Heleomyzidae

Family of flies

Heleomyzidae

Suillia sp.
Scientific classification
Domain:  | Eukaryota
Kingdom:  | Animalia
Phylum:  | Arthropoda
Class:  | Insecta
Order:  | Diptera
Section:  | Schizophora
Subsection:  | Acalyptratae
Superfamily:  | Sphaeroceroidea
Family:  | Heleomyzidae
Westwood, 1840
Subfamilies

* Borboropsinae
* Chiropteromyzinae
* Cinderellinae
* Cnemospathidinae
* Diaciinae
* Heleomyzinae
* Heteromyzinae
* Rhinotorinae
* Suilliinae
* Tapeigastrinae
* Trixoscelidinae

Diversity
At least 80 genera
Mating behaviour of Suillia cf. bicolor on fungus Tephrochlamys Eccoptomera longiseta

The Heleomyzidae is a small family of true flies in the insect order Diptera. Over 740 described species of Heleomyzidae occur in about 76 genera[1] distributed throughout the world.

## Description[edit]

Heleomyzids are small to medium-sized flies which vary in colour from yellow to reddish yellow or reddish brown to black. The wings often have small but distinctly longer, well-spaced spines mixed with the shorter spines along the leading edge and the crossveins are often clouded.

## Taxonomy[edit]

See also: List of Heleomyzidae genera

Over 740 described species of Heleomyzidae occur in about 76 genera[1] and 22 tribes distributed throughout the world; the greatest number occur in the Holarctic region. Around 100 species of Heleomyzidae are found in North America. Most of the subfamilies have been commonly recognized as families in the past, but are now included within the Heleomyzidae.[2] The composition and monophyly of the family continues to be controversial. McAlpine[3] recently combined Heleomyzidae and Sphaeroceridae into Heteromyzidae, but this arrangement has not been widely accepted.

## Ecology[edit]

Adults of Borboroides and Heleomicra are attracted to carcasses and faeces. Larvae feed on decaying plant and animal matter, mushrooms, and various fungi. The larvae of the Holarctic Suilliinae and Tapeigaster occur principally in fungi. Larvae of Cairnsimyia live in borer tunnels in trees.

## References[edit]

1. ^ a b Pape, Thomas, Vladimir Blagoderov, and Michail B. Mostovski. "Order Diptera Linnaeus, 1758." Animal Biodiversity. Magnolia Press, 2011.
2. ^ McAlpine, David K. "The Australian genera of Heleomyzidae (Diptera: Schizophora) and a reclassification of the family into tribes." Records of the Australian Museum 36.5 (1985): 203-251.
3. ^ McAlpine, David K. "Review of the Borboroidini or Wombat Flies (Diptera: Heteromyzidae), with reconsider-ation of the status of families Heleomyzidae and Sphaeroceridae, and descriptions of femoral gland-baskets." RECORDS-AUSTRALIAN MUSEUM 59.2/3 (2007): 143

## External links[edit]

Wikimedia Commons has media related to Heleomyzidae.

Wikispecies has information related to Heleomyzidae.

* Family description and figures

### Identification resources[edit]

#### Palaearctic[edit]

* Czerny, L., 1924. Monographie der Helomyziden. Abhandlung der Zoologischen-Botanischen Gesellschaft in Wien, 15(1): 1–166. Keys genera and species.
* Czerny, L., 1937. Ergänzungen zu meiner Monographie der Helomyziden. X. Konowia, 16(2): 137–142.
* Gorodkov, K.B., 1984. Family Heleomyzidae (Helomyzidae), 15–45.In: Soós Á. and L. Papp, (eds.), Catalogue of Palaearctic Diptera, vol.10, Budapest.
* Papp, T.,1978. Some cavernicolous Diptera of the Geneva Museum. Revue Suisse de Zoologie 85: 99-106.
* Papp, T., 1998. Heleomyzidae. Chapter 3.41, 439–455. In: Papp L. and B. Darvas, (eds.), Contribution to a manual of Palaearctic Diptera. Vol. 3. 1998. Science Herald, Budapest, 880 pp.

#### United Kingdom[edit]

* Collin, J.E. 1943b. The British species of Helomyzidae (Diptera). Entomologist's Monthly Magazine 79: 234–251.

