Cucujoidea

Superfamily of beetles

Cucujoidea

Temporal range: Jurassic–Recent

PreꞒ

Various Cucujoidea with larvae
and anatomical details
Scientific classification
Kingdom:  | Animalia
Phylum:  | Arthropoda
Class:  | Insecta
Order:  | Coleoptera
Suborder:  | Polyphaga
Infraorder:  | Cucujiformia
Superfamily:  | Cucujoidea
Latreille, 1802

Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea.[1] They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark beetles (Scolytinae), which are weevils (superfamily Curculionoidea).

## Morphology

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The morphology of Cucujoidea is varied and there are no features uniting all members of the superfamily. In terms of general appearance, they tend to be small, drab in colour and with clubbed antennae.[1] Even this is not universal; for example, Glischrochilus (Nitidulidae) have aposematic orange spots on their elytra.[2]

Adults can be recognised by the procoxal cavities being internally open in most taxa, females having tarsal formula 5-5-5 and males 5-5-5 or 5-5-4 (rarely 4-4-4), females with tergite VIII concealed dorsally by tergite VII, and males with tergite X completely membraneous.[1]

Larvae have frontal arms usually lyriform, the mandible mesal surface usually with well-developed mola, a maxillary articulating area usually present, a hypopharyngeal sclerome usually present, and two pretarsal setae.[1]

## Ecology

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Cucujoidea usually have cryptic habits, living in fungi, leaf litter or dead wood.[1] This is reflected in many families having "fungus" or "bark" in their common names. The Kateretidae[3] and some Phalacridae[4] feed on flowers instead. The Nitidulidae are quite varied: some are saprophagous and mycetophagous like typical cucujoids, but others are associated with carrion, flowers, insect nests or stored food products.[5]

## Taxonomy

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According to a 2015 revision, the following 25 families make up superfamily Cucujoidea:[1]

* Agapythidae Sen Gupta and Crowson, 1969
* Boganiidae Sen Gupta & Crowson, 1966
* Cavognathidae Sen Gupta & Crowson, 1966
* Cryptophagidae Kirby, 1826 - silken fungus beetles
* Cucujidae Latreille, 1802 - flat bark beetles
* Cybocephalidae Jacquelin du Val, 1858
* Cyclaxyridae Gimmel, Leschen & Ślipiński, 2009 - sooty mould beetles
* Erotylidae Latreille, 1802 - pleasing fungus beetles
* Helotidae Reitter, 1876
* Hobartiidae Sen Gupta & Crowson, 1966
* Kateretidae Erichson in Agassiz, 1846 - short-winged flower beetles (= Brachypteridae)
* Laemophloeidae Ganglbauer 1899 - lined flat bark beetles
* Lamingtoniidae Sen Gupta & Crowson, 1966
* Monotomidae Laporte, 1840 - root-eating beetles
* Myraboliidae Lawrence and Britton, 1991
* Nitidulidae Latreille, 1802 - sap beetles
* Passandridae Erichson, 1845 - parasitic flat bark beetles
* Phalacridae Leach, 1815 - shining flower beetles
* Phloeostichidae Reitter, 1911
* Priasilphidae Crowson, 1973
* Protocucujidae Crowson, 1954
* Silvanidae Kirby, 1937 - silvanid flat bark beetles
* Smicripidae Horn, 1879 - palmetto beetles
* Sphindidae Jacquelin du Val, 1860 - dry-fungus beetles
* Tasmosalpingidae Lawrence and Britton, 1991
* †Parandrexidae Kirejtshuk, 1994 (Middle Jurassic-Early Cretaceous)
* †Wabbelidae Alekseev, 2017[6] (Eocene)

### Extinct genera

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* †Alloterocucus Li et al., 2022 Burmese amber, Myanmar, mid-Cretaceous (latest Albian\- earliest Cenomanian)

## References

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1. ^ a b c d e f Robertson, James A.; Ślipiński, Adam; Moulton, Matthew; Shockley, Floyd W.; Giorgi, Adriano; Lord, Nathan P.; Mckenna, Duane D.; Tomaszewska, Wioletta; Forrester, Juanita; Miller, Kelly B.; Whiting, Michael F. (2015). "Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia): Systematics of Cucujoidea and Coccinelloidea". Systematic Entomology. 40 (4): 745–778\. doi:10.1111/syen.12138. S2CID 55206626.
2. ^ Clayhills, Tom; Audisio, Paolo; Cline, Andrew R.; Mancini, Emiliano; Trizzino, Marco; Sabatelli, Simone (2016-05-11). "Unraveling cryptic species diversity in an aposematic sap beetle genus (Coleoptera: Nitidulidae: Cryptarchinae) from northern Europe". Insect Systematics & Evolution. 47 (2): 131–148\. doi:10.1163/1876312X-47022137. ISSN 1399-560X.
3. ^ "Family Kateretidae - Short-winged Flower Beetles". bugguide.net. Retrieved 2023-08-25.
4. ^ "Family Phalacridae - Shining Flower Beetles". bugguide.net. Retrieved 2023-08-25.
5. ^ "Family Nitidulidae - Sap-feeding Beetles". bugguide.net. Retrieved 2023-08-25.
6. ^ Vitalii I. Alekseev (2017). "A new coleopterous family Wabbelidae fam. nov. (Coleoptera: Cucujoidea) from Baltic amber (Cenozoic, Paleogene, Eocene)". Baltic Journal of Coleopterology. 17 (1): 29–41.

