Beech marten

Species of carnivore

"House marten" redirects here. For the bird, see Common house martin.

Beech marten

Conservation status

Least Concern (IUCN 3.1)[1]

Scientific classification
Kingdom:  | Animalia
Phylum:  | Chordata
Class:  | Mammalia
Order:  | Carnivora
Suborder:  | Caniformia
Family:  | Mustelidae
Genus:  | Martes
Species:  |

M. foina

Binomial name
Martes foina

(Erxleben, 1777)

Beech marten range

The beech marten (Martes foina), also known as the stone marten, house marten or white breasted marten, is a species of marten native to much of Europe and Central Asia, though it has established a feral population in North America. It is listed as Least Concern on the IUCN Red List on account of its wide distribution, its large population, and its presence in a number of protected areas.[1] It is superficially similar to the European pine marten, but differs from it by its smaller size and habitat preferences. While the pine marten is a forest specialist, the beech marten is a more generalist and adaptable species, occurring in a number of open and forest habitats.[2]

## Evolution

[edit]

Its most likely ancestor is Martes vetus, which also gave rise to the pine marten. The earliest M. vetus fossils were found in deposits dated to the Würm glaciation in Lebanon and Israel. The beech marten likely originated in the Near East or southwestern Asia, and may have arrived in Europe by the Late Pleistocene or the early Holocene. Thus, the beech marten differs from most other European mustelids of the Quaternary, as all other species (save for the European mink) appeared during the Middle Pleistocene. Comparisons between fossil animals and their descendants indicate that the beech marten underwent a decrease in size beginning in the Würm period.[3] Beech martens indigenous to the Aegean Islands represent a relic population with primitive Asiatic affinities.[4]

The skull of the beech marten suggests a higher adaptation than the pine marten toward hypercarnivory, as indicated by its smaller head, shorter snout and its narrower post-orbital constriction and lesser emphasis on cheek teeth. Selective pressures must have acted to increase the beech marten's bite force at the expense of gape. These traits probably acted on male beech martens as a mechanism to avoid both intraspecific competition with females and interspecific competition with the ecologically overlapping pine marten.[2]

### Subspecies

[edit]

As of 2005[update],[5] eleven subspecies are recognised.

Subspecies  | Trinomial authority  | Description  | Range  | Synonyms

|
|
|
|

European beech marten Martes f. foina

(Nominate subspecies)

| Erxleben, 1777  | A small subspecies, with an average-sized skull. In winter, its back varies from light greyish tawny to completely dark brown. The guard hairs are tawny or chestnut brown, while the underfur is very light, pale-grey. The flanks and withers are slightly lighter than the back, and the belly darker. The legs are dark brown and the throat patch pure white. The patch is variable in size and shape.[6] | European Russia, Western Europe (except the Balkan Peninsula) and the Iberian Peninsula | alba (Bechstein, 1801)

domestica (Pinel, 1792)
fagorum (Fatio, 1869)

Balkan beech marten Martes f. bosniaca | Brass, 1911  |  | Balkan Peninsula |
Cretan beech marten Martes f. bunites | Bate, 1906  | A smaller subspecies than foina, with a less defined throat patch, which may be absent in some specimens.[7] | Crete, Skopelos, Naxos, Erimomilos, Karpathos, Samothrake, Seriphos and Kythnos |
Middle Asian beech marten Martes f. intermedia | Severtzov, 1873  | A smaller subspecies than nehringi, with lighter fur. The back is moderately dark greyish-tawny. The flanks are lighter, but of the same tone as the back. The guard hairs are dark-tawny, while the underfur is almost white. The tail is dark brown. The throat patch is very variable, being sometimes completely undefined.[8] | Montane Middle Asia, from Kopet Dag and Bolshoi Balkhas to Tarbagatai (in the Khangai Mountains) and Altai. Outside the former Soviet Union, its range includes northern Iran, Afghanistan, western Pakistan, western Himalayas, Tien Shan, Tibet and northern Mongolia | altaica (Satunin, 1914)

leucolachnaea (Blanford, 1879)

Tibetan beech marten Martes f. kozlovi | Ognev, 1931  |  | Eastern Tibet |
Iberian beech marten Martes f. mediterranea | Barrett-Hamilton, 1898.  | A lighter, less drab coloured form than foina.[9] | Iberian Peninsula |
Rhodes beech marten Martes f. milleri | Festa, 1914  |  | Rhodes |
Caucasian beech marten Martes f. nehringi | Satunin, 1906  | A large subspecies with a massive skull. The winter coat is quite dark, brownish-tawny or dark tawny with a greyish tint. The flanks are lighter than the back, and the tail and feet are dark brown. The throat patch varies in form and size, but shows a tendency towards reduction.[10] | Caucasus and contiguous parts of Turkey and Iran |
Crimean beech marten Martes f. rosanowi | Martino and Martino, 1917  | A smaller subspecies than foina, but with near identical colours.[10] | Montane Crimea |
Syrian beech marten Martes f. syriaca | Nehring, 1902  | A pale coloured subspecies with a smaller skull than the nominative form[11] | Syria |
Lhasa beech marten Martes f. toufoeus | Hodgson, 1842  |  | Lhasa, Tibet |

