List of organisms named after works of fiction

This is a dynamic list and may never be able to satisfy particular standards for completeness. You can help by adding missing items with reliable sources.

Newly created taxonomic names in biological nomenclature often reflect the discoverer's interests or honour those the discoverer holds in esteem, including fictional elements.[1][2][3]

This is a list of real organisms with scientific names chosen to reference works of fiction.

## Literature[edit]

### Greek mythology[edit]

Taxon | Type | Named for | Notes | Ref

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Antigone antigone (Linnaeus, 1758) | Crane | Antigone of Troy | The species was named after "Antigone, daughter of King Laomedon of Troy, who was metamorphosed into a stork for presuming to compare herself to the goddess Hera. Linnaeus confused this myth with that of Gerana, princess of the pygmies, who was changed into a crane by Hera for committing the same lèse-majesté"  | [4]
Saguinus oedipus (Linnaeus, 1758) | New World monkey | Oedipus Rex | "Linnaeus had a penchant for giving primates names derived from mythology, sometimes with little obvious rationale. So he may have named this one after the mythical tragic King of Thebes, who unknowingly married his own mother."  | [5]
Pseudoeurycea rex (Dunn, 1921) | Salamander | Oedipus Rex | Species formerly named Oedipus rex | [6]
Oedipodrilus oedipus Holt, 1967 | Segmented worm | Oedipus Rex |  | [7]
Amblytylus peitho Linnavuori, 1997 | Leaf bug | Peitho |  | [8]
Atomophora astraia Linnavuori, 1997 | Leaf bug | Dike/Astraea |
Pseudotanais gaiae Jakiel, Palero & Błażewicz, 2019 | Tanaid | Gaia | P. gaiae and P. uranos form a pair of sister species.  | [9]
Pseudotanais uranos Jakiel, Palero & Błażewicz, 2019 | Tanaid | Uranus
†Phyllodrepa daedali Shavrin & Yamamoto, 2019 | Rove beetle | Daedalus | Fossil species found in Eocene Baltic amber.  | [10]
†Phyllodrepa icari Shavrin & Yamamoto, 2019 | Rove beetle | Icarus
Hotwheels sisyphus Liu & Zhang, 2024 | Spider | Sisyphus | "The specific name is derived from Sisyphus, a king in Greek mythology who offended Zeus and whose punishment was to repeatedly roll a huge stone up a hill only to have it roll back down, because the circular copulatory ducts are like Sisyphus’s cyclic mission."  | [11]

### Gargantua and Pantagruel[edit]

Taxon | Type | Named for | Notes | Ref

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Gigantactis gargantua Bertelsen, Pietsch & Lavenberg, 1981 | Anglerfish | Gargantua |  | [12]
†Gargantuavis Buffetaut & Le Loeuff, 1998 | Bird | Gargantua | A genus of fossil flightless birds from the Cretaceous of Europe, and the largest known birds of the Mesozoic era; "generic name from Gargantua, the giant of French folklore made famous by François Rabelais, and avis, Latin for bird"  | [13]
†Notoetayoa gargantuai Gelfo, López & Bond, 2008 | Xenungulate mammal  | Gargantua | A fossil species of hoofed mammal from the Paleocene of Patagonia, Argentina, "Named after the literary character in François Rabelais’ sixteenth century story of two eccentric giants, Gargantua and Pantagruel. In allusion to the larger size of this species compared to [its relative] Etayoa bacatensis."  | [14]
Epimeria gargantua d’Udekem d’Acoz & Verheye, 2017 | Amphipod | Gargantua | "Gargantua is a giant and one of the main characters in the tales of François Rabelais, such as La vie très horrifique du grand Gargantua, père de Pantagruel. The name [...] alludes to the huge size of the species, which is the largest known Epimeria species."  | [15]

### William Shakespeare[edit]

Taxon | Type | Named for | Notes | Ref

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Oberonia Lindley (1859) | Fairy orchid | Oberon, A Midsummer Night's Dream |  | [16]
Sycorax Haliday, 1839 | Fly | Sycorax, The Tempest |  | [17]
Peneothello Mathews, 1920 | Australasian robin | Othello, Othello | Named for being almost (pene) black  | [18]
Pigrogromitus Calman, 1927 | Sea spider | Pigrogromitus, Twelfth Night |  | [19]
Queubus Barnard, 1946 | Sea spider | Queubus, Twelfth Night | "In sooth, we taxonomists are hard put to it to find names, but there have been far worse sources than the nonsense of Will Shakespeare."  | [20]
Iago Compagno & Springer, 1971 | Houndshark | Iago, Othello | "This shark, a namesake of the villain of Shakespeare's Othello, is a troublemaker for systematists and hence a kind of villain."  | [21]
Geocharidius romeoi Erwin, 1982 | Ground beetle | Romeo, Romeo and Juliet | "Romeoi, after Shakespeare's Romeo, who in Act 1 told Benvolio, "He that is stricken blind cannot forget the precious treasure of his eyesight lost," in reference to the demise of the Central American forests and its probable impact on these blind beetles, which are restricted to deep forest humus and litter."  | [22]
Agra othello Erwin, 2000 | Ground beetle | Othello, Othello | The specific name "honors the complicated Shakespearian character whose stage image is a large male singer with a dark complexion. Agra othello is very large, black, and belongs to a formerly complicated species group"  | [23]
Perdita desdemona Portman, 2016 | Bee | Desdemona, Othello | "The name comes from the character in Shakespeare's Othello."  | [24]
Perdita hippolyta Portman, 2016 | Bee | Hippolyta, A Midsummer Night's Dream | "The name comes from the character in Shakespeare's A Midsummer Night's Dream."
Perdita sycorax Portman, 2016 | Bee | Sycorax, The Tempest | "The name comes from the unseen character in Shakespeare's The Tempest."
Perdita titania Portman, 2016 | Bee | Titania, A Midsummer Night's Dream | "The name comes from the fairy queen Titania in Shakespeare's play A Midsummer Night's Dream. The name was chosen due to the beauty and rarity of this species."
†Alfaites romeo Valent, Fatka, & Marek 2019 | Hyolith | Romeo, Romeo and Juliet | "The specific epithet 'romeo' refers to the Shakespeare's protagonist Romeo Montague."  | [25]
Pseudotanais julietae Jakiel, Palero & Błażewicz, 2019 | Tanaid | Juliet, Romeo and Juliet | P. julietae and P. romeo form a pair of sister species.  | [9]
Pseudotanais romeo Jakiel, Palero & Błażewicz, 2019 | Tanaid | Romeo, Romeo and Juliet
Synagelides rosalindae Kanesharatnam & Benjamin, 2020 | Jumping spider | Rosalind, As You Like It | "The species is named after Rosalind Senior, the heroine of the play As You Like It by William Shakespeare. Generally noted for her resilience, quick wit, and beauty."  | [26]
Synagelides orlandoi Kanesharatnam & Benjamin, 2020 | Jumping spider | Orlando, As You Like It | "The species is named after Orlando de Bois, who at first sight falls in love with Rosalind. He is brave, chivalrous, tender, modest, smart, strong, handsome and beloved by all. However, he is unable to express his love for Rosalind, before he leaves to the forest of Arden."

### Don Quixote[edit]

Taxon | Type | Named for | Notes | Ref

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†Hipparion rocinantis Hernández-Pacheco, 1921 | Horse | Rocinante | A fossil species of primitive horse from the Pliocene of La Mancha, Spain, named "in memory of the horse that was the product of the fantasy and genius of our immortal Cervantes. [...] the name I have chosen associates the equine idea with the geographical one [...]"  | [27]
†Dulcineaia manchegana Babin & Gutiérrez-Marco, 1991 | Bivalve | Dulcinea del Toboso | A fossil species of saltwater clam from the Ordovician of La Mancha, Spain.  | [28]
Ardistomis quixotei Pavel Valdés, 2007 | Ground beetle | Don Quixote | "a patronymic, based on the Latinized surname of the fictional Don Quijote, immortalized in the famous novel Don Quijote de la Mancha written by Miguel de Cervantes Saavedra and first published in 1605. This species is named to commemorate the 400th anniversary of publication of this important piece of Spanish literature."  | [29]
†Lohuecotitan pandafilandi Díez Díaz et al., 2016 | Titanosaur | Pandafilando of the Scowl  | "The specific name pandafilandi refers to Pandafilando de la fosca vista, one of the characters in the novel The Ingenious Gentleman Don Quixote of La Mancha (El ingenioso hidalgo don Quijote de la Mancha) written by Miguel de Cervantes and published in the early seventeenth century. Pandafilando is, in the mind of the protagonist, a giant against who he must fight." The character is named "Pandafilando of the Scowl" in John Ormsby's translation.  | [30]
Carex quixotiana Ben.Benítez, Martín-Bravo, Luceño & Jim.Mejías (2023) | Sedge | Don Quixote | "The species epithet, quixotiana (pronounced kee·how·tee·a·na in English) is derived from Miguel de Cervantes's (1547–1616) masterpiece Don Quixote [...], globally considered one of the best works in the history of literature, and whose number of editions and translations is only surpassed by the Bible. The setting of Don Quixote is La Mancha, the region of Spain where almost all populations of Carex quixotiana occur."  | [31]

