Ethology

Not to be confused with ethnology or ecology.

"Animal behaviour" redirects here. For other uses, see Animal behaviour (disambiguation).

For the journal, see Ethology (journal).

Scientific objective study of non-human animal behaviour

Honeybee workers perform the waggle dance to indicate the range and direction of food. Great crested grebes perform a complex synchronised courtship display. Male impalas fighting during the rut

Ethology is a branch of zoology that studies the behaviour of non-human animals. It has its scientific roots in the work of Charles Darwin and of American and German ornithologists of the late 19th and early 20th century, including Charles O. Whitman, Oskar Heinroth, and Wallace Craig. The modern discipline of ethology is generally considered to have begun during the 1930s with the work of the Dutch biologist Nikolaas Tinbergen and the Austrian biologists Konrad Lorenz and Karl von Frisch, the three winners of the 1973 Nobel Prize in Physiology or Medicine. Ethology combines laboratory and field science, with a strong relation to neuroanatomy, ecology, and evolutionary biology.[1]

## Etymology

[edit]

The modern term ethology derives from the Greek language: ἦθος, ethos meaning "character" and -λογία, -logia meaning "the study of". The term was first popularized by the American entomologist William Morton Wheeler in 1902.[2]

## History

[edit]

### The beginnings of ethology

[edit]

Charles Darwin (1809–1882) explored the expression of emotions in animals.

Ethologists have been concerned particularly with the evolution of behaviour and its understanding in terms of natural selection. In one sense, the first modern ethologist was Charles Darwin, whose 1872 book The Expression of the Emotions in Man and Animals influenced many ethologists. He pursued his interest in behaviour by encouraging his protégé George Romanes, who investigated animal learning and intelligence using an anthropomorphic method, anecdotal cognitivism, that did not gain scientific support.[3]

Other early ethologists, such as Eugène Marais, Charles O. Whitman, Oskar Heinroth, Wallace Craig and Julian Huxley, instead concentrated on behaviours that can be called instinctive in that they occur in all members of a species under specified circumstances.[4][5][2] Their starting point for studying the behaviour of a new species was to construct an ethogram, a description of the main types of behaviour with their frequencies of occurrence. This provided an objective, cumulative database of behaviour.[2]

### Growth of the field

[edit]

Due to the work of Konrad Lorenz and Niko Tinbergen, ethology developed strongly in continental Europe during the years prior to World War II.[2] After the war, Tinbergen moved to the University of Oxford, and ethology became stronger in the UK, with the additional influence of William Thorpe, Robert Hinde, and Patrick Bateson at the University of Cambridge.[6]

Lorenz, Tinbergen, and von Frisch were jointly awarded the Nobel Prize in Physiology or Medicine in 1973 for their work of developing ethology.[7]

Ethology is now a well-recognized scientific discipline, with its own journals such as Animal Behaviour, Applied Animal Behaviour Science, Animal Cognition, Behaviour, Behavioral Ecology and Ethology. In 1972, the International Society for Human Ethology was founded along with its journal, Human Ethology.[8]

### Social ethology

[edit]

In 1972, the English ethologist John H. Crook distinguished comparative ethology from social ethology, and argued that much of the ethology that had existed so far was really comparative ethology—examining animals as individuals—whereas, in the future, ethologists would need to concentrate on the behaviour of social groups of animals and the social structure within them.[9]

E. O. Wilson's book Sociobiology: The New Synthesis appeared in 1975,[10] and since that time, the study of behaviour has been much more concerned with social aspects. It has been driven by the Darwinism associated with Wilson, Robert Trivers, and W. D. Hamilton. The related development of behavioural ecology has helped transform ethology.[11] Furthermore, a substantial rapprochement with comparative psychology has occurred, so the modern scientific study of behaviour offers a spectrum of approaches. In 2020, Tobias Starzak and Albert Newen from the Institute of Philosophy II at the Ruhr University Bochum postulated that animals may have beliefs.[12]

