Columella (botany)

Axis of sterile tissue

Not to be confused with Columellia.

Zygote fungus sporangium, with columella labelled

Columella (in plants) is an axis of sterile tissue which passes through the center of the spore-case of mosses.[1] In fungi, it refers to a centrally vacuolated part of a hypha, bearing spores. The word finds analogous usage in myxomycetes.[citation needed]

The term columella is also used to refer to story 1 to story 4 (S1 – S4) cells in the root cap, located apically of the quiescent centre.[citation needed] In vascular plants, columella tissue helps the roots to perceive gravity. Within columella cells, the amyloplasts and Golgi apparatus organelles tend to fall to the bottom third of the cell.[2]

## References

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1. ^ One or more of the preceding sentences incorporates text from a publication now in the public domain: Beach, Chandler B., ed. (1914). "Columella" . The New Student's Reference Work . Chicago: F. E. Compton and Co.
2. ^ Stoker, R.; Moore, R. (1984). "Structure of columella cells in primary and lateral roots of Helianthus annuus (Compositae)". New Phytologist. 97 (2): 205–212\. doi:10.1111/j.1469-8137.1984.tb04123.x. Retrieved 24 June 2024.

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