### Robinson Crusoe[edit]

Taxon | Type | Named for | Notes | Ref

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Robinsonia DC. (1833) | Flowering plant | Robinson Crusoe | This genus is endemic to the Juan Fernández Islands, where Alexander Selkirk was shipwrecked; he was the inspiration for Robinson Crusoe.  | [32]
Hemistomia fridayi Haase & Bouchet, 1998 | Freshwater snail | Friday, Robinson Crusoe | This species was found in the district of Robinson, New Caledonia.  | [33]
Oncopagurus crusoei Lemaitre, 2014 | Hermit crab | Robinson Crusoe | This species is endemic to the Juan Fernández Islands, where Alexander Selkirk was shipwrecked; he was the inspiration for Robinson Crusoe.  | [34]

### Gulliver's Travels[edit]

Taxon | Type | Named for | Notes | Ref

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Salticus lilliputanus Lucas, 1846 | Jumping spider | Lilliput |  | [35]
Holorusia brobdingnagia (Westwood, 1876) | Crane fly | Brobdingnag |  | [36]
Laputa Whitley, 1930 | Filefish | Laputa |  | [37]
Blossfeldia liliputana Werderm. (1937) | Cactus | Lilliput | Smallest cactus known to science.  | [38][39]
†Balnibarbi Fortey, 1974 | Trilobite | Balnibarbi |  | [40]
Dryadella lilliputiana Cogniaux (1978) | Orchid | Lilliput | "Many of the species names reflect their diminutive stature [...] D. lilliputiana to Gulliver's tiny captors"  | [41]
†Laputavis Dyke, 2001 | Swift | Laputa |  | [42]
Chorebus liliputanus Fischer, Tormos, Docavo & Pardo, 2004 | Wasp | Lilliput | "The name refers to the very small size of the species."  | [43]
Viola lilliputana Ballard & Iltis (2012) | Violet | Lilliput | "The specific epithet makes a fanciful reference to the tiny growth form of the new species as 'lilliputian'"  | [44]
Peruphorticus gulliveri Erwin & Zamorano, 2014 | Ground beetle | Lemuel Gulliver | "We so name this species because of its very large size in comparison to its congeners, reminding us of Gulliver's travels on the island of Lilliput."  | [45]
Meoneura lilliputensis Stuke & Freidberg, 2017 | Fly | Lilliput | A tiny fly (length<2 mm) "named after the fictional island Lilliput that was introduced in the novel Gulliver's Travels by Jonathan Swift (1726). Lilliput is inhabited by tiny people, who are about one-twelfth the height of ordinary human beings."  | [46]
Anthobium liliputense Shavrin & Smetana, 2018 | Rove beetle | Lilliput | "The specific epithet is derived from Lilliput, the fictional island with small people that appear in the novel Gulliver's Travels by the famous English writer Jonathan Swift. It refers to the very small size of the species."  | [47]
Pluteus liliputianus E.F. Malysheva & Malysheva (2019) | Mushroom | Lilliput | "The epithet reflects very small, diminutive size of basidiocarps"  | [48]
Hypothenemus liliputianus Bright, 2019 | Bark beetle | Lilliput | "Latinized form of liliputian, referring to the very small size of the female."  | [49]

### Victor Hugo[edit]

Taxon | Type | Named for | Notes | Ref

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Paracalanus quasimodo Bowman, 1971 | Copepod | Quasimodo, The Hunchback of Notre Dame | "The name is derived from the protagonist of Victor Hugo's classic novel, The Hunchback of Notre Dame, and alludes to the distinctive shape of the prosome."  | [50]
Stylaclista quasimodo Early, 1980 | Wasp | Quasimodo, The Hunchback of Notre Dame | "Stylaclista quasimodo can be recognised by its strongly humped scutellum (hence the name), setation of the head and mesosoma, and the scarcity of setae between the epomia."  | [51]
Tetragnatha quasimodo Gillespie, 1992 | Long-jawed orb weaver | Quasimodo, The Hunchback of Notre Dame | "The common name of this species is "Humpback Spiny", because of the prominent mid-dorsal peak of the abdomen. The specific epithet, regarded as a noun in apposition, refers to Victor Hugo's 'Hunchback of Notre Dame'."  | [52]
Agra eponine Erwin, 2000 | Ground beetle | Éponine, Les Misérables | "The specific epithet, eponine, is the name of the unfortunate street urchin in Victor Hugo's Les Miserables, who, in the Broadway version of the story, personified tragic beauty. Such is the state of the tropical forests where these beetles live"  | [23]
Schistura quasimodo Kottelat, 2000 | Stone loach | Quasimodo, The Hunchback of Notre Dame | The body of this species "sometimes [shows] a conspicuous hump".  | [53]
Paradynomene quasimodo McLay & Ng, 2004 | Crab | Quasimodo, The Hunchback of Notre Dame | "The name is derived from Victor Hugo's character, Quasimodo, the Hunchback of Notre Dame; alluding to its tumescent, hunchback carapace shape."  | [54]
Pseudione quasimodo Boyko & Williams, 2004 | Isopod | Quasimodo, The Hunchback of Notre Dame | "The specific name quasimodo is derived from Victor Hugoʼs famous bell-ringer of Notre Dame, most famously portrayed in film by the great Lon Chaney (1883–1930). This appellation is appropriate due to the shape of the female isopod and the bulge it creates in the branchial chamber of the host"  | [55]
Neopantopsalis quasimodo Taylor & Hunt, 2009 | Harvestman | Quasimodo, The Hunchback of Notre Dame |  | [56]
Harpirhynchus quasimodo Bochkov & Mertins, 2010 | Mite | Quasimodo, The Hunchback of Notre Dame | "Anterior part of propodonotal shield bearing hump-like projection..."  | [57]
Apseudes quasimodo Błażewicz-Paszkowycz & Bamber, 2012 | Tanaid | Quasimodo, The Hunchback of Notre Dame | "Named after Quasimodo, a central character from French author Victor Hugo’s 1831 novel Notre Dame de Paris, who also had a distinctive dorsal hump."  | [58]
Chaleponcus quasimodo Enghoff, 2014 | Millipede | Quasimodo, The Hunchback of Notre Dame |  | [59]
Ephemeroporus quasimodo Elmoor-Loureiro, 2014 | Water flea | Quasimodo, The Hunchback of Notre Dame | "clearly differs from all other species of the genus in the high dorsal keel (hunchback)".  | [60]
Temnothorax quasimodo Snelling, Borowiec & Prebus, 2014 | Ant | Quasimodo, The Hunchback of Notre Dame | "named for the Victor Hugo character in his novel The Hunchback of Notre Dame. [...] The short, deep mesosoma with distinctly convex dorsum results in a "hunched" profile that is characteristic for this species".  | [61]
Squalus quasimodo Viana, Carvalho & Gomes, 2016 | Dogfish shark | Quasimodo, The Hunchback of Notre Dame | "Named for the hunchback of Notre-Dame from the 19th Century novel by Victor Hugo, in obvious reference to its most noticeable character" — "body conspicuously robust and humped dorsally".  | [62]
Epimeria quasimodo d'Udekem d'Acoz & Verheye, 2017 | Amphipod | Quasimodo, The Hunchback of Notre Dame | "Quasimodo, the hunchback of Notre-Dame, is a well-known character from the novel Notre-Dame de Paris by Victor Hugo. The name [...] alludes to the humpbacked silhouette of the species."  | [15]
Haplochromis quasimodo Vranken et al., 2022 | Cichlid fish | Quasimodo, The Hunchback of Notre-Dame | "Specific name from Quasimodo, hunchbacked character in Victor Hugo's novel Notre-Dame de Paris (1831); referring to rather shallow head and deep and rhomboid bodies of large specimens."  | [63]