## Determinants of behaviour

[edit]

Behaviour is determined by three major factors, namely inborn instincts, learning, and environmental factors. The latter include abiotic and biotic factors. Abiotic factors such as temperature or light conditions have dramatic effects on animals, especially if they are ectothermic or nocturnal. Biotic factors include members of the same species (e.g. sexual behavior), predators (fight or flight), or parasites and diseases.[13]

### Instinct

[edit]

Main article: Instinct

Kelp gull chicks peck at red spot on mother's beak to stimulate regurgitating reflex

Webster's Dictionary defines instinct as "A largely inheritable and unalterable tendency of an organism to make a complex and specific response to environmental stimuli without involving reason".[14] This covers fixed action patterns like beak movements of bird chicks,[15] and the waggle dance of honeybees.[16]

#### Fixed action patterns

[edit]

Main article: Fixed action pattern

An important development, associated with the name of Konrad Lorenz though probably due more to his teacher, Oskar Heinroth, was the identification of fixed action patterns. Lorenz popularized these as instinctive responses that would occur reliably in the presence of identifiable stimuli called sign stimuli or "releasing stimuli". Fixed action patterns are now considered to be instinctive behavioural sequences that are relatively invariant within the species and that almost inevitably run to completion.[15]

One example of a releaser is the beak movements of many bird species performed by newly hatched chicks, which stimulates the mother to regurgitate food for her offspring.[17] Other examples are the classic studies by Tinbergen on the egg-retrieval behaviour and the effects of a "supernormal stimulus" on the behaviour of graylag geese.[18][19]

One investigation of this kind was the study of the waggle dance ("dance language") in bee communication by Karl von Frisch.[16]

### Learning

[edit]

#### Habituation

[edit]

Main article: Habituation

Habituation is a simple form of learning and occurs in many animal taxa. It is the process whereby an animal ceases responding to a stimulus. Often, the response is an innate behavior. Essentially, the animal learns not to respond to irrelevant stimuli. For example, prairie dogs (Cynomys ludovicianus) give alarm calls when predators approach, causing all individuals in the group to quickly scramble down burrows. When prairie dog towns are located near trails used by humans, giving alarm calls every time a person walks by is expensive in terms of time and energy. Habituation to humans is therefore an important behavior in this context.[20][21][22]

#### Associative learning

[edit]

Main article: Association (psychology)

Further information: Classical conditioning and Operant conditioning

Associative learning in animal behaviour is any learning process in which a new response becomes associated with a particular stimulus.[23] The first studies of associative learning were made by the Russian physiologist Ivan Pavlov, who observed that dogs trained to associate food with the ringing of a bell would salivate on hearing the bell.[24]

#### Imprinting

[edit]

Main article: Imprinting (psychology)

Imprinting in a moose.

Imprinting enables the young to discriminate the members of their own species, vital for reproductive success. This important type of learning only takes place in a very limited period of time. Konrad Lorenz observed that the young of birds such as geese and chickens followed their mothers spontaneously from almost the first day after they were hatched, and he discovered that this response could be imitated by an arbitrary stimulus if the eggs were incubated artificially and the stimulus were presented during a critical period that continued for a few days after hatching.[25]

#### Cultural learning

[edit]

Main article: Cultural transmission in animals

##### Observational learning

[edit]

Main article: Observational learning

##### Imitation

[edit]

Main article: Imitation

Imitation is an advanced behavior whereby an animal observes and exactly replicates the behavior of another. The National Institutes of Health reported that capuchin monkeys preferred the company of researchers who imitated them to that of researchers who did not. The monkeys not only spent more time with their imitators but also preferred to engage in a simple task with them even when provided with the option of performing the same task with a non-imitator.[26] Imitation has been observed in recent research on chimpanzees; not only did these chimps copy the actions of another individual, when given a choice, the chimps preferred to imitate the actions of the higher-ranking elder chimpanzee as opposed to the lower-ranking young chimpanzee.[27]