## Description

[edit]

Skull, as illustrated in Merriam's Synopsis of the weasels of North America Various throat patch variations, as illustrated in Pocock, R. I. The Fauna of British India, including Ceylon and Burma. Vol. Mammalia 2. The elongated body combined with short legs enable the beech marten to move in narrow spaces easily Skull of a beech marten

The beech marten is superficially similar to the pine marten, but has a somewhat longer tail, a more elongated and angular head and has shorter, more rounded and widely spaced ears. Its nose is also of a light peach or grey colour, whereas that of the pine marten is dark black or greyish-black.[12] Its feet are not as densely furred as those of the pine marten, thus making them look less broad, with the paw pads remaining visible even in winter. Because of its shorter limbs, the beech marten's manner of locomotion differs from that of the pine marten; the beech marten moves by creeping in a polecat-like manner, whereas the pine marten and sable move by bounds.[13] The load per 1 cm2 of the supporting surface of the beech marten's foot (30.9 g) is double that of the pine marten (15.2 g), thus it is obliged to avoid snowy regions.[14]

Its skull is similar to that of the pine marten, but differs in its shorter facial region, more convex profile, its larger carnassials and smaller molars.[15] The beech marten's penis is larger than the pine marten's, with the bacula of young beech martens often outsizing those of old pine martens. Males measure 430–590 mm in body length, while females measure 380–470 mm. The tail measures 250–320 mm in males and 230–275 mm in females. Males weigh 1.7–1.8 kg in winter and 2–2.1 kg in summer, while females weigh 1.1–1.3 kg in winter and 1.4–1.5 kg in summer.[16]

The beech marten's fur is coarser than the pine marten's, with elastic guard hairs and less dense underfur. Its summer coat is short, sparse and coarse, and the tail is sparsely furred. The colour tone is lighter than the pine marten's. Unlike the pine marten, its underfur is whitish, rather than greyish. The tail is dark-brown, while the back is darker than that of the pine marten. The throat patch of the beech marten is always white. The patch is large and generally has two projections extending backwards to the base of the forelegs and upward on the legs. The dark colour of the belly juts out between the forelegs as a line into the white colour of the chest and sometimes into the neck. In the pine marten, by contrast, the white colour between the forelegs juts backwards as a protrusion into the belly colour.[13]

## Behaviour and ecology

[edit]

A litter of beech marten kits in а farm outbuilding in the village of Orlintzi, Bulgaria Beech marten fighting a European otter, as illustrated in Brehm's Life of Animals

The beech marten is mainly a crepuscular and nocturnal animal, though to a much lesser extent than the European polecat. It is especially active during moonlit nights. Being a more terrestrial animal than the pine marten, the beech marten is less arboreal in its habits, though it can be a skilled climber in heavily forested areas. It is a skilled swimmer, and may occasionally be active during daytime hours, particularly in the summer, when nights are short. It typically hunts on the ground. During heavy snowfalls, the beech marten moves through paths made by hares or skis.[17]

### Social and territorial behaviours

[edit]

In an area of northeastern Spain, where the beech marten still lives in relatively unmodified habitats, one specimen was recorded to have had a home range of 52.5 ha (130 acres) with two centres of activity. Its period of maximum activity occurred between 6 PM and midnight. Between 9 AM and 6 PM, the animal was found to be largely inactive.[18] In urban areas, beech marten's dens are almost entirely in buildings, particularly during winter.[19] The beech marten does not dig burrows, nor does it occupy those of other animals. Instead, it nests in naturally occurring fissures and clefts in rocks, spaces between stones in rock slides and inhabited or uninhabited stone structures. It may live in tree holes at a height of up to 9 metres.[20]

### Reproduction and development

[edit]

Estrus and copulation occur at the same time as in the pine marten.[21] Copulation can last longer than 1 hour.[22] Mating occurs in the June–July period, and takes place in the morning or in moonlit nights on the ground or on the roofs of houses. The gestation period lasts as long as the pine marten's, lasting 236–237 days in the wild, and 254–275 days in fur farms. Parturition takes place in late March-early April, with the average litter consisting of 3–7 kits. The kits are born blind, and begin to see at the age of 30–36 days. The lactation period lasts 40–45 days. In early July, the young are indistinguishable from the adults.[21]

### Diet

[edit]

The beech marten's diet includes a much higher quantity of plant food than that of the pine marten and sable. Plant foods eaten by the beech marten include cherries, apples, pears, plums, black nightshade, tomatoes, grapes, raspberries and mountain ash. Plant food typically predominates during the winter months. Rats, mice and chickens are also eaten. Among bird species preyed upon by the beech marten, sparrow-like birds predominate, though snowcocks and partridges may also be taken. The marten likes to plunder nests of birds including passerines, galliformes and small owls, preferring to kill the parents in addition to the fledglings. Although it rarely attacks poultry, some specimens may become specialized poultry raiders, even when wild prey is abundant.[14] Males tend to target large, live prey more than females, who feed on small prey and carrion with greater frequency.[2]