### The Three Musketeers[edit]

Taxon | Type | Named for | Notes | Ref

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Schistura aramis Kottelat, 2000 | Stone loach | Aramis |  | [53]
Schistura athos Kottelat, 2000 | Stone loach | Athos |
Schistura porthos Kottelat, 2000 | Stone loach | Porthos |

### Moby-Dick[edit]

Taxon | Type | Named for | Notes | Ref

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Daggoo Sime & Wahl, 2002 | Wasp | Daggoo | "The genus is named after one of the harpooners in Melville's novel Moby Dick."  | [64]
Queequeg Sime & Wahl, 2002 | Wasp | Queequeg
Tashtego Sime & Wahl, 2002 | Wasp | Tashtego
†Thaleops mobydicki Amati & Westrop, 2004 | Trilobite | Moby Dick | "At times, the senior author's analysis of relationships among illaenid trilobites felt like Ahab's pursuit of the Great White Whale."  | [65]
Voeltzkowia mobydick (Miralles et al, 2012) | Skink | Moby Dick | "The specific epithet refers to Moby Dick, the famous albino sperm whale imagined by Herman Melville (1851), with whom the new species shares several uncommon characteristics, such as the lack of hindlimbs, the presence of flipper-like forelimbs, highly reduced eyes, and the complete absence of pigmentation"  | [66]
†Albicetus oxymycterus Boersma & Pyenson, 2015 | Toothed whale | Moby Dick | "Combining the Latin words albus (white) and cetus (whale). The name pays tribute to H. Melville's classic American novel Moby-Dick; or, The Whale. In the novel, Melville refers to Moby-Dick as “the White Whale”, a creature of “unwonted magnitude” with a “remarkable hue” and “deformed lower jaw”. These traits are coincidentally similar to the type specimen of Albicetus, a white fossil sperm whale whose jaws have been displaced due to diagenetic processes."  | [67]
Vrijenhoekia ahabi Summers, Pleijel & Rouse, 2015 | Segmented worm | Captain Ahab | "Ahabi is in reference to Herman Melville's Captain Ahab, as both he and this species of worm will go to the ends of the earth to find a whale. This name was a winner of a 'Name a Species' public contest organised by the Birch Aquarium, Scripps Institution of Oceanography, submitted by Andy Fyfe."  | [68]

### Lewis Carroll[edit]

Taxon | Type | Named for | Notes | Ref

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†Borogovia gracilicrus Osmólska, 1987 | Theropod dinosaur | Borogove, "Jabberwocky"  | Named after the "borogove — the name of a fantastic creature from "Alice in Wonderland" by Lewis Carroll"  | [69]
†Hemignathus vorpalis James & Olsen, 2003 | Finch | Vorpal blade, "Jabberwocky"  | Named "in reference to the long scimitar-like maxillary rostrum of the bird"  | [70]

### Mark Twain[edit]

Taxon | Type | Named for | Notes | Ref

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Phanuromyia pauper Nesheim and Masner, 2017 | Wasp | The Prince and the Pauper | "The name pauper refers to the lack of longitudinal costae on the base of T2."  | [71]
Phanuromyia princeps Nesheim and Masner, 2017 | Wasp | The Prince and the Pauper | "The name princeps is derived from the prince character in the book The Prince and the Pauper by Mark Twain in reference to its similarity to P. pauper."

### The Adventures of Pinocchio[edit]

Taxon | Type | Named for | Notes | Ref

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Walckenaeria pinocchio (Kaston, 1945) | Spider | Pinocchio | "From all other known species this one can be distinguished by its very long cephalic horn."
Originally described as Cornicularia pinocchio; subsequently, genus Cornicularia was synonymised with Walckenaeria.  | [72][73]
Masdevallia pinocchio Luer & Andreeta, 1978 | Orchid | Pinocchio | "Named for Pinocchio, the long-nosed, wooden puppet that wanted to be a little boy", because of the appearance of the flowers.  | [74]
Oxypleurodon pinocchio (Guinot & Richer de Forges, 1985) | Crab | Pinocchio | "the long and "turned-up" rostrum of this species recall [sic] to mind the long and snub nose of the mischievious [sic] Pinocchio."
Originally described as Sphenocarcinus pinocchio, and subsequently transferred to genus Oxypleurodon.  | [75]
Uroptychus pinocchio Poore & Andreakis, 2011 | Squat lobster | Pinocchio | "For Pinocchio, a wooden puppet that dreamt of becoming a real boy in the 1883 novel Le avventure di Pinocchio by Carlo Collodi. Pinocchio's nose grew longer when he told lies." This species belongs to the Uroptychus naso complex, distinctive for their particularly long and broad rostrum.  | [76]
Eviota pinocchioi Greenfield & Winterbottom, 2012 | Dwarfgoby | Pinocchio | "Named after Carlo Collodi's fictional character Pinocchio, who had a nose that grew long when he lied, alluding to the exceptionally long anterior tubular nares in this species."  | [77]
Anchylorhynchus pinocchio De Madeiros & Nunez-Avellaneda, 2013 | Weevil | Pinocchio | "Named after the popular fictional character Pinocchio, because of the extremely elongate rostrum as compared to other species of the genus."  | [78]
Monodelphis pinocchio Pavan, 2015 | Short-tailed opossum | Pinocchio | "The specific epithet is a noun in apposition and refers to the fictional wooden doll [...], in allusion to the elongated rostrum shared by this species and its namesake."  | [79]
Litoria pinocchio Oliver et al, 2019 | Frog | Pinocchio | Males have a "distinct rostral spike" and is named in "reference to Carlo Collodi's fictional character Pinocchio, who had a nose that became longer when under stress or lying."  | [80]
Pinoquio Carvalho & Huber, 2022 | Spider | Pinocchio | A genus of Brazilian cellar spiders; "The generic name is taken from Carlo Collodi's fictional character, as written in Portuguese, whose famous nose reminds of the projecting clypeus in the type species". This genus was originally named Pinocchio Huber & Carvalho, 2019, but this name turned out to be a junior homonym that had been used before, for the harvestman genus Pinocchio Mello-Leitão, 1940 (now considered a synonym of Bresslauius Mello-Leitão, 1935). Therefore, the genus name was amended to Pinoquio.  | [81][82]

### Arthur Conan Doyle[edit]

Taxon | Type | Named for | Notes | Ref

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†Irritator challengeri Martill, et al., 1996 | Theropod dinosaur | Professor Challenger, The Lost World | Named after "Professor Challenger, the fictitious hero and dinosaur discoverer of Sir Arthur Conan-Doyle's [The] Lost World"  | [83]
Mandevilla sherlockii L.O.Alvarado & Lozada-Pérez (2017) | Rocktrumpet | Sherlock Holmes | "The name is dedicated to the fictional character Sherlock Holmes [who] reflects much of the work of taxonomists and scientists in general, which is entirely detective work. Additionally, he was conceived as an amateur botanist, mainly interested in plants that can be used as poisons"  | [84]

### Rudyard Kipling[edit]

Taxon | Type | Named for | Notes | Ref

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Akela Peckham & Peckham, 1896 | Jumping spider | Akela, The Jungle Book |  | [85]
Bagheera kiplingi Peckham & Peckham, 1896 | Jumping spider | Bagheera and Rudyard Kipling, The Jungle Book |
Messua Peckham & Peckham, 1896 | Jumping spider | Messua, The Jungle Book |
Nagaina Peckham & Peckham, 1896 | Jumping spider | Nagaina, "Rikki-Tikki-Tavi"  |

### Cyrano de Bergerac[edit]

Taxon | Type | Named for | Notes | Ref

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Garra cyrano Kottelat, 2000 | Log sucker fish  | Cyrano de Bergerac, Cyrano de Bergerac (play) | "From Cyrano de Bergerac, the main character in a comedy of Edmond Rostand, characterized by a long nose"; this species has a "snout with a conspicuous, deeply notched secondary rostrum with large tubercles"  | [53]
Eremobates cyranoi Cushing & Brookhart, 2016 | Camel spider | Cyrano de Bergerac, Cyrano de Bergerac (play) | "Referencing the twisted upturned fixed finger reminiscent of the reputed nose of Edmond Rostand's Cyrano de Bergerac."  | [86]
Epimeria cyrano d’Udekem d’Acoz & Verheye, 2017 | Amphipod | Cyrano de Bergerac, Cyrano de Bergerac (play) | "Cyrano de Bergerac is the central character of the eponymous play by Edmond Rostand. In the play, Cyrano is described as endowed with a prominent nose. The name [...] alludes to the long, straight and anteriorly directed rostrum of the species."  | [15]

### Dracula[edit]

Taxon | Type | Named for | Notes | Ref

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†Bradycneme draculae Harrison & Walker, 1975 | Theropod dinosaur | Count Dracula | "The specific name is derived from the Romanian word dracul meaning evil one."