## External links

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* Media related to Cucujoidea at Wikimedia Commons
* Data related to Cucujoidea at Wikispecies

* v
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Extant Coleoptera families

* Kingdom: Animalia
* Phylum: Arthropoda
* Class: Insecta
* Subclass: Pterygota
* Infraclass: Neoptera
* Superorder: Holometabola

Suborder Archostemata

* Crowsoniellidae (Crowsoniella relicta)
* Cupedidae (reticulated beetles)
* Jurodidae (Sikhotealinia zhiltzovae)
* Micromalthidae
* Ommatidae

Suborder Adephaga

| Extant families|

* Amphizoidae (trout-stream beetles)
* Aspidytidae
* Carabidae (ground beetles)
* Cicindelidae (tiger beetles)
* Dytiscidae (predaceous diving beetles)
* Gyrinidae (whirligig beetles)
* Haliplidae (crawling water beetles)
* Hygrobiidae
* Meruidae (Meru phyllisae)
* Noteridae (burrowing water beetles)
* Trachypachidae (false ground beetles)

|

Suborder Myxophaga

* Hydroscaphidae (skiff beetles)
* Lepiceridae
* Sphaeriusidae
* Torridincolidae

Suborder Polyphaga

| Bostrichiformia|

| Bostrichoidea|

* Bostrichidae (auger beetles)
* Dermestidae (skin beetles)
* Endecatomidae
* Jacobsoniidae (Jacobson's beetles)
* Nosodendridae (wounded-tree beetles)
* Ptiniidae (furniture beetles, death watch beetles, spider beetles)

|

Derodontoidea|

* Derodontidae (tooth-necked fungus beetles)

Cucujiformia|

| Chrysomeloidea|

* Cerambycidae (longhorn beetles)
* Chrysomelidae (leaf beetles)
* Disteniidae
* Megalopodidae
* Orsodacnidae
* Oxypeltidae
* Vesperidae

|

Cleroidea|

* Acanthocnemidae (Acanthocnemus nigricans)
* Biphyllidae (false skin beetles)
* Byturidae (fruitworm beetles)
* Chaetosomatidae
* Cleridae (checkered beetles)
* Lophocateridae
* Mauroniscidae
* Melyridae (soft-wing flower beetles)
* Metaxinidae (Metaxina ornata)
* Peltidae
* Phloiophilidae (Phloiophilus edwardsi)
* Phycosecidae
* Prionoceridae
* Protopeltidae
* Rentoniidae
* Rhadalidae
* Thanerocleridae
* Thymalidae
* Trogossitidae (bark-gnawing beetles)

Coccinelloidea|

* Akalyptoischiidae
* Alexiidae
* Anamorphidae
* Bothrideridae (dry bark beetles)
* Cerylonidae (minute bark beetles)
* Coccinellidae (lady beetles, or God's cows)
* Corylophidae (minute fungus beetles)
* Discolomatidae
* Endomychidae (handsome fungus beetles)
* Eupsilobiidae
* Euxestidae (well polished beetles)
* Latridiidae (minute brown scavenger beetles)
* Murmidiidae
* Mycetaeidae
* Teredidae

Cucujoidea|

* Agapythidae (Agapytho foveicollis)
* Boganiidae
* Cavognathidae
* Cryptophagidae (silken fungus beetles)
* Cucujidae (flat bark beetles)
* Cybocephalidae
* Cyclaxyridae
* Erotylidae (pleasing fungus beetles)
* Helotidae
* Hobartiidae
* Kateretidae (short-winged flower beetles)
* Laemophloeidae (lined flat bark beetles)
* Lamingtoniidae (Lamingtonium binnaberrense)
* Monotomidae (root-eating beetles)
* Myraboliidae
* Nitidulidae (sap beetles)
* Passandridae (parasitic flat bark beetles)
* Phalacridae (shining flower beetles)
* Phloeostichidae
* Priasilphidae
* Propalticidae
* Protocucujidae
* Silvanidae (silvanid flat bark beetles)
* Smicripidae (palmetto beetles)
* Sphindidae (dry-fungus beetles)
* Tasmosalpingidae

Curculionoidea
(weevils)|

* Anthribidae (fungus weevils)
* Attelabidae (leaf-rolling weevils)
* Belidae (primitive weevils)
* Brentidae (straight snout weevils, New York weevil)
* Caridae
* Curculionidae (true weevils, bark beetles, ambrosia beetles)
* Nemonychidae (pine flower weevils)