### Relationships with other predators

[edit]

In areas where the beech marten is sympatric with the pine marten, the two species avoid competing with one another by assuming different ecological niches; the pine marten feeds on birds and rodents more frequently, while the beech marten feeds on fruits and insects.[23] However, in one known case, a subadult beech marten was killed by a pine marten. The beech marten has been known to kill European polecats on rare occasions. Red foxes, lynxes, mountain lions, golden eagles, and Eurasian eagle-owls may prey on adults, and juveniles are vulnerable to attack by birds of prey and wildcats. There is, however, one case, from Germany, of a beech marten killing a domestic cat,[21][24][25] although there is a report of a domestic cat killing a beech marten after a struggle lasting more than half an hour.[26]

## Range

[edit]

The beech marten is a widespread species which occurs throughout much of Europe and Central Asia. It occurs from Spain and Portugal in the west, through Central and Southern Europe, the Middle East and Central Asia, extending as far east as the Altai and Tien Shan mountains and northwest China. Within Europe, the species is absent in the British Isles, Scandinavian Peninsula, Finland, the northern Baltic and northern European Russia. It occurs in Afghanistan, Pakistan, India, Nepal, Bhutan and was recently confirmed to inhabit northern Burma.[1]

### Introduction in North America

[edit]

The beech marten is present in Wisconsin, particularly near the urban centres surrounding Milwaukee. It is also present in several wooded, upland areas in the Kettle Moraine State Forest, and in nearby woodlands of Walworth, Racine, Waukesha and probably Jefferson Counties. North American beech martens are likely descended from feral animals that escaped a private fur farm in Burlington during the 1940s.[27] They have also been listed as being released or having escaped in 1972.[28]

## Relationships with humans

[edit]

### Tameability

[edit]

British zoologist George Rolleston theorised that the "domestic cat" of the Ancient Greeks and Romans was in fact the beech marten.[29] Pioneering marine biologist Jeanne Villepreux-Power kept two tame beech martens.[30]

### Hunting and fur use

[edit]

Although the beech marten is a valuable animal to the fur trade, its pelt is inferior in quality to that of the pine marten and sable. Beech marten skins on the fur markets of the Soviet Union accounted for only 10–12% of the market presence of pine marten skins. Beech martens were caught only in the Caucasus, in the Montane part of Crimea and (in very small numbers) in the rest of Ukraine, and in the republics of Middle Asia. Because animals with more valuable pelts are rare in those areas, the beech marten is of value to hunters on the local market. Beech martens are captured with jaw traps, or, for live capture, with cage traps. The shooting of beech martens is inefficient, and trailing them with dogs is only successful when the animal can be trapped in a tree hollow.[31]

### Car damage

[edit]

Since the mid-1970s, the beech marten has been known to occasionally cause damage to cars. Cars attacked by martens typically have cut tubes and cables. The reason for this behaviour is not fully known, as the damaged items are not eaten. There is, however, a seasonal peak in marten attacks on cars in spring, when young martens explore their surroundings more often and have yet to learn which items in their habitat are edible or not.[32]

### Large Hadron Collider

[edit]

On 29 April and 21 November 2016, two beech martens shut down the Large Hadron Collider, the world's most powerful particle accelerator, by climbing on 18–66 kV electrical transformers located above ground near the LHCb and ALICE experiments, respectively.[33][34][35] The second marten was stuffed and put on display in the Rotterdam Natural History Museum.[36]

## References

[edit]