The fossil was discovered in Transylvania, the setting of Bram Stoker's Dracula.

| [87]
Dracula vampira Luer (1978) | Orchid | Count Dracula |  | [88]
Liparis draculoides Greenwood (1982) | Orchid | Count Dracula |  | [89]
†Desmodus draculae Morgan, Linares and Ray, 1988 | Bat | Count Dracula | "The specific epithet of this largest known chiropteran vampire commemorates Count Dracula, the greatest human vampire of folklore"  | [90]
Draculoides bramstokeri Harvey & Humphreys, 1995 | Whip scorpion | Count Dracula, Bram Stoker | "The generic name alludes to another toothed creature"  | [91]
Danionella dracula Britz, Conway & Rüber, 2009 | Ray-finned fish | Count Dracula | "The species name dracula alludes to the long tooth-like fangs in the jaws in males of the new species and was inspired by Count Dracula in Bram Stoker's novel."  | [92]
†Deinocroton draculi Peñalver, et al, 2017 | Tick | Count Dracula | "Patronym for the main character of the gothic horror novel by Irish writer Abraham “Bram” Stoker, which is a fictionalised account of Vlad III, or Vlad Dracula"  | [93]
†Supraserphites draculi Rasnitsyn & Öhm-Kühnle, 2019 | Wasp | Count Dracula | "Species name derived from dracul, Romanian for devil, and the literary figure Count Dracula"  | [94]
Patrera dracula Martínez, Brescovit, Villarreal & Oliveira, 2021 | Spider | Count Dracula | "The epithet in apposition referring to the mythological vampire Dracula, alluding to the large chelicerae, which resemble Dracula's fangs."  | [95]

### Peter Pan[edit]

Taxon | Type | Named for | Notes | Ref

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Tinkerbella nana Huber & Noyes, 2013 | Fairyfly | Tinker Bell and Nana  | "After the fairy Tinker Bell in the 1904 play 'Peter Pan' by J.M. Barrie [and] the dog Nana in Peter Pan and coincidentally from nanos, the Greek word for dwarf."  | [96]
Ischnothyreus hooki Kranz-Baltensperger, 2011 | Goblin spider | Captain Hook | "The specific name is derived from Captain Hook from the novel Peter Pan, written by James M. Barrie, and refers to the similarity of the distal part of the male palp with the left arm of Captain Hook."  | [97]
Meoneura tinkerbellae Stucke, 2016 | Fly | Tinker Bell | "Tinker Bell is a long haired and very small fairy as Meoneura tinkerbellae is a very small, hardly visible fly with characteristically long setulae on tergite 5."  | [98]

### H. P. Lovecraft[edit]

Taxon | Type | Named for | Notes | Ref

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Pimoa cthulhu Hormiga, 1994 | Spider | Cthulhu | "Named after H.P. Lovecraft's mythological deity Cthulhu, akin to the powers of Chaos"  | [99]
†Millerocaulis tekelili Vera (2012) | Fern | "Tekeli-li"  | "The specific epithet derives from the word Tekeli-li, pronounced by fictional Antarctic inhabitants in the book 'The Narrative of Arthur Gordon Pym of Nantucket' by Edgar [Allan] Poe and in the book 'At the [M]ountains of [M]adness' by Howard Phillip Lovecraft.  | [100]
Nanocthulhu lovecrafti Buffington, 2012 | Wasp | Cthulhu, H. P. Lovecraft | "Cthulhu's [sic] is described as having 'a pulpy, tentacled head,' and the clypeal fuscina described herein is reminiscent of Cthulhu's head"  | [101]
Cthulhu macrofasciculumque James & Keeling, 2012 | Parabasalid | Cthulhu | "The tentacle-headed appearance given by the coordinated beat pattern of the anterior flagellar bundle of Cthulhu cells is reminiscent of this demon"  | [102]
Cthylla microfasciculumque James & Keeling, 2012 | Parabasalid | Cthylla | Cthylla "was the secret daughter of Cthulhu [...] It is here named as a smaller and simpler relative of the parabasalian genus Cthulhu"
Ochyrocera atlachnacha Brescovit et al., 2018 | Spider | Atlach-Nacha | The specific name "refers to Atlach-Nacha, a supernatural entity from Cthulhu mythology that resembles a huge spider with an almost human face"  | [103]
Yogsothoth Shishkin & Zlatogursky, 2018 | Centrohelid | Yog-Sothoth | Yog-Sothoth "was described as a conglomeration of glowing spheres".

The genus is in the family Yogsothothidae.

| [104]
†Sollasina cthulhu Rahman et al., 2019 | Echinoderm | Cthulhu mythos | "Named for the Cthulhu mythos of H. P. Lovecraft, a fictional universe populated with bizarre tentacled monsters"  | [105]
Fujuriphyes dagon Cepeda, Pardos, Zeppilli & Sánchez, 2020 | Mud dragon | Dagon (Cthulhu Mythos) | "The species is named after the fictional deity Dagon (also known as Father Dagon), created by the American writer of horror fiction H.P. Lovecraft (1890–1937) and firstly introduced in the short story "Dagon," published in 1919. In the pantheon of Lovecraftian cosmic entities, Dagon presides over the Deep Ones, an amphibious humanoid race indigenous to Earth's oceans."  | [106]
Fujuriphyes hydra Cepeda, Pardos, Zeppilli & Sánchez, 2020 | Mud dragon | Hydra (Cthulhu Mythos) | "The species is named after the fictional deity Hydra (also known as Mother Hydra), created by the American writer of cosmic horror fiction H.P. Lovecraft (1890–1937) and firstly introduced in the short story The Shadow over Innsmouth, published in 1936. In the pantheon of Lovecraftian cosmic entities, Mother Hydra is the consort of Father Dagon."
Fujuriphyes cthulhu Cepeda, Pardos, Zeppilli & Sánchez, 2020 | Mud dragon | Cthulhu | "The species is named after the fictional cosmic entity Cthulhu, created by the American writer of horror fiction H.P. Lovecraft (1890–1937) and firstly introduced in the short story "The Call of Cthulhu," published in 1928. Considered a Great Old One within the pantheon of Lovecraftian cosmic entities, Cthulhu is a gigantic being of great power described as looking like an octopus or a dragon that lies in a death-like torpor in the sunken city of R'lyeh."

### Winnie-the-Pooh[edit]

Taxon | Type | Named for | Notes | Ref

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Eeyorius hutchinsi Paulin, 1986 | Ray-finned fish | Eeyore | "Named for Eeyore, a literary character who lived in damp places."  | [107]
Potamalpheops tigger Yeo & Ng, 1997 | Shrimp | Tigger | Named after Tigger to reference "the bold striped appearance of freshly caught live specimens".  | [108]