##### Stimulus and local enhancement

[edit]

Animals can learn using observational learning but without the process of imitation. One way is stimulus enhancement in which individuals become interested in an object as the result of observing others interacting with the object.[28] Increased interest in an object can result in object manipulation which allows for new object-related behaviours by trial-and-error learning. Haggerty (1909) devised an experiment in which a monkey climbed up the side of a cage, placed its arm into a wooden chute, and pulled a rope in the chute to release food. Another monkey was provided an opportunity to obtain the food after watching a monkey go through this process on four occasions. The monkey performed a different method and finally succeeded after trial-and-error.[29] In local enhancement, a demonstrator attracts an observer's attention to a particular location.[30] Local enhancement has been observed to transmit foraging information among birds, rats and pigs.[31] The stingless bee (Trigona corvina) uses local enhancement to locate other members of their colony and food resources.[32]

##### Social transmission

[edit]

See also: Cultural transmission in animals

A well-documented example of social transmission of a behaviour occurred in a group of macaques on Hachijojima Island, Japan. The macaques lived in the inland forest until the 1960s, when a group of researchers started giving them potatoes on the beach: soon, they started venturing onto the beach, picking the potatoes from the sand, and cleaning and eating them.[10] About one year later, an individual was observed bringing a potato to the sea, putting it into the water with one hand, and cleaning it with the other. This behaviour was soon expressed by the individuals living in contact with her; when they gave birth, this behaviour was also expressed by their young—a form of social transmission.[33]

#### Teaching

[edit]

See also: Animal culture § Teaching

Teaching is a highly specialized aspect of learning in which the "teacher" (demonstrator) adjusts their behaviour to increase the probability of the "pupil" (observer) achieving the desired end-result of the behaviour. For example, orcas are known to intentionally beach themselves to catch pinniped prey.[34] Mother orcas teach their young to catch pinnipeds by pushing them onto the shore and encouraging them to attack the prey. Because the mother orca is altering her behaviour to help her offspring learn to catch prey, this is evidence of teaching.[34] Teaching is not limited to mammals. Many insects, for example, have been observed demonstrating various forms of teaching to obtain food. Ants, for example, will guide each other to food sources through a process called "tandem running," in which an ant will guide a companion ant to a source of food.[35] It has been suggested that the pupil ant is able to learn this route to obtain food in the future or teach the route to other ants. This behaviour of teaching is also exemplified by crows, specifically New Caledonian crows. The adults (whether individual or in families) teach their young adolescent offspring how to construct and utilize tools. For example, Pandanus branches are used to extract insects and other larvae from holes within trees.[36]

## Mating and the fight for supremacy

[edit]

Courtship display of a sarus crane

Individual reproduction is the most important phase in the proliferation of individuals or genes within a species: for this reason, there exist complex mating rituals, which can be very complex even if they are often regarded as fixed action patterns. The stickleback's complex mating ritual, studied by Tinbergen, is regarded as a notable example.[37]

Often in social life, animals fight for the right to reproduce, as well as social supremacy. A common example of fighting for social and sexual supremacy is the so-called pecking order among poultry. Every time a group of poultry cohabitate for a certain time length, they establish a pecking order. In these groups, one chicken dominates the others and can peck without being pecked. A second chicken can peck all the others except the first, and so on. Chickens higher in the pecking order may at times be distinguished by their healthier appearance when compared to lower level chickens.[citation needed] While the pecking order is establishing, frequent and violent fights can happen, but once established, it is broken only when other individuals enter the group, in which case the pecking order re-establishes from scratch.[38]

## Social behaviour

[edit]

Several animal species, including humans, tend to live in groups. Group size is a major aspect of their social environment. Social life is probably a complex and effective survival strategy. It may be regarded as a sort of symbiosis among individuals of the same species: a society is composed of a group of individuals belonging to the same species living within well-defined rules on food management, role assignments and reciprocal dependence.[39]