#### Nearctic[edit]

* Gill, G.D., 1962. The Heleomyzid flies of America north of Mexico(Diptera: Heleomyzidae). Proceedings of the U.S. National Museum, 113(3465): 495–603. Keys genera and species.
* Gill, G.D and B.V. Peterson, 1987.1989. Heleomyzidae, 973–980. In:McAlpine J. F., (ed.), Manual of Nearctic Diptera Vol. 2, Research Branch Agric. Canada, Monograph no. 28, IV + 675–1332. Keys genera and species.

#### Australasia[edit]

* McAlpine, D.K., 1985. The Australian genera of Heleomyzidae (Diptera: Schizophora) and a reclassification of the family into tribes. Records of the Australian Museum, 36: 203–251.

### Taxonomic lists[edit]

* World list
* The European subfamilies, tribes, genera and species - Fauna Europaea
* West Palaearctic including Russia
* Nearctic
* Australasian/Oceania
* Japan

* v
* t
* e

Extant Diptera families

* Kingdom: Animalia
* Phylum: Arthropoda
* Class: Insecta
* Subclass: Pterygota
* Infraclass: Neoptera
* Superorder: Endopterygota

Suborder Nematocera

| Axymyiomorpha|

* Axymyiidae

|

| Culicomorpha|

| Culicoidea|

* Dixidae (meniscus midges)
* Corethrellidae (frog-biting midges)
* Chaoboridae (phantom midges)
* Culicidae (mosquitoes)

|

Chironomoidea|

* Thaumaleidae (solitary midges)
* Simuliidae (black flies)
* Ceratopogonidae (biting midges)
* Chironomidae (non-biting midges)

| Blephariceromorpha|

* Blephariceridae (net-winged midges)
* Deuterophlebiidae (mountain midges)
* Nymphomyiidae

|

| Bibionomorpha|

| Bibionoidea|

* Bibionidae (march flies, lovebugs)

|

Anisopodoidea|

* Anisopodidae (wood gnats)

Sciaroidea
(fungus gnats)|

* Bolitophilidae
* Cecidomyiidae (gall midges)
* Diadocidiidae
* Ditomyiidae
* Keroplatidae (predatory fungus gnats)
* Lygistorrhinidae (long-beaked fungus gnats)
* Mycetophilidae
* Rangomaramidae (long-winged fungus gnats)
* Sciaridae (dark-winged fungus gnats)

| Perissommatomorpha|

* Perissommatidae

|

| Psychodomorpha|

| Scatopsoidea|

* Canthyloscelidae
* Scatopsidae (minute black scavenger flies, or dung midges)
* Valeseguyidae

|

Psychodoidea|

* Psychodidae (moth flies)

| Ptychopteromorpha|

* Ptychopteridae (phantom crane flies)
* Tanyderidae (primitive crane flies)

|

| Tipulomorpha|

| Trichoceroidea|

* Trichoceridae (winter crane flies)

|

Tipuloidea|

* Cylindrotomidae (long-bodied crane flies)
* Limoniidae (limoniid crane flies)
* Pediciidae (hairy-eyed craneflies)
* Tipulidae (large crane flies)

Suborder Brachycera

| Asilomorpha|

| Asiloidea|

* Apioceridae (flower-loving flies)
* Apsilocephalidae
* Apystomyiidae
* Asilidae (robber flies)
* Bombyliidae (bee flies)
* Evocoidae
* Hilarimorphidae (hilarimorphid flies)
* Mydidae (mydas flies)
* Mythicomyiidae
* Scenopinidae (window flies)
* Therevidae (stiletto flies)

|

Empidoidea|

* Atelestidae
* Dolichopodidae (long-legged flies)
* Empididae (dagger flies, balloon flies)
* Homalocnemiidae
* Hybotidae (dance flies)
* Oreogetonidae
* Ragadidae

Nemestrinoidea|

* Acroceridae (small-headed flies)
* Nemestrinidae (tangle-veined flies)

Muscomorpha|

| Aschiza|

| Platypezoidea|

* Ironomyiidae (ironic flies)
* Lonchopteridae (spear-winged flies)
* Opetiidae (flat-footed flies)
* Phoridae (scuttle flies, coffin flies, humpbacked flies)
* Platypezidae (flat-footed flies)

|

Syrphoidea|

* Pipunculidae (big-headed flies)
* Syrphidae (hoverflies)

| Schizophora|

| Acalyptratae|

| Conopoidea|

* Conopidae (thick-headed flies)

|

Tephritoidea|

* Pallopteridae (flutter flies)
* Piophilidae (cheese flies)
* Platystomatidae (signal flies)
* Pyrgotidae
* Richardiidae
* Tephritidae (peacock flies)
* Ulidiidae (picture-winged flies)