### Vladimir Nabokov[edit]

Taxon | Type | Named for | Notes | Ref

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Pseudolucia aureliana Bálint & Johnson, 1993 | Butterfly | "The Aurelian"  | "Named from Nabokov's short story 'The Aurelian' in which a devoted butterfly collector is unable to fulfill his dream of journeying to the tropics."  | [109]
Pseudolucia charlotte Bálint & Johnson, 1993 | Butterfly | Charlotte Haze, Lolita | "Named for 'Charlotte Haze' Lolita's mother in Nabokov's novel Lolita."
Pseudolucia clarea Bálint & Johnson, 1993 | Butterfly | Charlotte Haze, Lolita | "Named for 'Clare Quilty', the rival of Lolita's paramour 'Humbert' in Nabokov's novel Lolita."
Pseudolucia hazeorum Bálint & Johnson, 1993 | Butterfly | Haze family, Lolita | "A double meaning, the denotative of 'haze' referring to the dark wing bands characterizing the species, but signifying also Nabokov's characters of the Haze family in Lolita\-- the infamous Lolita and her mother Charlotte."
Pseudolucia zembla Bálint & Johnson, 1993 | Butterfly | Zembla, Pale Fire | "Named for the mythical kingdom 'Zembla' claimed to have been ruled by Nabokov's character Kinbote in the novel Pale Fire."
Itylos pnin Bálint, 1993 | Butterfly | Timofey Pavlovich Pnin, Pnin | The species is "named for the Russian emigre professor in Nabokov's novel Pnin."  | [110]
Madeleinea lolita Bálint, 1993 | Butterfly | Lolita | "Named for 'Lolita', the nickname of the Nabokov's best-known character— the pre-teen nymphet Dolores in the well-known novel Lolita."
Madeleinea mashenka Bálint, 1993 | Butterfly | Mashen'ka, the Russian title of Mary | "'Mashenka' (Mary) was the title of the first novel published by Nabokov in Russian."
Paralycaeides shade Bálint, 1993 | Butterfly | John Shade, Pale Fire | The species is "named for 'John Shade', the imaginary New England poet and author of Nabokov's Pale Fire."
Nabokovia ada Bálint & Johnson, 1994 | Butterfly | Ada Veen, Ada or Ardor: A Family Chronicle |  | [111]
Madeleinea cobaltana Bálint & Lamas, 1994 | Butterfly | Kobaltana, Pale Fire |  | [112]
Madeleinea nodo Bálint & Johnson, 1994 | Butterfly | Nodo, Pale Fire | "Named for 'Nodo', half-brother of Odon, signifying the sister relationship with M. odon and the fact that, of the two, 'Nodo' occurs geographically to the [no]rth."  | [113]
Madeleinea odon Bálint & Johnson, 1994 | Butterfly | Odon, Pale Fire | "Named for 'Odon', half brother of Nodo and further suggesting the interesting diversity of the koa Species Group."
Madeleinea tintarrona Bálint & Johnson, 1994 | Butterfly | Tintarron, Pale Fire | "From Nabokov's 'Tintarron', a precious deep blue glass made in the mountains of Zembla, here referring to the dorsal ground color of this species"
Leptotes krug Bálint et al., 1995 | Butterfly | Adam Krug, Bend Sinister | Named for "the 'unfortunate professor' Krug in Vladimir Nabokov's novel Bend Sinister [...] 'krug' also means 'circle' in Russian. [The] ventral maculation in this genus includes circles or ellipses of white which enclose patches of brown ground color."  | [114]
Leptotes delalande Bálint et al., 1995 | Butterfly | Pierre Delalande, Invitation to a Beheading | "Named for 'Delalande', a philosopher invented by Nabokov as his alter-ego, here reflecting the sister relationship of this new species with L. lamasi."
Pseudolucia humbert Bálint & Johnson, 1995 | Butterfly | "Humbert Humbert", Lolita | "Named from Humbert, a central character in Nabokov's novel Lolita. We refrain here from adding the genitive 'i' since the character was imaginary and obviously masculine."  | [115]
Pseudolucia zina Benyamini, Bálint & Johnson, 1995 | Butterfly | Zina Mertz, The Gift | "Zina is perhaps the closest Nabokov came to placing his own wife, Vera, into his fictions. In brief, Zina is the most ideal of all Nabokov's female characters."  | [116]
Patalycaeides hazelea Bálint and Johnson, 1995 | Butterfly | Hazel Shade, Pale Fire | "[T]he name is after Shade's daughter Hazel, in Nabokov's Pale Fire, whose suicide is at [the] center of Shade's poem and who turns up, after her death and just before Shade's, in the form of a butterfly."  | [117]
Madeleinea ardisensis Bálint & Lamas, 1996 | Butterfly | Ardis Hall, Ada or Ardor: A Family Chronicle |  | [118]
Plebejus ardis Bálint & Johnson, 1997 | Butterfly | Ardis Hall, Ada or Ardor: A Family Chronicle | "Dr. Proffer notes that 'Ardis' was the name of the great estate ('Ardis Hall') in Nabokov's novel Ada, where Van and Ada (Nabokov's only lepidopterist heroine) began their life-long love affair."  | [119]
Plebejus pilgram Bálint & Johnson, 1997 | Butterfly | Paul Pilgram, "The Aurelian"  | Dr. Simon Karlinsk: "'Pilgram' was the protagonist in Nabokov's story 'The Aurelian' The German owner of a butterfly shop, he dreamed all his life of hunting butterflies in exotic locales but, as he was about to realize his dreams, died of a heart attack."
Plebejus pilgram Bálint & Lamas, 1998 | Butterfly | Mira Belochkin, Pnin | "Gayla Diment [...] proposed 'Mira', the first name of Pnin's beloved who died in a Nazi camp. It is fitting that a butterfly, often representing the human soul, should bear her name."  | [120]
Plebejus fyodor Hsu, Bálint & Johnson, 2000 | Butterfly | Fyodor Godunov-Cherdyntsev, The Gift | "Fyodor narrates Nabokov's novel The Gift, a record of his love of Russian literature, his lepidopterist father, butterflies, and a young woman named Zina."  | [121]
Humbert humberti Sime & Wahl, 2002 | Wasp | "Humbert Humbert", Lolita | "The genus is after the hapless Humbert Humbert of Nabokov's novel, Lolita"  | [64]

### J. R. R. Tolkien[edit]

Main article: List of things named after J. R. R. Tolkien and his works § Taxonomy

### Enid Blyton[edit]

Taxon | Type | Named for | Notes | Ref

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Cavisternum bom Ranasinghe & Benjamin, 2018 | Goblin spider | Bom, "The Goblins Looking-Glass"  | "This specific name is a noun in apposition named after 'Bom' a magnificent goblin in the story of 'The Goblins Looking-Glass' by Enid Blyton (1947)."  | [122]
Pelicinus tumpy Ranasinghe & Benjamin, 2018 | Goblin spider | Tumpy, "The Goblins Looking-Glass"  | "This species is a noun in apposition named after 'Tumpy' a little goblin in the story, The Goblins Looking-Glass by Blyton (1947)."
Pelicinus snooky Ranasinghe & Benjamin, 2018 | Goblin spider | Snooky, "The Firework Goblins"  | "This species a noun in apposition named after 'Snooky' a goblin and a main character in the story 'The Firework Goblins' written by Blyton in 1971."
Ischnothyreus chippy Ranasinghe & Benjamin, 2018 | Goblin spider | Chippy, "Billy's Little Boats"  | "This species name is a noun in apposition named after 'Chippy' a brownie and one of the characters in the story 'Billy's Little Boats' by Blyton (1971). Brownies possess queer little pointed feet like goblins."
Silhouettella snippy Ranasinghe & Benjamin, 2018 | Goblin spider | Snippy, "Billy's Little Boats"  | "This species is a noun in apposition named after 'Snippy' a brownie and one of the characters in the story 'Billy's Little Boats' by Blyton in 1971."
Silhouettella tiggy Ranasinghe & Benjamin, 2018 | Goblin spider | Tiggy, "Billy's Little Boats"  | "This species is a noun in apposition named after “Tiggy” a brownie and one of the characters in the story 'Billy's Little Boats' by Blyton in 1971."

### Dune[edit]

Taxon | Type | Named for | Notes | Ref

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Synothele arrakis Raven, 1994 | Brushed trapdoor spider | Arrakis | "A noun taken from the Frank Herbert novel, Dune, in which the desert planet is known as Arrakis. Sands of the desert planet yield a 'spice' which changes the eye colour in users to deep blue. The anterior median eyes of the types have a deep blue colour, unusual in spider eyes."  | [123]
Lycosa gesserit Steinpress et al., 2022 | Wolf spider | Bene Gesserit | "Species named after the order of Bene Gesserit, from Frank Herbert's 1965 novel, Dune."  | [124]
Shaihuludia Kimmig et al., 2023 | Polychaete | Sandworms | "Shai-Hulud: the indigenous name for the sandworms on the planet Arrakis in the Dune novels written by Frank Herbert."  | [125]
Tulostoma shaihuludii Finy et al., 2023 | Fungus | Sandworms | "Reminiscent of the sandworm Shai-Hulud of the fictional planet Arrakis from the science fiction novel series Dune by Frank Herbert."  | [126]
Arrakis Robillard, Tan & Su, 2024 | Cricket | Arrakis | "The genus is named after the fictional desert planet featured in the Dune series of novels by Frank Herbert, in reference to the desert regions where the members of this genus are distributed."  | [127]

### Aubrey–Maturin series[edit]

Taxon | Type | Named for | Notes | Ref

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Ledermanniella maturiniana Beentje (2005) | Riverweed | Stephen Maturin |  | [128]
Theognete maturini Anderson, 2010 | Weevil | Stephen Maturin | "The dedication was arranged through a donation to Nature Discovery Fund of the Canadian Museum of Nature by The Gunroom of HMSSurprise.org."  | [129][130]

