005.
12. ^ Waloszek, Dieter; Repetski, John E.; Maas, Andreas (June 2005). "A new Late Cambrian pentastomid and a review of the relationships of this parasitic group". Transactions of the Royal Society of Edinburgh: Earth Sciences. 96 (2): 163–176. doi:10.1017/S0263593300001280.
13. ^ Siveter, David J.; Williams, Mark; Waloszek, Dieter (20 July 2001). "A Phosphatocopid Crustacean with Appendages from the Lower Cambrian". Science. 293 (5529): 479–481. doi:10.1126/science.1061697.
14. ^ Vaccari, N. E.; Edgecombe, G. D.; Escudero, C. (July 2004). "Cambrian origins and affinities of an enigmatic fossil group of arthropods". Nature. 430 (6999): 554–557. doi:10.1038/nature02705.
15. ^ Zhang, Xi-guang; Siveter, David J.; Waloszek, Dieter; Maas, Andreas (4 October 2007). "An epipodite-bearing crown-group crustacean from the Lower Cambrian". Nature. 449 (7162): 595–598. doi:10.1038/nature06138.

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