Copiula

Genus of amphibians

Copiula

Scientific classification
Domain:  | Eukaryota
Kingdom:  | Animalia
Phylum:  | Chordata
Class:  | Amphibia
Order:  | Anura
Family:  | Microhylidae
Subfamily:  | Asterophryinae
Genus:  | Copiula
Méhelÿ, 1901
Type species
Phrynixalus oxyrhinus

Boulenger, 1898

Species

14 species (see text)

Copiula is a genus of microhylid frogs endemic to New Guinea. The common name Mehely frogs has been coined for them.[1] They are leaf-litter inhabitants.[2]

## Taxonomy

[edit]

Copiula is probably not monophyletic. Some former Austrochaperina species have already been transferred to this genus, and further ones might follow when more data became available.[1][3]

## Species

[edit]

There are at present 14 species in this genus:[1]

* Copiula alpestris (Zweifel, 2000)
* Copiula annanoreenae Günther, Richards, and Dahl, 2014
* Copiula derongo (Zweifel, 2000)
* Copiula exspectata Günther, 2002
* Copiula fistulans Menzies and Tyler, 1977 — Lae Mehely Frog
* Copiula guttata (Zweifel, 2000)
* Copiula lennarti Günther, Richards, and Dahl, 2014
* Copiula major Günther, 2002
* Copiula minor Menzies and Tyler, 1977 — Milne Bay Mehely Frog
* Copiula obsti Günther, 2002
* Copiula oxyrhina (Boulenger, 1898) — Misima Island Mehely Frog
* Copiula pipiens Burton and Stocks, 1986 — Wirui Mehely Frog
* Copiula rivularis (Zweifel, 2000)
* Copiula tyleri Burton, 1990 — Sepik Mehely Frog

The AmphibiaWeb[4] reports fewer species, with species that Peloso and colleagues moved in 2016 from Austrochaperina and Oxydactyla missing.[3]

## References

[edit]

1. ^ a b c Frost, Darrel R. (2016). "Copiula Méhely, 1901". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 20 July 2016.
2. ^ Zweifel, R. G. (2000). "Partition of the Australopapuan microhylid frog genus Sphenophryne with descriptions of new species". Bulletin of the American Museum of Natural History. 253: 1–130\. doi:10.1206/0003-0090(2000)253<0001:POTAMF>2.0.CO;2. hdl:2246/1600. S2CID 85621508.1-130&rft.date=2000&rft_id=info:hdl/2246/1600&rft_id=https://api.semanticscholar.org/CorpusID:85621508#id-name=S2CID&rft_id=info:doi/10.1206/0003-0090(2000)253<0001:POTAMF>2.0.CO;2&rft.aulast=Zweifel&rft.aufirst=R.+G.&rft_id=http://digitallibrary.amnh.org/handle/2246/1600&rfr_id=info:sid/en.wikipedia.org:Copiula" class="Z3988">
3. ^ a b Peloso, Pedro L.V.; Frost, Darrel R.; Richards, Stephen J.; Rodrigues, Miguel T.; Donnellan, Stephen; Matsui, Masafumi; Raxworthy, Cristopher J.; Biju, S.D.; Lemmon, Emily Moriarty; Lemmon, Alan R.; Wheeler, Ward C. (2016). "The impact of anchored phylogenomics and taxon sampling on phylogenetic inference in narrow-mouthed frogs (Anura, Microhylidae)". Cladistics. 32 (2): 113–140\. doi:10.1111/cla.12118. PMID 34732021. S2CID 84925667.113-140&rft.date=2016&rft_id=https://api.semanticscholar.org/CorpusID:84925667#id-name=S2CID&rft_id=info:pmid/34732021&rft_id=info:doi/10.1111/cla.12118&rft.aulast=Peloso&rft.aufirst=Pedro+L.V.&rft.au=Frost,+Darrel+R.&rft.au=Richards,+Stephen+J.&rft.au=Rodrigues,+Miguel+T.&rft.au=Donnellan,+Stephen&rft.au=Matsui,+Masafumi&rft.au=Raxworthy,+Cristopher+J.&rft.au=Biju,+S.D.&rft.au=Lemmon,+Emily+Moriarty&rft.au=Lemmon,+Alan+R.&rft.au=Wheeler,+Ward+C.&rft_id=https://doi.org/10.1111%2Fcla.12118&rfr_id=info:sid/en.wikipedia.org:Copiula" class="Z3988">
4. ^ "Microhylidae". AmphibiaWeb: Information on amphibian biology and conservation. [web application]. Berkeley, California: AmphibiaWeb. 2016. Retrieved 20 July 2016.

Taxon identifiers

Copiula|

* Wikidata: Q2566288
* Wikispecies: Copiula
* ADW: Copiula
* ASW: Copiula
* CoL: 3TQC
* EoL: 42960
* GBIF: 2421966
* iNaturalist: 24933
* IRMNG: 1314203
* ITIS: 661500
* NCBI: 83968
* Open Tree of Life: 442618

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