### The Hitchhiker's Guide to the Galaxy[edit]

Taxon | Type | Named for | Notes | Ref

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Erechthias beeblebroxi Robinson & Nelson, 1993 | Moth | Zaphod Beeblebrox |  | [131]
Bidenichthys beeblebroxi Paulin, 1995 | Viviparous brotula fish | Zaphod Beeblebrox | "Named for Zaphod Beeblebrox, President of the Galaxy in The [Hitchhiker's] Guide to the Galaxy by Douglas Adams."  | [132]
Fiordichthys slartibartfasti Paulin, 1995 | Viviparous brotula fish | Slartibartfast | "Named for Slartibartfast, designer of fiords in The [Hitchhiker's] Guide to the Galaxy by Douglas Adams."
†Babelichthys Davesne, 2017 | Crestfish | Babelfish  | Named after the "teleost-like, ear-dwelling, polyglot" Babel Fish "in reference to the very peculiar, almost alien-like, appearance of the genus".  | [133]

### Discworld[edit]

Taxon | Type | Named for | Notes | Ref

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†Czekanowskia anguae Watson et al., 2001 | Gymnosperm | Angua von Überwald | "In tribute to the author Terry Pratchett OBE, all the new fossil plant species diagnosed and described in this paper are named for fictional characters who appear in his series of Discworld novels."  | [134]
†Ginkgoites weatherwaxiae Watson et al., 2001 | Ginkgo | Granny Weatherwax
†Ginkgoites nannyoggiae Watson et al., 2001 | Ginkgo | Nanny Ogg
†Ginkgoites garlickianus Watson et al., 2001 | Ginkgo | Magrat Garlick
†Phoenicopsis rincewindii Watson et al., 2001 | Gymnosperm | Rincewind
†Pseudotorellia vimesiana Watson et al., 2001 | Conifer | Sam Vimes
†Sciadopityoides greeboana Watson et al., 2001 | Gymnosperm | Greebo, Nanny Ogg's cat
†Sulcatocladus dibbleri Watson et al., 2001 | Conifer | C.M.O.T. Dibbler
†Torreyites detriti Watson et al., 2001 | Conifer | Detritus
Apseudes atuini Bamber, 2005 | Crustacean | Great A'Tuin |  | [135]
Bathytanais greebo Bamber, 2005 | Crustacean | Greebo, Nanny Ogg's cat  |  | [136]
Tanaella dongo Bamber, 2005 | Crustacean | Crocodile Dongo  |  | [137]
Aleiodes achingae Butcher et al., 2012 | Wasp | Tiffany Aching |  | [138]
Aleiodes adorabelleae Butcher et al., 2012 | Wasp | Adora Belle Dearheart |
Aleiodes anguaae Butcher et al., 2012 | Wasp | Angua von Überwald |
Aleiodes atuin Butcher et al., 2012 | Wasp | Great A'Tuin |
Aleiodes binkyi Butcher et al., 2012 | Wasp | Binky, Death's horse  |
Aleiodes conina Butcher et al., 2012 | Wasp | Conina |
Aleiodes deathi Butcher et al., 2012 | Wasp | Edward d'Eath |
Aleiodes deyoyoi Butcher et al., 2012 | Wasp | Kompt de Yoyo  |
Aleiodes downeyi Butcher et al., 2012 | Wasp | Lord Downey |
Aleiodes flannelfooti Butcher et al., 2012 | Wasp | Zlorf Flannelfoot  | "Named after the assassin character Zlorf Flannelfoot in the Terry Pratchett Discworld novel and in allusion to the pectinate claws."
Aleiodes gaspodei Butcher et al., 2012 | Wasp | Gaspode the Wonder Dog  |
Aleiodes herrena Butcher et al., 2012 | Wasp | Herrena |
Aleiodes lavaeolous Butcher et al., 2012 | Wasp | Lavaeolous |
Aleiodes liessa Butcher et al., 2012 | Wasp | Liessa Wyrmbidder  |
Aleiodes lipwigi Butcher et al., 2012 | Wasp | Moist von Lipwig |
Aleiodes lipwigduplicitus Butcher et al., 2012 | Wasp | Moist von Lipwig | "In reference to the duplicitous nature of the character Mr Lipwig in the Terry Pratchett Discworld novel Going Postal and hence it's easy confusion with A. lipwigi"
Aleiodes magratae Butcher et al., 2012 | Wasp | Magrat Garlick |
Aleiodes malichi Butcher et al., 2012 | Wasp | Alberto Malich |
Aleiodes mericeti Butcher et al., 2012 | Wasp | Mericet  |
Aleiodes morti Butcher et al., 2012 | Wasp | Mort |
Aleiodes nivori Butcher et al., 2012 | Wasp | Grunworth Nivor  |
Aleiodes ponderi Butcher et al., 2012 | Wasp | Ponder Stibbons | "Named after the character Ponder Stibbons from the Terry Pratchett Discworld novels, and a pun on 'to ponder'."
Aleiodes prillae Butcher et al., 2012 | Wasp | Lady Prill  |
Aleiodes pteppicymoni Butcher et al., 2012 | Wasp | Pteppicymon XXVIII | "Named after the assassin character 'His Greatness the King Pteppicymon XXVIII, Lord of the Heavens ...' from the Terry Pratchett Discworld novel, in reference to lethal parasitoid biology of the wasp."
Aleiodes ptraci Butcher et al., 2012 | Wasp | Ptraci I |
Aleiodes ridcullyi Butcher et al., 2012 | Wasp | Mustrum Ridcully |
Aleiodes rincewindi Butcher et al., 2012 | Wasp | Rincewind |
Aleiodes sacharissa Butcher et al., 2012 | Wasp | Sacharissa Cripslock |
Aleiodes selachiii Butcher et al., 2012 | Wasp | Robert Selachii  |
Aleiodes stibbonsi Butcher et al., 2012 | Wasp | Ponder Stibbons |
Aleiodes stohelit Butcher et al., 2012 | Wasp | Susan Sto-Helit |
Aleiodes teatimei Butcher et al., 2012 | Wasp | Jonathan Teatime |
Aleiodes tmaliaae Butcher et al., 2012 | Wasp | T'malia  |
Aleiodes vetinarii Butcher et al., 2012 | Wasp | Havelock Vetinari |
Periclimenes rincewindi De Grave, 2014 | Shrimp | Rincewind | "Rincewind is a fictional wizard in several Discworld novels by Terry Pratchett, who is well known for his ability to blend in with any situation, despite his penchant for colourful clothing."  | [139]
Dimophora rhysi Klopfstein, 2016 | Wasp | Rhys Rhysson | "This particularly small Dimophora species is named after the Low King of the dwarfs, Rhys Rhysson, from the late Terry Pratchett's novel 'The Fifth Elephant', and is dedicated to its inventor and father of the Discworld series. He will be sorely missed."  | [140]
Pristomerus fourecksensis Klopfstein, 2016 | Wasp | Fourecks | "This name refers to the continent 'Fourecks' in the discworld novel by the late Terry Pratchett. Fourecks has many similarities with Australia, and the name is most probably based on the XXXX beer from Queensland."  | [141]

### The Witcher[edit]

Taxon | Type | Named for | Notes | Ref

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Pseudotanais geralti Jakiel, Palero & Błażewicz, 2019 | Tanaid | Geralt of Rivia | P. geralti and P. yenneferae form a pair of sister species.  | [9]
Pseudotanais yenneferae Jakiel, Palero & Błażewicz, 2019 | Tanaid | Yennefer of Vengerberg