Nerioidea|

* Cypselosomatidae
* Micropezidae (stilt-legged flies)
* Neriidae (cactus flies, banana stalk flies)

Diopsoidea|

* Diopsidae (stalk-eyed flies)
* Gobryidae
* Megamerinidae
* Nothybidae
* Psilidae (rust flies)
* Somatiidae
* Strongylophthalmyiidae
* Syringogastridae
* Tanypezidae

Sciomyzoidea|

* Coelopidae (kelp flies)
* Dryomyzidae
* Helcomyzidae
* Helosciomyzidae
* Heterocheilidae
* Huttoninidae
* Natalimyzidae
* Phaeomyiidae
* Ropalomeridae
* Sciomyzidae (marsh flies)
* Sepsidae (black scavenger flies)

Sphaeroceroidea|

* Chyromyidae
* Heleomyzidae
* Nannodastiidae
* Sphaeroceridae (small dung flies)

Lauxanioidea|

* Celyphidae (beetle-backed flies)
* Chamaemyiidae (aphid flies)
* Cremifaniidae
* Lauxaniidae

Opomyzoidea|

* Agromyzidae (leaf miner flies)
* Anthomyzidae
* Asteiidae
* Aulacigastridae (sap flies)
* Clusiidae (lekking, or druid flies)
* Fergusoninidae
* Marginidae
* Neminidae
* Neurochaetidae (upside-down flies)
* Odiniidae
* Opomyzidae
* Periscelididae
* Teratomyzidae
* Xenasteiidae

Ephydroidea|

* Camillidae
* Curtonotidae (quasimodo flies)
* Diastatidae (bog flies)
* Drosophilidae (vinegar and fruit flies)
* Ephydridae (shore flies)
* Mormotomyiidae (frightful hairy fly)

Carnoidea|

* Acartophthalmidae
* Australimyzidae
* Braulidae (bee lice)
* Canacidae (beach flies)
* Carnidae
* Chloropidae (frit flies)
* Inbiomyiidae
* Milichiidae (freeloader flies)

Lonchaeoidea|

* Cryptochetidae
* Lonchaeidae (lance flies)

Calyptratae|

| Muscoidea|

* Anthomyiidae (cabbage flies)
* Fanniidae (little house flies)
* Muscidae (house flies, stable flies)
* Scathophagidae (dung flies)

|

Oestroidea|

* Calliphoridae (blow-flies: bluebottles, greenbottles)
* Mesembrinellidae
* Mystacinobiidae (New Zealand batfly)
* Oestridae (botflies)
* Rhiniidae
* Rhinophoridae
* Sarcophagidae (flesh flies)
* Tachinidae (tachina flies)
* Ulurumyiidae

Hippoboscoidea|

* Glossinidae (tsetse flies)
* Hippoboscidae (louse flies)
* Nycteribiidae (bat flies)
* Streblidae (bat flies)

| Stratiomyomorpha|

| Stratiomyoidea|

* Pantophthalmidae (timber flies)
* Stratiomyidae (soldier flies)
* Xylomyidae (wood soldier flies)

|

Tabanomorpha|

| Rhagionoidea|

* Austroleptidae
* Bolbomyiidae
* Rhagionidae (snipe flies)

|

Tabanoidea|

* Athericidae (water snipe flies)
* Oreoleptidae
* Pelecorhynchidae
* Tabanidae (horse and deer flies)

Vermileonomorpha|

| Vermileonoidea|

* Vermileonidae

|

Xylophagomorpha|

| Xylophagoidea|

* Xylophagidae (awl flies)

|

List of families of Diptera

Taxon identifiers|

* Wikidata: Q2233992
* Wikispecies: Heleomyzidae
* BioLib: 17428
* BOLD: 565
* BugGuide: 12760
* EoL: 451
* EPPO: 1HELEF
* Fauna Europaea: 10916
* Fauna Europaea (new): 361a98c1-09e0-4ad7-84de-f66c17f1cac3
* Fossilworks: 139301
* GBIF: 5551
* iNaturalist: 154606
* IRMNG: 101781
* ITIS: 145445
* NBN: NBNSYS0000160133
* NCBI: 219548
* NZOR: ec4d3076-fb44-4a04-be50-5083f256c4c8
* Plazi: 5D446B59-FFAD-FF80-27BD-FD4A2DB5F861
* WoRMS: 150967

|

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