Lymexyloidea|

* Lymexylidae (ship-timber beetles)

Tenebrionoidea|

* Aderidae (ant-like leaf beetles)
* Anthicidae (ant-like flower beetles)
* Archeocrypticidae (cryptic fungus beetles)
* Boridae (conifer bark beetles)
* Chalcodryidae
* Ciidae (minute tree-fungus beetles)
* Melandryidae (false darkling beetles)
* Meloidae (blister beetles)
* Mordellidae (tumbling flower beetles)
* Mycetophagidae (hairy fungus beetles)
* Mycteridae (palm and flower beetles)
* Oedemeridae (false blister beetle)
* Perimylopidae, or Promecheilidae
* Prostomidae (jugular-horned beetles)
* Pterogeniidae
* Pyrochroidae (fire-coloured beetles)
* Pythidae (dead log bark beetles)
* Ripiphoridae (wedge-shaped beetles)
* Salpingidae (narrow-waisted bark beetles)
* Scraptiidae (false flower beetles)
* Stenotrachelidae (false longhorn beetles)
* Synchroidae (synchroa bark beetles)
* Tenebrionidae (darkling beetles)
* Tetratomidae (polypore fungus beetles)
* Trictenotomidae
* Ulodidae
* Zopheridae (ironclad beetles, cylindrical bark beetles)

Elateriformia|

| Buprestoidea|

* Buprestidae (jewel beetles, or metallic wood-boring beetles)
* Schizopodidae

|

Byrrhoidea|

* Byrrhidae (pill beetles)
* Callirhipidae (cedar beetles)
* Chelonariidae (turtle beetles)
* Cneoglossidae
* Dryopidae (long-toed water beetles)
* Elmidae (riffle beetles)
* Eulichadidae (forest stream beetles)
* Heteroceridae (variegated mud-loving beetles)
* Limnichidae (minute mud beetles)
* Lutrochidae (travertine beetles)
* Psephenidae (water-penny beetles)
* Ptilodactylidae

Dascilloidea|

* Dascillidae (soft bodied plant beetles)
* Rhipiceridae (cicada beetle, cicada parasite beetles)

Elateroidea|

* Artematopodidae (soft-bodied plant beetles)
* Brachypsectridae (Texas beetles)
* Cantharidae (soldier beetles)
* Cerophytidae (rare click beetles)
* Elateridae (click beetles)
* Eucnemidae (false click beetles)
* Jurasaidae
* Lampyridae (fireflies)
* Lycidae (net-winged beetles)
* Omethidae (false fireflies, long-lipped beetles)
* Phengodidae (glowworm beetles)
* Rhagophthalmidae
* Sinopyrophoridae
* Throscidae (false metallic wood-boring beetles)

Rhinorhipoidea|

* Rhinorhipidae (Rhinorhipus tamborinensis)

Scirtoidea|

* Clambidae
* Decliniidae (Declinia relicta)
* Eucinetidae (plate-thigh beetles)
* Scirtidae

Scarabaeiformia|

| Scarabaeoidea|

* Belohinidae (Belohina inexpectata)
* Bolboceratidae
* Diphyllostomatidae (false stag beetles)
* Geotrupidae (dor beetles)
* Glaphyridae (bumble bee scarab beetles)
* Glaresidae (enigmatic scarab beetles)
* Hybosoridae (scavenger scarab beetles)
* Lucanidae (stag beetles)
* Ochodaeidae (sand-loving scarab beetles)
* Passalidae (betsy beetles)
* Pleocomidae (rain beetles)
* Scarabaeidae (scarabs)
* Trogidae (hide beetles)

|

Staphyliniformia|

| Histeroidea|

* Histeridae (clown beetles)
* Sphaeritidae (false clown beetles)
* Synteliidae

|

Hydrophiloidea|

* Epimetopidae
* Georissidae (minute mud-loving beetles)
* Helophoridae
* Hydrochidae
* Hydrophilidae (water scavenger beetles)
* Spercheidae

Staphylinoidea|

* Agyrtidae (primitive carrion beetles)
* Hydraenidae
* Leiodidae (round fungus beetles)
* Ptiliidae (feather-winged beetles)
* Silphidae (carrion beetles)
* Staphylinidae (rove beetles)

List of subgroups of the order Coleoptera

Taxon identifiers

Cucujoidea|

* Wikidata: Q651038
* Wikispecies: Cucujoidea
* ADW: Cucujoidea
* AFD: Cucujoidea
* BugGuide: 40618
* CoL: 4ZR
* EoL: 354
* Fauna Europaea: 11037
* Fauna Europaea (new): a4bb42d6-a457-4214-af69-26add8279d9f
* iNaturalist: 48485
* ITIS: 114289
* NBN: NHMSYS0001717771
* NCBI: 71526
* NZOR: 0d7853e6-38f8-47c4-a37f-acc7483a1d63
* Open Tree of Life: 765495
* Paleobiology Database: 69407

*[†]: Extinct
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