### A Song of Ice and Fire[edit]

Taxon | Type | Named for | Notes | Ref

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Laelius arryni Azevedo & Barbosa, 2014 | Wasp | House Arryn | "The specific epithets of all new species are derived from some families of the book 'A Song of Ice and Fire, [A] Game of Thrones'" by George R. R. Martin | [142]
Laelius baratheoni Azevedo & Barbosa, 2014 | Wasp | House Baratheon
Laelius lannisteri Azevedo & Barbosa, 2014 | Wasp | House Lannister
Laelius martelli Azevedo & Barbosa, 2014 | Wasp | House Martell
Laelius targaryeni Azevedo & Barbosa, 2014 | Wasp | House Targaryen
Laelius tullyi Azevedo & Barbosa, 2014 | Wasp | House Tully
Laelius starki Azevedo & Barbosa, 2014 | Wasp | House Stark
Marianina khaleesi (Silva, Azevedo & Matthews-Cascon, 2014) | Sea slug | Khaleesi, title of Daenerys Targaryen | Daenerys "is described as being short and having long white hair, features that resemble the white band on the notum" of the species  | [143]
Pheidole drogon Sarnat, Fischer & Economo, 2016 | Ant | Drogon | "The species name refers to Drogon, the black-colored dragon of Daenerys Targaryen"  | [144]
Pheidole viserion Sarnat, Fischer & Economo, 2016 | Ant | Viserion | "The species name refers to Viserion, the cream and gold colored dragon of Daenerys Targaryen"
Eadya daenerys Ridenbaugh, 2018 | Wasp | Daenerys Targaryen | "This species is named for Daenerys Stormborn of House Targaryen, the First of Her Name, Queen of the Andals and the First Men, Protector of the Seven Kingdoms, the Mother of Dragons, Khaleesi of the Great Grass Sea, the Unburnt, the Breaker of Chains"  | [145]
Ochyrocera varys Brescovit et al., 2018 | Spider | Varys | "Lorde Varys is a character with a venomous spirit, known as a spider in the plot."  | [103]
Paramonovius nightking Li & Yeates, 2018 | Bee fly | Night King | "This species is named after the Night King in the American fantasy drama Game of Thrones, because all the specimens were collected in winter and the fly is mostly covered in thick pale pruinescence." Also, the fly, like the fictional villain, turns its victims into zombies.  | [146]
Abyssarya Bonifácio & Menot, 2018 | Segmented worm | Arya Stark | "This genus is dedicated to Arya Stark, one of P.B.’s favourite characters in the novel 'A song of ice and fire' by George R. R. Martin. The name is composed by 'abyss' from the Latin word 'ăbyssus' meaning 'bottomless' and Arya."  | [147]
Hodor Bonifácio & Menot, 2018 | Segmented worm | Hodor | "This genus is dedicated to Hodor, one of P.B.’s favourite characters in the novel 'A song of ice and fire' by George R. R. Martin."
Fujuriphyes viserioni Sánchez, Sørensen & Landers, 2019 | Mud dragon | Viserion |  | [148]
Gymnetis drogoni Ratcliffe, 2019 | Scarab beetle | Drogon |  | [149]
Gymnetis rhaegali Ratcliffe, 2019 | Scarab beetle | Rhaegal |
Gymnetis viserioni Ratcliffe, 2019 | Scarab beetle | Viserion
†Targaryendraco Pêgas, Holgado & Leal, 2019 | Pterosaur | Dragons of House Targaryen | "A combination of Targaryen and draco (from the Latin word for dragon), in reference to the fictional dragons of the saga A Song of Ice and Fire that exhibit dark-coloured bones. This refers to the dark colour of the type specimen of Targaryendraco wiedenrothi comb. nov., as well as to the classic association between pterosaurs and dragons [...]. This reference also honours the fact that pterosaurs have inspired some biological aspects of the dragons featured in the novels."  | [150]
†Meraxes Canale et al., 2022 | Dinosaur | Meraxes  | Named after a dragon ridden by Queen Rhaenys Targaryen.  | [151]
Lannisterella Gellert, Palero & Błażewicz, 2022 | Tanaid | House Lannister |  | [152]
Lannisterella cerseiae Gellert, Palero & Błażewicz, 2022 | Tanaid | Cersei Lannister |

### Harry Potter[edit]

Main article: List of organisms named after the Harry Potter series

### Rumo and His Miraculous Adventure[edit]

Taxon | Type | Named for | Notes | Ref

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Tetramorium rumo Hita Garcia & Fisher, 2014 | Ant | Rumo  | "The new species is named after the fictional character 'Rumo' from Walter Moers' fantasy novel 'Rumo and His Miraculous Adventures'. Tetramorium rumo is a very bright species, almost white, with distinct propodeal spines reminiscent of 'Rumo', who is a white wolperting with short but acute horns."  | [153]
Tetramorium rala Hita Garcia & Fisher, 2014 | Ant | Rala  | "The new species is named after the fictional character 'Rala' from Walter Moers' fantasy novel 'Rumo and His Miraculous Adventures'."

### Ready Player One[edit]

Taxon | Type | Named for | Notes | Ref

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Strumigenys anorak Sarnat et al, 2019 | Ant | Anorak  | "Anorak refers to the online avatar of James Donovan Halliday, creator of the virtual reality world OASIS, in the fictional work Ready Player One"  | [154]
Strumigenys artemis Sarnat et al, 2019 | Ant | Art3mis  | "Artemis is the Latinized version of Art3mis, a famous gunter in the fictional work Ready Player One. Artemis is also the Greek goddess of the hunt and is an apt name for a Strumigenys species."
Strumigenys avatar Sarnat et al, 2019 | Ant | Avatar | "Avatar refers to the three-dimensional representation of OASIS users in the fictional work Ready Player One"
Strumigenys gunter Sarnat et al, 2019 | Ant | Gunter  | "Gunter is a contraction of ‘egg’ and ‘hunter’ in the fictional work Ready Player One and refers to particular users of the virtual reality world OASIS."
Strumigenys oasis Sarnat et al, 2019 | Ant | OASIS  | "Oasis (Ontologically Anthropocentric Sensory Immersive Simulation) refers to the virtual reality world featured in the fictional work Ready Player One"
Strumigenys parzival Sarnat et al, 2019 | Ant | Parzival  | "Parzival refers to the name of the protagonist's virtual reality avatar in the fictional work Ready Player One"

### Other literature[edit]

Taxon | Type | Named for | Notes | Ref

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†Ophiodon ozymandias (Jordan, 1907) | Ray-finned fish | "Ozymandias", Percy Bysshe Shelley | "The generic name refers to the heroic giant noticed by Shelley, of whom nothing is known save the feet of his gigantic statue in the desert."

The species has been renamed with Ozymandias as the specific name.