1. ^ a b c Abramov, A.V.; Kranz, A.; Herrero, J.; Choudhury, A.; Maran, T. (2016). "Martes foina". IUCN Red List of Threatened Species. 2016: e.T29672A45202514. doi:10.2305/IUCN.UK.2016-1.RLTS.T29672A45202514.en. Retrieved 19 November 2021.
2. ^ a b c Loy, A.; Spinosi, O. & Carlini, R. (2004). "Cranial morphology of Martes foina and M. martes (Mammalia, Carnivora, Mustelidae): The role of size and shape in sexual dimorphism and interspecific differentiation". Italian Journal of Zoology. 71 (1): 27–34\. doi:10.1080/11250000409356547. S2CID 83563512.
3. ^ Spagnesi & De Marina Marinis 2002, p. 238
4. ^ Schreiber, A. (1999). "On the status of Martes foina bunites Bate, 1905" (PDF). Small Carnivore Conservation. 20: 20–21\. Archived from the original (PDF) on 2011-07-28. Retrieved 2011-07-09.
5. ^ Wozencraft, W. C. (2005). "Order Carnivora". In Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. ISBN 978-0-8018-8221-0. OCLC 62265494.
6. ^ Heptner & Sludskii 2002, pp. 892
7. ^ Miller 1912, pp. 381
8. ^ Heptner & Sludskii 2002, pp. 894
9. ^ Miller 1912, pp. 380
10. ^ a b Heptner & Sludskii 2002, pp. 893
11. ^ Harrison, D.L. & Bates, P.J.J. (1991). The Mammals of Arabia. Sevenoaks, Kent: Harrison Zoological Museum. ISBN 9780951731307.
12. ^ Heptner & Sludskii 2002, p. 876
13. ^ a b Heptner & Sludskii 2002, p. 877
14. ^ a b Heptner & Sludskii 2002, pp. 896–899
15. ^ Heptner & Sludskii 2002, p. 879
16. ^ Heptner & Sludskii 2002, p. 881
17. ^ Heptner & Sludskii 2002, pp. 900–902
18. ^ López-Martín, J.M.; Ruiz-Olmo, J. & Cahill, S. (1992). "Autumn home range and activity of a Stone Marten (Martes foina Erxleben, 1777) in Northeastern Spain". Miscellània Zoològica. 16: 258–260.
19. ^ Herra, J.; Schley, L.; Engel, E. & Roper, T. J. (2010). "Den preferences and denning behaviour in urban stone martens (Martes foina)". Mammalian Biology - Zeitschrift für Säugetierkunde. 75 (2): 138–145\. Bibcode:2010MamBi..75..138H. doi:10.1016/j.mambio.2008.12.002.
20. ^ Heptner & Sludskii 2002, pp. 899–900
21. ^ a b c Heptner & Sludskii 2002, pp. 902
22. ^ Lodé, T. (1991). "Conspecific recognition and mating in stone marten Martes foina" (PDF). Acta Theriologica. 36 (3–4): 275–283\. doi:10.4098/AT.arch.91-28.3–4&rft.pages=275-283&rft.date=1991&rft_id=info:doi/10.4098/AT.arch.91-28&rft.au=Lodé,+T.&rft_id=http://rcin.org.pl/Content/11763/BI002_2613_Cz-40-2_Acta-T36-nr24-275-283_o.pdf&rfr_id=info:sid/en.wikipedia.org:Beech+marten" class="Z3988">
23. ^ Posłuszny, M.; Pilot, M.; Goszczyński, J. & Gralak, B. (2007). "Diet of sympatric Pine Marten (Martes martes) and Stone Marten (Martes foina) identified by genotyping of DNA from faeces" (PDF). Annales Zoologici Fennici. 44 (4): 269–284\. JSTOR 23736771.
24. ^ Mañosa, Santi, et al. "Diet of the Golden Eagle Aquila chrysaetos in two areas on the southern slopes of the Catalan Pyrenees." (2021).
25. ^ Hadad, Ezra, et al. "Prey-Base Does Not Influence Breeding Success in Eagle Owls (Bubo bubo) in Judea, Israel." Animals 12.10 (2022): 1280.
26. ^ Leyhausen, Paul (1979). Cat behavior: the predatory and social behavior of domestic and wild cats. Garland series in ethology. New York: Garland. p. 76\. ISBN 978-0-8240-7017-5.
27. ^ Long, C.A. (1995). "Stone marten (Martes foina) in southeast Wisconsin, U.S.A." (PDF). Small Carnivore Conservation. 13: 14. Archived from the original (PDF) on 2012-03-16. Retrieved 2011-07-09.
28. ^ Bill Frederickson (3 July 2007). "Martes foina: Current status and distribution" (PDF). Wisconsin Department of Natural Resources.
29. ^ Hamilton, Edward (1896) The Wild Cat of Europe, pp. 80-81, London, HR Porter
30. ^ "The 1830s seamstress who solved Aristotle's octopus mystery".
31. ^ Heptner & Sludskii 2002, pp. 904
32. ^ Lachat, N. (1991). "Stone martens and cars: a beginning war?" (PDF). Small Carnivore Conservation. 5: 4–6\. Archived from the original (PDF) on 2012-03-16. Retrieved 2011-07-09.
33. ^ Sample, I. (29 April 2016). "Large Hadron Collider On Paws After Creature Chews Through Wiring", The Guardian. Retrieved 30 April 2016.
34. ^ Bertolasi, Stefano and Lamont, Mike for the LHC team (May 2016). "LHC report: Stoat-ally back on track!" CERN Bulletin Issue No. 20-21/2016
35. ^ "LHC Morning Meeting" (PDF). 21 November 2016.
36. ^ Sample, Ian (2017-01-27). "Totally stuffed: Cern's electrocuted weasel to go on display". The Guardian.

Wikispecies has information related to Martes foina.

### Bibliography

[edit]

Wikimedia Commons has media related to Martes foina.