When biologists interested in evolution theory first started examining social behaviour, some apparently unanswerable questions arose, such as how the birth of sterile castes, like in bees, could be explained through an evolving mechanism that emphasizes the reproductive success of as many individuals as possible, or why, amongst animals living in small groups like squirrels, an individual would risk its own life to save the rest of the group. These behaviours may be examples of altruism.[40] Not all behaviours are altruistic, as indicated by the table below. For example, revengeful behaviour was at one point claimed to have been observed exclusively in Homo sapiens. However, other species have been reported to be vengeful including chimpanzees,[41] as well as anecdotal reports of vengeful camels.[42]

Classification of social behaviours[39] Type of behaviour  | Effect on the donor  | Effect on the receiver

|
|

Egoistic | Neutral to Increases fitness  | Decreases fitness
Cooperative | Neutral to Increases fitness  | Neutral to Increases fitness
Altruistic | Decreases fitness  | Neutral to Increases fitness
Revengeful | Decreases fitness  | Decreases fitness

Altruistic behaviour has been explained by the gene-centred view of evolution.[43][44]

### Benefits and costs of group living

[edit]

One advantage of group living is decreased predation. If the number of predator attacks stays the same despite increasing prey group size, each prey has a reduced risk of predator attacks through the dilution effect.[11][page needed] Further, according to the selfish herd theory, the fitness benefits associated with group living vary depending on the location of an individual within the group. The theory suggests that conspecifics positioned at the centre of a group will reduce the likelihood predations while those at the periphery will become more vulnerable to attack.[45] In groups, prey can also actively reduce their predation risk through more effective defence tactics, or through earlier detection of predators through increased vigilance.[11]

Another advantage of group living is an increased ability to forage for food. Group members may exchange information about food sources, facilitating the process of resource location.[11][page needed] Honeybees are a notable example of this, using the waggle dance to communicate the location of flowers to the rest of their hive.[46] Predators also receive benefits from hunting in groups, through using better strategies and being able to take down larger prey.[11][page needed]

Some disadvantages accompany living in groups. Living in close proximity to other animals can facilitate the transmission of parasites and disease, and groups that are too large may also experience greater competition for resources and mates.[47]

### Group size

[edit]

Theoretically, social animals should have optimal group sizes that maximize the benefits and minimize the costs of group living. However, in nature, most groups are stable at slightly larger than optimal sizes.[11][page needed] Because it generally benefits an individual to join an optimally-sized group, despite slightly decreasing the advantage for all members, groups may continue to increase in size until it is more advantageous to remain alone than to join an overly full group.[48]

## Tinbergen's four questions for ethologists

[edit]

Main article: Tinbergen's four questions

Tinbergen argued that ethology needed to include four kinds of explanation in any instance of behaviour:[49][50]

* Function – How does the behaviour affect the animal's chances of survival and reproduction? Why does the animal respond that way instead of some other way?
* Causation – What are the stimuli that elicit the response, and how has it been modified by recent learning?
* Development – How does the behaviour change with age, and what early experiences are necessary for the animal to display the behaviour?
* Evolutionary history – How does the behaviour compare with similar behaviour in related species, and how might it have begun through the process of phylogeny?

These explanations are complementary rather than mutually exclusive—all instances of behaviour require an explanation at each of these four levels. For example, the function of eating is to acquire nutrients (which ultimately aids survival and reproduction), but the immediate cause of eating is hunger (causation). Hunger and eating are evolutionarily ancient and are found in many species (evolutionary history), and develop early within an organism's lifespan (development). It is easy to confuse such questions—for example, to argue that people eat because they are hungry and not to acquire nutrients—without realizing that the reason people experience hunger is because it causes them to acquire nutrients.[51]

## See also

[edit]

* Psychology portal
* Zoology portal

* Animal behavior consultant
* Anthrozoology
* Behavioral ecology
* Cognitive ethology
* Deception in animals
* Human ethology
* List of abnormal behaviours in animals
* Tool use by non-human animals

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[edit]

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## Further reading

[edit]

* Burkhardt, Richard W. Jr. "On the Emergence of Ethology as a Scientific Discipline." Conspectus of History 1.7 (1981).

