Gnaphosoidea

Superfamily of spiders

Gnaphosoidea

Gnaphosa bicolor
Scientific classification
Domain:  | Eukaryota
Kingdom:  | Animalia
Phylum:  | Arthropoda
Subphylum:  | Chelicerata
Class:  | Arachnida
Order:  | Araneae
Infraorder:  | Araneomorphae
Clade:  | Entelegynae
Clade:  | Dionycha
Superfamily:  | Gnaphosoidea
Simon, 1893[1]
Families

See text

The Gnaphosoidea or gnaphosoids are a superfamily of araneomorph spiders with seven families. A 2014 study did not find the group to be monophyletic.

## Phylogeny

[edit]

Gnaphosoidea has been circumscribed to contain the following families:[1]

* Ammoxenidae
* Cithaeronidae
* Gallieniellidae
* Gnaphosidae
* Lamponidae
* Trochanteriidae

Gnaphosoidea has been placed in the Dionycha clade, itself part of the RTA clade:[2]

RTA clade  |

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basal clades

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| Dionycha  |

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other clades

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Gnaphosoidea

Grate-shaped tapetum clade (including lycosoids)

The Prodidomidae, Lamponidae and Gnaphosidae have been considered "higher gnaphosoids", sharing anterior lateral spinnerets consisting of only a single "joint" (article); the "lower gnaphosoids" (Ammoxenidae, Cithaeronidae, Gallieniellidae and Trochanteriidae) retain a distal article that is represented by an entire ring of hardened (sclerotized) cuticle.[3] (Earlier the Lamponidae were grouped with the "lower gnaphosoids", having spinnerets of an intermediate kind.[4]) One hypothesis for the internal phylogeny of the gnaphosoids, defined in this way, is:[2]

Gnaphosoidea  |

| "lower gnaphosoids"  |

|  |

|  |

Gallieniellidae

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Trochanteridae

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Cithaeronidae

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Ammoxenidae

"higher gnaphosoids"  |

|  |

Lamponidae

|

|  |

Gnaphosidae

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Prodidomidae

A 2014 study of dionychan spiders did not recover Gnaphosoidea as a monophyletic group, instead finding "gnaphosoid" families other than Gnaphosidae and Prodidomidae to be part of a larger clade, mixed in with three other dionychan families, Liocranidae, Trachelidae and Phrurolithidae. Forcing Gnaphosoidea to be monophyletic produced results described as "quite suboptimal".[5]

## References

[edit]

1. ^ a b Dunlop, Jason A. & Penney, David (2011). "Order Araneae Clerck, 1757" (PDF). In Zhang, Z.-Q. (ed.). Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa. Auckland, New Zealand: Magnolia Press. ISBN 978-1-86977-850-7. Retrieved 2015-10-31.
2. ^ a b Nentwig, Wolfgang, ed. (2013), "Appendix : Spider Phylogeny" (PDF), Spider Ecophysiology, Springer, ISBN 978-3-642-33988-2, retrieved 2015-11-03
3. ^ Platnick, Norman I. & Baehr, Barbara C. (2006). "A revision of the Australasian ground spiders of the family Prodidomidae (Araneae, Gnaphosoidea)" (PDF). Bulletin of the American Museum of Natural History. 298 (298): 1–287. doi:10.1206/0003-0090(2006)298[1:AROTAG]2.0.CO;2. S2CID 83420288.
4. ^ Platnick, N.I. (2002), "A revision of the Australasian ground spiders of the families Ammoxenidae, Cithaeronidae, Gallieniellidae, and Trochanteriidae (Araneae: Gnaphosoidea)", Bulletin of the American Museum of Natural History, 271 (271): 1–244, doi:10.1206/0003-0090(2002)271<0001:AROTAG>2.0.CO;2, hdl:2246/494, S2CID 86321070, retrieved 2015-11-20
5. ^ Ramírez, M. (2014). "The morphology and phylogeny of Dionychan spiders (Araneae: Araneomorphae)". Bulletin of the American Museum of Natural History. 390 (390): 1–374. doi:10.1206/821.1. hdl:2246/6537. S2CID 86146467. pp. 323–327.