* Brehm, A. E. (1895). Brehm's Life of Animals. Chicago: A. N. Marquis & Company.
* Heptner, V. G.; Sludskii, A. A. (2002). Mammals of the Soviet Union. Vol. II, part 1b, Carnivores (Mustelidae and Procyonidae). Washington, D.C. : Smithsonian Institution Libraries and National Science Foundation. ISBN 978-90-04-08876-4.
* Miller, Gerrit Smith (1912). Catalogue of the mammals of Western Europe (Europe exclusive of Russia) in the collection of the British museum. London : printed by order of the Trustees.`{{cite book}}`: CS1 maint: publisher location (link)
* Pocock, R. I. (1941). The Fauna of British India, Including Ceylon and Burma. Vol. Mammals Volume 2. Taylor and Francis.
* Spagnesi, M.; De Marina Marinis, M. (2002). "Mammiferi d'Italia" (PDF). Quaderni di Conservazione della Natura. ISSN 1592-2901.[permanent dead link]

* v
* t
* e

Extant Carnivora species

* Kingdom: Animalia
* Phylum: Chordata
* Class: Mammalia
* Infraclass: Eutheria
* Superorder: Laurasiatheria

Feliformia ("cat-like" carnivorans)

Feliformia

* African palm civet (Nandinia binotata)

Feloidea

| Prionodon (Asiatic linsangs)|

* Banded linsang (P. linsang)
* Spotted linsang (P. pardicolor)

|

Felidae (cats)

| Pantherinae|

| Neofelis|

* Sunda clouded leopard (N. diardi)
* Clouded leopard (N. nebulosa)

|

Panthera|

* Snow leopard (P. uncia)
* Tiger (P. tigris)
* Jaguar (P. onca)
* Lion (P. leo)
* Leopard (P. pardus)

Felinae sensu stricto

| Bay cat
lineage|

| Pardofelis|

* Marbled cat (P. marmorata)

|

Catopuma|

* Bay cat (C. badia)
* Asian golden cat (C. temminckii)

Caracal
lineage|

* Serval (Leptailurus serval)

Caracal|

* African golden cat (C. aurata)
* Caracal (C. caracal)

Leopardus|

* Pampas cat (L. colocola)
* Geoffroy's cat (L. geoffroyi)
* Kodkod (L. guigna)
* Southern tiger cat (L. guttulus)
* Andean mountain cat (L. jacobita)
* Ocelot (L. pardalis)
* Oncilla (L. tigrinus)
* Margay (L. wiedii)

Lynx|

* Canada lynx (L. canadensis)
* Eurasian lynx (L. lynx)
* Iberian lynx (L. pardinus)
* Bobcat (L. rufus)

Puma
lineage|

| Acinonyx|

* Cheetah (A. jubatus)

|

* Jaguarundi (Herpailurus yagouaroundi)

Puma|

* Cougar (P. concolor)

Leopard cat
lineage|

* Pallas's cat (Otocolobus manul)

Prionailurus|

* Leopard cat (P. bengalensis)
* Sunda leopard cat (P. javanensis)
* Flat-headed cat (P. planiceps)
* Rusty-spotted cat (P. rubiginosus)
* Fishing cat (P. viverrinus)

Felis|

* Chinese mountain cat (F. bieti)
* Domestic cat (F. catus)
* Jungle cat (F. chaus)
* African wildcat (F. lybica)
* Sand cat (F. margarita)
* Black-footed cat (F. nigripes)
* European wildcat (F. silvestris)

Viverroidea|

*     * see below↓

Viverroidea

Viverridae

Palm civets

| Hemigalinae|

* Owston's palm civet (Chrotogale owstoni)
* Otter civet (Cynogale bennettii)
* Hose's palm civet (Diplogale hosei)
* Banded palm civet (Hemigalus derbyanus)

|

Paradoxurinae|

* Binturong (Arctictis binturong)
* Small-toothed palm civet (Arctogalidia trivirgata)
* Sulawesi palm civet (Macrogalidia musschenbroekii)
* Masked palm civet (Paguma larvata)

Paradoxurus|

* Asian palm civet (P. hermaphroditus)
* Brown palm civet (P. jerdoni)
* Golden palm civet (P. zeylonensis)

Viverrinae sensu lato

| Viverrinae
sensu stricto|

* Small Indian civet (Viverricula indica)
* African civet (Civettictis civetta)

Viverra|

* Malabar large-spotted civet (V. civettina)
* Large-spotted civet (V. megaspila)
* Malayan civet (V. tangalunga)
* Large Indian civet (V. zibetha)