## External links

[edit]

* Media related to Ethology at Wikimedia Commons

* v
* t
* e

Ethology

Branches|

* Animal cognition
* Animal communication
* Animal consciousness
* Animal culture
* Animal sexual behaviour
* Animal welfare science
* Anthrozoology
* Bee learning and communication
* Behavioural ecology
* Behavioral endocrinology
* Behavioural genetics
* Breed
* Cognitive ethology
* Comfort behaviour
* Grooming
* Comparative psychology
* Emotion in animals
* Ethogram
* Evolutionary neuroscience
* Feeding
* Hover
* Human ethology
* Instinct
* Learning
* Neuroethology
* Pain in animals
* Philosophical ethology
* Sociobiology
* Species-typical behavior
* Stereotypy
* Structures
* Hive
* Honeycomb
* Nest
* Instinct
* Swarm
* Tool use by non-humans
* Zoosemiotics
* Zoomusicology

Ethologists|

* Patrick Bateson
* Marc Bekoff
* Donald Broom
* John B. Calhoun
* Charles Darwin
* Marian Dawkins
* Richard Dawkins
* Irenäus Eibl-Eibesfeldt
* Dian Fossey
* Karl von Frisch
* Jane Goodall
* Heini Hediger
* Julian Huxley
* Konrad Lorenz
* Desmond Morris
* Thomas Sebeok
* William Homan Thorpe
* Nikolaas Tinbergen
* Jakob von Uexküll
* Wolfgang Wickler
* E. O. Wilson
* Solly Zuckerman

Societies|

* Association for the Study of Animal Behaviour
* International Society for Applied Ethology

Journals|

* Animal Behaviour
* Animal Cognition
* Animal Welfare
* Behavioral Ecology
* Behaviour

* Category

* v
* t
* e

Biological rhythms

Internal rhythms|

* Circadian rhythm
* Infradian rhythm
* Ultradian rhythm
* Lunar rhythm
* Annual rhythm

External cycles|

* Diurnality
* Diurnal cycle
* Nocturnality
* Cathemerality
* Crepuscular
* Matutinal
* Vespertine
* Seasonality
* Photoperiodism
* Vernalization

Fields|

* Behavioural ecology
* Chronobiology
* Ethology

See also|

* Chronotype
* Diel vertical migration
* Light pollution
* Solunar theory
* Parts of a day

* v
* t
* e

Sociobiology

Topics|

* Behavioural genetics
* Challenge hypothesis
* Dual inheritance theory
* Ethology
* Evolutionary psychology
* Evolution of morality
* Evolutionary models of food sharing
* Group selection
* Kin selection
* Inbreeding avoidance
* Kin recognition
* Male warrior hypothesis
* Reciprocal altruism
* Sexual selection
* in humans
* Sex and psychology
* Sociality
* eusociality
* evolution
* presociality
* Dunbar's number
* Polyethism

Supporters|

* Anne Campbell
* Noam Chomsky
* Richard Dawkins
* Daniel Dennett
* Sarah Blaffer Hrdy
* Steven Pinker
* Frans de Waal
* E. O. Wilson
* Sociobiology: The New Synthesis
* Robert Wright

Opponents|

* Stephen Jay Gould
* Leon Kamin
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* Not in Our Genes

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* Commons
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Animal communication

Concepts|

* Animal training
* Animal language
* Animal cognition
* Anecdotal cognitivism
* Bioacoustics
* Deception in animals
* Tactical
* Ethology
* Evolutionary linguistics
* FOXP2
* FOXP2 and human evolution
* Human–animal communication
* Mating call
* Origin of language
* Seismic communication
* Self-anointing in animals
* Talking animals