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Extant Araneae families

* Kingdom: Animalia
* Phylum: Arthropoda
* Subphylum: Chelicerata
* Class: Arachnida

Suborder Mesothelae

* Liphistiidae (segmented spiders)
* Heptathelidae (segmented spiders)

Suborder Opisthothelae

| Mygalomorphae|

* Actinopodidae (mouse spiders and relatives)
* Antrodiaetidae (folding trapdoor spiders)
* Atracidae (Australian funnel-web spiders)
* Atypidae (atypical tarantulas or purseweb spiders)
* Barychelidae (brushed trapdoor spiders)
* Ctenizidae (cork-lid trapdoor spiders)
* Cyrtaucheniidae (wafer trapdoor spiders)
* Dipluridae (funnel-web tarantulas)
* Euctenizidae
* Halonoproctidae
* Hexathelidae (funnel-webs or venomous funnel-web tarantulas)
* Idiopidae
* Macrothelidae
* Mecicobothriidae (dwarf tarantulas)
* Microstigmatidae
* Migidae (tree trapdoor spiders)
* Nemesiidae (funnel-web tarantulas)
* Paratropididae (bald-legged spiders)
* Porrhothelidae
* Theraphosidae (true tarantulas)

Araneomorphae|

| Non-entelegynes|

* Archaeidae (pelican spiders)
* Austrochilidae
* Caponiidae
* Diguetidae (coneweb spiders)
* Drymusidae (false violin spiders)
* Dysderidae (woodlouse hunters)
* Filistatidae (crevice weaver spiders)
* Gradungulidae (large-clawed spiders)
* Huttoniidae
* Hypochilidae (lampshade spiders)
* Leptonetidae
* Mecysmaucheniidae
* Ochyroceratidae (midget ground weavers)
* Oonopidae (goblin spiders)
* Orsolobidae
* Pacullidae
* Palpimanidae (palp-footed spiders)
* Periegopidae
* Pholcidae (cellar spiders)
* Plectreuridae
* Scytodidae (spitting spiders)
* Segestriidae (tube-dwelling spiders)
* Sicariidae (violin spiders, assassin spiders)
* Stenochilidae
* Telemidae (long-legged cave spiders)
* Tetrablemmidae (armored spiders)
* Trogloraptoridae (Trogloraptor marchingtoni)

Entelegynae|

* Agelenidae (araneomorph funnel weavers)
* Amaurobiidae (tangled nest spiders)
* Ammoxenidae (termite hunters)
* Anapidae
* Anyphaenidae (anyphaenid sac spiders)
* Araneidae (orb-weaver spiders)
* Arkyidae
* Cheiracanthiidae
* Cithaeronidae
* Clubionidae (sac spiders)
* Corinnidae (dark sac spiders)
* Ctenidae (wandering spiders or tropical wolf spiders)
* Cyatholipidae
* Cybaeidae
* Cycloctenidae
* Deinopidae (net-casting spiders)
* Desidae (intertidal spiders)
* Dictynidae
* Eresidae (velvet spiders)
* Gallieniellidae
* Gnaphosidae (flat-bellied ground spiders)
* Hahniidae (dwarf sheet spiders)
* Hersiliidae (tree trunk spiders)
* Homalonychidae
* Lamponidae (white-tailed spiders)
* Linyphiidae (sheet weavers or money spiders)
* Liocranidae (liocranid sac spiders)
* Lycosidae (wolf spiders)
* Malkaridae (shield spiders)
* Megadictynidae
* Mimetidae (pirate spiders)
* Miturgidae
* Mysmenidae (spurred orb-weavers)
* Nesticidae (cave cobweb spiders)
* Nicodamidae (red and black spiders)
* Oecobiidae (disc web spiders)
* Oxyopidae (lynx spiders)
* Penestomidae
* Philodromidae (running crab spiders)
* Phrurolithidae
* Physoglenidae
* Phyxelididae
* Pimoidae
* Pisauridae (nursery web spiders) (including Halidae)
* Psechridae
* Salticidae (jumping spiders)
* Selenopidae
* Senoculidae (bark hunters)
* Sparassidae (huntsman spiders)
* Stiphidiidae
* Symphytognathidae (dwarf orb-weavers)
* Synaphridae
* Synotaxidae
* Tetragnathidae (long jawed orb-weavers)
* Theridiidae (tangle-web spiders)
* Theridiosomatidae (ray spiders)
* Thomisidae (crab spiders)
* Titanoecidae
* Toxopidae
* Trachelidae
* Trechaleidae
* Trochanteriidae
* Udubidae
* Uloboridae (cribellate orb weavers)
* Viridasiidae
* Xenoctenidae
* Zodariidae (ant spiders)
* Zoropsidae

* Spider taxonomy
* Bold are families with more than 1000 species

Taxon identifiers

Gnaphosoidea|

* Wikidata: Q54895
* Wikispecies: Gnaphosoidea
* GBIF: 4660193
* iNaturalist: 367173
* Paleobiology Database: 268858

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