Genettinae

| Poiana
(African linsangs)|

* Central African oyan (P. richardsonii)
* West African oyan (P. leightoni)

|

Genetta (genets)

| subgenus Genetta
(paraphyletic)|

* Abyssinian genet (G. abyssinica)
* Common genet (G. genetta)
* South African small-spotted genet (G. felina)

|

subgenus Eugenetta
(paraphyletic)|

* Angolan genet (G. angolensis)
* Cape genet (G. tigrina)

subgenus Herpailuropoda
(paraphyletic)|

* Bourlon's genet (G. bourloni)
* Johnston's genet (G. johnstoni)
* Hausa genet (G. thierryi)
* Giant forest genet (G. victoriae)

subgenus Pardogale
(paraphyletic)|

* Pardine genet (G. pardina)
* King genet (G. poensis)

subgenus Prionailuropoda|

* Letaba genet (G. letabae)
* Rusty-spotted genet (G. maculata)
* Schouteden's genet (G. schoutedeni)

subgenus Leptailuropoda
(paraphyletic)|

* Crested servaline genet (G. cristata)
* Servaline genet (G. servalina)

subgenus Osbornictis|

* Aquatic genet (G. piscivora)

| Herpestoidea|

*     * see below↓

|

Herpestoidea

| Hyaenidae
(hyenas)|

| Proteles|

* Aardwolf (P. cristata)

|

Hyaeninae
(bone-crushing hyenas)|

* Striped hyena (Hyaena hyaena)
* Brown hyena (Parahyaena brunnea)

Crocuta|

* Spotted hyena (C. crocuta)

Herpestidae sensu lato

| Eupleridae
(Malagasy
carnivorans)|

| Euplerinae
(Malagasy civets)|

* Fossa (Cryptoprocta ferox)
* Malagasy civet (Fossa fossana)

Eupleres (falanoucs)|

* Eastern falanouc (E. goudotii)
* Western falanouc (E. major)

Galidiinae
(vontsira)|

* Ring-tailed vontsira (Galidia elegans)

Galidictis|

* Broad-striped Malagasy mongoose (G. fasciata)
* Grandidier's mongoose (G. grandidieri)

* Narrow-striped mongoose (Mungotictis decemlineata)

Salanoia|

* Brown-tailed mongoose (S. concolor)
* Durrell's vontsira (S. durrelli)

Herpestidae sensu stricto (mongooses)

Mungotinae

| Suricata|

* Meerkat (S. suricatta)

|

* Liberian mongoose (Liberiictus kuhni)

Mungos|

* Gambian mongoose (M. gambianus)
* Banded mongoose (M. mungo)

* Pousargues's mongoose (Dologale dybowskii)

Helogale|

* Ethiopian dwarf mongoose (H. hirtula)
* Common dwarf mongoose (H. parvula)

Crossarchus
(kusimanses)|

* Alexander's kusimanse (C. alexandri)
* Angolan kusimanse (C. ansorgei)
* Common kusimanse (C. obscurus)
* Flat-headed kusimanse (C. platycephalus)

Herpestinae

* Marsh mongoose (Atilax paludinosus)
* Long-nosed mongoose (Xenogale naso)

Urva
(Asian mongooses)|

* Small Indian mongoose (U. auropunctata)
* Short-tailed mongoose (U. brachyura)
* Indian grey mongoose (U. edwardsii)
* Indian brown mongoose (U. fusca)
* Javan mongoose (U. javanica)
* Collared mongoose (U. semitorquata)
* Ruddy mongoose (U. smithii)
* Crab-eating mongoose (U. urva)
* Stripe-necked mongoose (U. vitticolla)

* White-tailed mongoose (Ichneumia albicauda)
* Yellow mongoose (Cynictis penicillata)
* Selous's mongoose (Paracynictis selousi)
* Meller's mongoose (Rhynchogale melleri)

Bdeogale|

* Bushy-tailed mongoose (B. crassicauda)
* Jackson's mongoose (B. jacksoni)
* Black-footed mongoose (B. nigripes)

Herpestes
(slender mongooses)|

* Angolan slender mongoose (H. flavescens)
* Egyptian mongoose (H. ichneumon)
* Somalian slender mongoose (H. ochracea)
* Cape gray mongoose (H. pulverulenta)
* Common slender mongoose (H. sanguinea)

Caniformia ("dog-like" carnivorans)

Canidae (dogs)

| Urocyon|

* Gray fox (U. cinereoargenteus)
* Island fox (U. littoralis)

|

Vulpini

* Bat-eared fox (Otocyon megalotis)

Nyctereutes
(raccoon dogs)|

* Common raccoon dog (N. procyonoides)
* Japanese raccoon dog (N. viverrinus)

Vulpes
(true foxes)|

* Bengal fox (V. bengalensis)
* Blanford's fox (V. cana)
* Cape fox (V. chama)
* Corsac fox (V. corsac)
* Tibetan fox (V. ferrilata)
* Arctic fox (V. lagopus)
* Kit fox (V. macrotis)
* Pale fox (V. pallida)
* Rüppell's fox (V. rueppelli)
* Swift fox (V. velox)
* Red fox (V. vulpes)
* Fennec fox (V. zerda)

Canini (true dogs)

Cerdocyonina
(zorro)

* Maned wolf (Chrysocyon brachyurus)

Speothos|

* Bush dog (S. venaticus)

* Short-eared dog (Atelocynus microtis)
* Crab-eating fox (Cerdocyon thous)

Lycalopex
(South American foxes)|

* Culpeo (L. culpaeus)
* Darwin's fox (L. fulvipes)
* South American gray fox (L. griseus)
* Pampas fox (L. gymnocercus)
* Sechuran fox (L. sechurae)
* Hoary fox (L. vetulus)