Animal-specific|

* Aquatic animals
* Whale vocalization
* Bees
* Bumblebees
* Birds
* Talking birds
* Cats
* Dogs
* Elephants
* Frogs
* Great ape language
* Yerkish
* Insects
* Lizards
* Wolves

Related|

* Roar
* Category:Talking birds
* Category:Individual apes involved in language studies

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Zoology

* Outline

Branches|

* Anthrozoology
* Arthropodology (Arachnology (Acarology)
* Carcinology
* Entomology (Coleopterology
* Lepidopterology
* Melittology
* Myrmecology
* Orthopterology)
* Myriapodology)
* Biological anthropology
* Bryozoology
* Cnidariology
* Ethnozoology
* Ethology
* Helminthology
* Herpetology (Batrachology
* Ophiology
* Testudinology)
* Ichthyology
* Malacology (Conchology
* Teuthology)
* Mammalogy (Cetology
* Cynology
* Felinology
* Hippology
* Primatology)
* Nematology
* Neuroethology
* Ornithology
* Parasitology
* Paleozoology
* Planktology
* Zooarchaeology

Animal groups|

* Porifera
* Ctenophora
* Placozoa
* Cnidaria
* Xenacoelomorpha
* Ambulacraria
* Chordata
* Aschelminthes
* Arthropoda
* Platyhelminthes
* Mollusca
* Annelida

Animal anatomy
Animal morphology|

| Histology|

* Tissues
* Epithelial tissue
* Muscular tissue
* Nervous tissue
* Connective tissues
* Mineralized tissues
* Molecular anatomy

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Anatomy
and morphology|

* Mammalian anatomy and morphology
* Human
* Dog
* Elephant
* Cat
* Bird anatomy
* Fish anatomy
* Shark anatomy
* Spider anatomy
* Insect morphology

Animal physiology|

| General physiology|

* Respiration
* Respiratory system
* Breathing
* Gas exchange
* Respiratory pigment
* Cellular respiration
* Vascular system
* Blood
* Lymph
* Blood vessels
* Arteries
* Veins
* Capillaries
* Heart
* Water vascular system

|

By species|

* Insect physiology
* Fish physiology

Zoologists|

* Karl Ernst von Baer
* Georges Cuvier
* Charles Darwin
* Jean-Henri Fabre
* William Kirby
* Carl Linnaeus
* Konrad Lorenz
* Thomas Say
* Jakob von Uexküll
* Alfred Russel Wallace

History|

* Pre-Darwin
* Post-Darwin
* Timeline of zoology

* Category

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Evolutionary psychology

* History
* Evolutionary thought
* Theoretical foundations
* Adaptationism
* Cognitive revolution
* Cognitivism
* Gene selection theory
* Modern synthesis
* Criticism

Evolutionary
processes|

* Adaptations
* Altruism
* Cheating
* Hamiltonian spite
* Reciprocal
* Baldwin effect
* By-products
* Evolutionarily stable strategy
* Exaptation
* Fitness
* Inclusive
* Kin selection
* Inbreeding avoidance
* Kin recognition
* Parental investment
* Parent–offspring conflict
* Mismatch
* Natural selection
* Sexual selection
* Costly signaling
* Male/female intrasexual competition
* Mate choice
* Sexual dimorphism
* Social selection
* Species-typical behavior

Areas|

| Cognition /
Emotion|

* Affect
* Display
* Display rules
* Facial expression
* Behavioral modernity
* Cognitive module/modularity of mind
* Adaptive unconscious/Unconscious cognition
* Automatic and controlled processes
* Cognitive specialization
* Computational theory of mind
* Domain generality
* Domain specificity/learning
* Dual process theory
* Cognitive tradeoff hypothesis
* Evolution of the brain
* Evolution of nervous systems
* Fight-or-flight response
* Arachnophobia
* Basophobia
* Ophidiophobia
* Folk biology/taxonomy
* Folk psychology/theory of mind
* Intelligence
* Flynn effect
* Wason selection task
* Motor control/skill
* Multitasking
* Numerical cognition
* Number sense
* Approximate number system
* Parallel individuation system
* Sleep
* Visual perception
* Color vision
* Eye
* Naïve physics