Canina
(wolf-like canids)

| Lupulella|

* Side-striped jackal (L. adustus)
* Black-backed jackal (L. mesomelas)

|

Lycaon|

* African wild dog (L. pictus)

* Dhole (Cuon alpinus)

Canis|

* Golden jackal (C. aureus)
* Domestic dog (C. familiaris)
* Coyote (C. latrans)
* African wolf (C. lupaster)
* Wolf (C. lupus)
* Eastern wolf (C. lycaon)
* Red wolf (C. rufus)
* Ethiopian wolf (C. simensis)

Arctoidea

Ursidae (bears)

| Ailuropoda|

* Giant panda (A. melanoleuca)

|

Tremarctos|

* Spectacled bear (T. ornatus)

Ursinae|

* Sloth bear (Melursus ursinus)
* Sun bear (Helarctos malayanus)

Ursus|

* American black bear (U. americanus)
* Asian black bear (U. thibetanus)
* Brown bear (U. arctos)
* Polar bear (U. maritimus)

| Mustelida|

| Pinnipedia (seals)|

*     * see below↓

|

Musteloidea|

*     * see below↓

Pinnipedia (seals)

| Odobenidae|

* Walrus (Odobenus rosmarus)

|

Otariidae (eared seals)

| Callorhinus
(northern fur seals)|

* Northern fur seal (C. ursinus)

|

Otariinae
(sea lions)|

* Steller sea lion (Eumetopias jubatus)

Zalophus|

* California sea lion (Z. californianus)
* Galápagos sea lion (Z. wollebaeki)

* South American sea lion (Otaria flavescens)

Neophoca|

* Australian sea lion (N. cinerea)

* New Zealand sea lion (Phocarctos hookeri)

Arctocephalus
(southern fur seals)|

* South American fur seal (A. australis)
* Australasian fur seal (A. forsteri)
* Galápagos fur seal (A. galapagoensis)
* Antarctic fur seal (A. gazella)
* Juan Fernández fur seal (A. philippii)
* Brown fur seal (A. pusillus)
* Guadalupe fur seal (A. townsendi)
* Subantarctic fur seal (A. tropicalis)

Phocidae (earless seals or true seals)

Phocinae
("northern seals")

* Bearded seal (Erignathus barbatus)
* Hooded seal (Cystophora cristata)

Phocini

* Harp seal (Pagophilus groenlandicus)
* Ribbon seal (Histriophoca fasciata)
* Grey seal (Halichoerus grypus)

Phoca|

* Spotted seal (P. largha)
* Harbor seal (P. vitulina)

Pusa|

* Caspian seal (P. caspica)
* Ringed seal (P. hispida)
* Baikal seal (P. sibirica)

Monachinae
("southern seals")

| Monachini
(monk seals)|

* Mediterranean monk seal (Monachus monachus)

Neomonachus|

* Hawaiian monk seal (N. schauinslandi)

Mirounga
(elephant seals)|

* Northern elephant seal (M. angustirostris)
* Southern elephant seal (M. leonina)

Lobodontini
(Antarctic seals)|

* Leopard seal (Hydrurga leptonyx)
* Weddell seal (Leptonychotes weddellii)
* Crabeater seal (Lobodon carcinophagus)
* Ross seal (Ommatophoca rossi)

Musteloidea

| Ailuridae|

* Red panda (Ailurus fulgens)

|

Mephitidae (skunks)

| Conepatus
(hog-nosed skunks)|

* Molina's hog-nosed skunk (C. chinga)
* Humboldt's hog-nosed skunk (C. humboldtii)
* American hog-nosed skunk (C. leuconotus)
* Striped hog-nosed skunk (C. semistriatus)

|

Mephitis|

* Hooded skunk (M. macroura)
* Striped skunk (M. mephitis)

Mydaus
(stink badgers)|

* Sunda stink badger (M. javanensis)
* Palawan stink badger (M. marchei)

Spilogale
(spotted skunks)|

* Southern spotted skunk (S. angustifrons)
* Western spotted skunk (S. gracilis)
* Eastern spotted skunk (S. putorius)
* Pygmy spotted skunk (S. pygmaea)

Procyonidae

* Kinkajou (Potos flavus)

Bassariscus|

* Ringtail (B. astutus)
* Cacomistle (B. sumichrasti)

Procyon
(raccoons)|

* Crab-eating raccoon (P. cancrivorus)
* Raccoon (P. lotor)
* Cozumel raccoon (P. pygmaeus)

Bassaricyon
(olingos)|

* Eastern lowland olingo (B. alleni)
* Northern olingo (B. gabbii)
* Western lowland olingo (B. medius)
* Olinguito (B. neblina)

Nasuina
(coatis)|

| Nasua|

* White-nosed coati (N. narica)
* South American coati (N. nasua)

|

Nasuella
(mountain coatis)|

* Eastern mountain coati (N. meridensis)
* Western mountain coati (N. olivacea)

| Mustelidae|

*     * see below↓

|

Mustelidae

Mustelidae

* American badger (Taxidea taxus)

Mellivora|

* Honey badger (M. capensis)

Melinae

| Arctonyx
(hog badgers)|

* Northern hog badger (A. albogularis)
* Greater hog badger (A. collaris)
* Sumatran hog badger (A. hoevenii)

|

Meles
(Eurasian badgers)|

* Japanese badger (M. anakuma)
* Caucasian badger (M. canescens)
* Asian badger (M. leucurus)
* European badger (M. meles)

Melogale
(ferret-badgers)|

* Vietnam ferret-badger (M. cucphuongensis)
* Bornean ferret badger (M. everetti)
* Chinese ferret-badger (M. moschata)
* Javan ferret-badger (M. orientalis)
* Burmese ferret-badger (M. personata)
* Formosan ferret-badger (M. subaurantiaca)

Guloninae

* Tayra (Eira barbara)

Pekania|

* Fisher (P. pennanti)

Gulo|

* Wolverine (G. gulo)

Martes
(martens)|

* American marten (M. americana)
* Pacific marten (M. caurina)
* Yellow-throated marten (M. flavigula)
* Beech marten (M. foina)
* Nilgiri marten (M. gwatkinsii)
* European pine marten (M. martes)
* Japanese marten (M. melampus)
* Sable (M. zibellina)

Ictonychinae

| Lyncodontini|

* Patagonian weasel (Lyncodon patagonicus)

Galictis
(grisons)|

* Lesser grison (G. cuja)
* Greater grison (G. vittata)

Ictonychini
(African polecats)|

| Vormela|

* Marbled polecat (V. peregusna)

|

* African striped weasel (Poecilogale albinucha)

Ictonyx|

* Saharan striped polecat (I. libycus)
* Striped polecat (I. striatus)

Lutrinae (otters)

* Giant otter (Pteronura brasiliensis)

Lontra|

* North American river otter (L. canadensis)
* Marine otter (L. felina)
* Neotropical otter (L. longicaudis)
* Southern river otter (L. provocax)

Enhydra|

* Sea otter (E. lutris)

* Spotted-necked otter (Hydrictis maculicollis)

Lutra|

* Eurasian otter (L. lutra)
* Hairy-nosed otter (L. sumatrana)

Lutrogale|

* Smooth-coated otter (L. perspicillata)

Aonyx|

* African clawless otter (A. capensis)
* Asian small-clawed otter (A. cinereus)
* Congo clawless otter (A. congicus)

Mustelinae

| Neogale
(New World weasels)|

* Amazon weasel (N. africana)
* Colombian weasel (N. felipei)
* Long-tailed weasel (N. frenata)
* American mink (N. vison)

|

Mustela
(weasels)

| subgenus Mustela
(paraphyletic)|

* Sichuan weasel (M. aistoodonnivalis)
* Mountain weasel (M. altaica)
* Stoat/Beringian ermine (M. erminea)
* Haida ermine (M. haidarum)
* Yellow-bellied weasel (M. kathiah)
* Least weasel (M. nivalis)
* American ermine (M. richardsonii)

|

subgenus Lutreola
(paraphyletic)|

* Japanese weasel (M. itatsi)
* European mink (M. lutreola)
* Indonesian mountain weasel (M. lutreolina)
* Malayan weasel (M. nudipes)
* Siberian weasel (M. sibirica)
* Back-striped weasel (M. strigidorsa)

subgenus Putorius|

* Steppe polecat (M. eversmannii)
* Ferret (M. furo)
* Black-footed ferret (M. nigripes)
* European polecat (M. putorius)

* See also: Mink
* Polecat

Taxon identifiers

Martes foina|

* Wikidata: Q27366
* Wikispecies: Martes foina
* ADW: Martes_foina
* BioLib: 1749
* BOLD: 210835
* CoL: 3Y9VW
* EoL: 311514
* EPPO: MRTSFO
* Fauna Europaea: 305307
* Fauna Europaea (new): affc380a-0232-4dbb-936e-b0c0d20a2844
* GBIF: 5218887
* iNaturalist: 41800
* IRMNG: 11057184
* ITIS: 621941
* IUCN: 29672
* MDD: 1005820
* MSW: 14001228
* NBN: NHMSYS0000376874
* NCBI: 9659
* Observation.org: 421
* Open Tree of Life: 348047
* Paleobiology Database: 157448
* SeaLifeBase: 71092
* TSA: 10023
* Xeno-canto: Martes-foina
* ZooBank: 53503791-49F9-44DB-9F5D-4BEBF4FC8F4D

Mustela foina|

* Wikidata: Q109646715
* CoL: 44QWN
* EUNIS: 11615
* GBIF: 8207313
* ITIS: 1086077

Authority control databases

National|

* Germany
* United States
* France
* BnF data
* Czech Republic
* Israel

Other|

* Yale LUX

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