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Culture|

* Aesthetics
* Literary criticism
* Musicology
* Anthropology
* Biological
* Crime
* Language
* Origin
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* Speech
* Morality
* Moral foundations
* Religion
* Origin
* Universals

Development|

* Attachment
* Bonding
* Affectional/maternal/paternal bond
* Caregiver deprivation
* Childhood attachment
* Cinderella effect
* Cognitive development
* Education
* Language acquisition
* Personality development
* Socialization

Human factors /
Mental health|

* Cognitive ergonomics
* Computer-mediated communication
* Engineering psychology
* Human–computer interaction
* Media naturalness theory
* Neuroergonomics
* Depression
* Digital media use and mental health
* Hypophobia
* Imprinted brain hypothesis
* Mind-blindness
* Psychological effects of Internet use
* Rank theory of depression
* Schizophrenia
* Screen time
* Smartphones and pedestrian safety
* Social aspects of television
* Societal effects of cars
* Distracted driving
* Lead–crime hypothesis
* Mobile phones and driving safety
* Texting while driving

Sex|

* Activity
* Adult attachment
* Age disparity
* Arousal
* Concealed ovulation
* Coolidge effect
* Desire
* Fantasy
* Hormonal motivation
* Incest taboo
* Westermarck effect
* Jealousy
* Mate guarding
* Mating preferences
* Mating strategies
* Orientation
* Ovulatory shift hypothesis
* Pair bond
* Physical/Sexual attraction
* Sexuality/male/female
* Sexy son hypothesis

Sex differences|

* Aggression
* Autism
* Cognition
* Crime
* Division of labour
* Emotional intelligence
* Empathising–systemising theory
* Gender role
* Intelligence
* Memory
* Mental health
* Narcissism
* Neuroscience
* Schizophrenia
* Substance abuse
* Suicide
* Variability hypothesis

Related subjects|

| Academic disciplines|

* Behavioral/evolutionary economics
* Behavioral epigenetics/genetics
* Affective/behavioral/cognitive/evolutionary neuroscience
* Biocultural anthropology
* Biolinguistics
* Biological psychiatry
* Cognitive psychology
* Cognitive science
* Cross-cultural psychology
* Ethology
* Evolutionary biology
* Evolutionary medicine
* Functional psychology
* Neuropsychology
* Philosophy of mind
* Population genetics
* Primatology
* Sociobiology

|

Research topics|

* Cultural evolution
* Evolutionary epistemology
* Great ape language
* Human–animal communication
* Missing heritability problem
* Number sense in animals
* Primate cognition
* Unit of selection
* Coevolution
* Cultural group selection
* Dual inheritance theory
* Fisher's principle
* Group selection
* Hologenome theory
* Lamarckism/acquired characteristics
* Population
* Punctuated equilibrium
* Recent human evolution
* Species
* Species complex
* Transgenerational epigenetic inheritance
* Trivers–Willard hypothesis

Theoretical positions|

* Cultural selection theory
* Determinism/indeterminism
* Biological determinism
* Connectionism
* Cultural determinism
* Environmental determinism
* Nature versus nurture
* Psychological nativism
* Social constructionism
* Social determinism
* Standard social science model
* Functionalism
* Memetics
* Multilineal evolution
* Neo-Darwinism
* Neoevolutionism
* Sociocultural evolution
* Unilineal evolution

* Evolutionary psychologists
*  Evolutionary psychology
*  Psychology portal
*  Evolutionary biology portal

Authority control databases

International|

* FAST

National|

* United States
* France
* BnF data
* Japan
* Czech Republic
* 2
* Spain
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Other|

* Encyclopedia of Modern Ukraine

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