| [155]
Paramphientomum yumyum Enderlein, 1907 | Barklouse | Yum-Yum, The Mikado | The species is native to Japan, the setting of The Mikado.  | [156]
Grendelius McGowan, 1976 | Ichthyosaur | Grendel, Beowulf | Named after "Grendel [...] a legendary creature from the Old English epic, Beowulf"  | [157]
Morlockia Garcia-Valdecasas, 1984 | Crustacean | Morlocks, The Time Machine | "H. J. Wells described in 'The Time Machine' the Morlocks as inhabitants of the interior of the earth. They had lost their eyes and showed other adaptations to subterranean life."  | [158]
Agra ichabod Erwin, 2002 | Ground beetle | Ichabod Crane, "The Legend of Sleepy Hollow"  | "The specific epithet, ichabod [...] refers to the fact that the Holotype is missing its head and the illusion is that of the frightened Schoolteacher Ichabod Crane's phantom nemesis, the Headless Horseman, in The Legend of Sleepy Hollow [by] Washington Irving, published in 1819"  | [159]
Micropholcus evaluna (Huber, Pérez & Baptista, 2005) | Cellar spider | Eva Luna, Eva Luna and The Stories of Eva Luna | A Venezuelan species "Named for Isabel Allende's Venezuela-born protagonist and fantastic story-teller." Originally described as Leptopholcus evaluna, and subsequently transferred to Micropholcus.  | [160][161]
Ituglanis macunaima Datovo & Landim, 2005 | Catfish | Macunaíma | "From the modernist Brazilian masterpiece by Mário de Andrade – "Macunaíma: o herói sem nenhum caráter" – meaning the hero without any character, in reference of the absence of any exclusive (taxonomic) character for the new species. Mário de Andrade's Macunaíma was based in folk Amazonian indian myth, and also presents infantile features, in allusion to the paedomorphic characters of the new species."  | [162]
Boccacciomymar decameron Triapitsyn & Berezovskiy, 2007 | Fairyfly | The Decameron |  | [163]
"Candidatus Desulforudis audaxviator" Chivian et al., 2008 | Bacteria | "Audax viator", Journey to the Center of the Earth | "[I]n Jules Verne's Journey to the Center of the Earth, in a message [...] deciphered by Verne's protagonist, Professor Lidenbrock, which reads in part, 'descende, Audax viator, et terrestre centrum attinges.' It means 'descend, Bold traveler, and attain the center of the Earth.'"  | [164][165]
Wukongopterus lii Wang et al., 2009 | Pterosaur | Sun Wukong, Journey to the West | "Wukongopterus, from Sun Wukong (the Monkey King), one of the most famous and beloved fictional characters of the classical Chinese literature "Journey to the West", and pterus from the Greek meaning wing."  | [166]
Calumma tarzan Gehring et al, 2010 | Chameleon | Tarzan | "[W]e dedicate the new species to the fictional forest man 'Tarzan' in the hope that this famous name will promote awareness and conservation activities for this apparently highly threatened new species and its habitats, in the mid-altitude rainforest."  | [167]
Aleiodes dresdeni Butcher et al, 2012 | Wasp | Harry Dresden, The Dresden Files | "Named after the wizard character, Harry Dresden, from the Jim Butcher novels."  | [138]
Auplopus charlesi Waichert & Pitts, 2012 | Wasp | Nick Charles, The Thin Man | "Named in honor of Samuel Dashiell Hammett (1894–1961), who was a well-known American author of hardboiled detective novels and short stories, and creator of the famous protagonist, Nick Charles."  | [168]
Dipogon marlowei Waichert & Pitts, 2012 | Wasp | Philip Marlowe | "Named in honor of Raymond Thornton Chandler (1888–1959), an American crime writer who greatly influenced the modern private eye story and created the famous protagonist, Philip Marlowe."
Priocnessus vancei Waichert & Pitts, 2012 | Wasp | Philo Vance | "Named in honor of Willard Huntington Wright (1888–1939), an American crime writer who created the fictional detective, Philo Vance."
Inconnivus billibunteri Błażewicz-Paszkowycz & Bamber, 2012 | Tanaid | Billy Bunter | This species, "much less slender" than its closest relatives, is "named after William George ("Billy") Bunter, a proportionately-fat schoolboy character in books written by Charles Hamilton using the pen-name Frank Richards."  | [58]
Oileus gasparilomi Cano & Schuster, 2012 | Bess beetle | Gaspar Ilóm, Men of Maize | "Named after Gaspar Ilóm, a native hero of the novel 'Men of Maize' by Miguel Ángel Asturias. The collection locality is called 'mountains of Ilóm'."  | [169]
†Millerocaulis tekelili Vera (2012) | Fern | "Tekeli-li", The Narrative of Arthur Gordon Pym of Nantucket | "The specific epithet derives from the word Tekeli-li, pronounced by fictional Antarctic inhabitants in the book 'The Narrative of Arthur Gordon Pym of Nantucket' by Edgar [Allan] Poe and in the book 'At the [M]ountains of [M]adness' by Howard Phillip Lovecraft.  | [100]
Magelona sinbadi Mortimer, Cassà, Martin & Gil, 2012 | Segmented worm | Sinbad the Sailor | "From the name Sinbad, the fictional sailor with Persian origins, referring to the region in which this species was first sampled."  | [170]
Ophryotrocha langstrumpae Wiklund et al., 2012 | Polychaete worm | Pippi Longstocking, Pippi Longstocking (novel) | "Ophryotrocha langstrumpae sp. nov. has long antennae and palps resembling braids, and it is named after a famous braid-bearing girl, Pippi Longstocking, in children's books by Astrid Lindgren. In the original books in Swedish, she is called Pippi Långstrump."  | [171]
Calochaete cimrmanii Hauer, Bohunická & Mühlsteinová, 2013 | Cyanobacteria | Jára Cimrman | "The species is named after the mythical Czech scientist, dramatist, poet, and writer Jára Cimrman."  | [172]
Simlops jamesbondi Bonaldo et al, 2014 | Spider | James Bond | "The species is named after the famous fictional character James Bond because its provisory name in the PBI database, CR007, included Bond's code number (007)."  | [173]
Chimerella corleone Twomey, Delia & Castroviejo-Fisher, 2014 | Glass frog | Corleone family, The Godfather | "The specific name is a patronym for the Corleone family depicted in Mario Puzo's novel The Godfather and the trilogy of films directed by Francis Ford Coppola."  | [174]
Hyorhinomys stuempkei Esselstyn et al., 2015 | True mouse | Harald Stümpke, The Snouters: Form and Life of the Rhinogrades | "The species is named in honor of Gerolf Steiner, who used the pseudonym Harald Stümpke, to publish a small book (Bau und Leben der Rhinogradentia) commonly known in English as The Snouters (Stümpke 1967). The Snouters describes a fictional island radiation of mammals with extraordinary nasal and aural adaptations and seemingly anticipates the discovery of H. stuempkei, with its large pink nose and long pinnae."  | [175]
Solanum watneyi Martine & Frawley | Bush tomato | Mark Watney, The Martian | "The specific epithet of "watneyi" is inspired by the book and film, The Martian, in which the protagonist finds himself stranded on Mars surrounded by the planet's harsh terrain and reddish soils. In a shelter, he manages to grow a crop of potatoes (Solanum tuberosum) before finally being rescued by his astronaut colleagues. We’ve chosen to name Solanum watneyi after this character, Mark Watney, in part because of the similarly reddish soils of its habitat and the congeneric nature of the potato – but, most notably, as a way to honor the creation of a sci-fi hero botanist by author Andy Weir and to acknowledge perhaps the finest paean to botanical science (and botanical field work) that Hollywood has yet presented."  | [176]
†Aquilonifer spinosus Briggs et al., 2016 | Arthropod | The Kite Runner | "The name of the new taxon refers to the fancied resemblance between the tethered individuals and kites, and echoes the title of the 2003 novel The Kite Runner by Khaled Hosseini (aquila, eagle or kite; -fer, suffix meaning carry; thus aquilonifer, kite bearer)"  | [177]
Fujuriphyes Sánchez et al., 2016 | Mud dragon | Falkor (named Fujur in the Spanish version; the lead author is Spanish), The Neverending Story | "From Fujur, the dog-dragon in the novel The Neverending Story by M. Ende \+ Greek phyes, the commonly used suffix in names of Allomalorhagid genera. The name adds to the list of kinorhynch (mud dragons) species named after dragons and also refers to the study of kinorhynch phylogeny as a "never-ending story"."  | [178]
Epimeria cinderella d’Udekem d’Acoz & Verheye, 2017 | Amphipod | Cinderella | "Cinderella, heroin of humble origin in a well-known folk tale. The name [...] alludes to the modest size and the absence of ornamentation of the species, which contrasts with the extravagant adornment and the impressive size of many 'rival' Epimeria species."  | [15]
Meoneura goldemari Stuke & Freidberg, 2017 | Fly | King Goldemar, Goldemar | A tiny fly (length<2 mm) "named after King Goldemar, a dwarf who imprisoned the maid Hertlin, who later became the wife of the gothic King Dietrich von Bern. The story was told in the poem by Albrecht von Kemenaten."  | [46]
Meoneura nilsholgerssoni Stuke & Freidberg, 2017 | Fly | Nils Holgersson, The Wonderful Adventures of Nils | A tiny fly (length=2.4 mm) "dedicated to Nils Holgersson, the main character of The Wonderful Adventures of Nils by Selma Lagerlöf. Nils was enchanted, shrunk to the gnome size and travelled with wild geese across Sweden."
Meoneura oskari Stuke & Freidberg, 2017 | Fly | Oskar Matzerath, The Tin Drum | A tiny fly (length=2.3 mm) "named after Oskar Matzerath from the novel The Tin Drum (Die Blechtrommel) by Günter Grass. Born with the adult's level of spiritual development, Oskar decided at his third birthday never to grow up and stayed little for his whole life."
Myrmecium oompaloompa Candiani & Bonaldo, 2017 | Spider | Oompa-Loompas, Charlie and the Chocolate Factory | "This species is named after the Oompa Loompas, fictional characters from the motion picture Charlie and the Chocolate Factory [despite the reference to the film, they were literary characters first]. They are workers in Willie Wonka's [sic] factory and are paid solely with their preferred food, cocoa. The name refers to the fact that most specimens were collected in cocoa plantations in southern Bahia."  | [179]
Ochyrocera charlotte Brescovit et al, 2018 | Spider | Charlotte, Charlotte's Web | "The specific name refers to Charlotte, the spider from the classic 'Charlotte's Web' by E.B. White and a great friend of the pig named Wilbur."  | [103]
Ochyrocera misspider Brescovit et al, 2018 | Spider | Little Miss Spider | "The specific name refers to Little Miss Spider, a very popular spider around the world and the main character of the children's books by David Kirk."
†Buratina truncata Khramov, 2019 | Spongefly | Buratino | "Genus named after Buratina, a long-nosed character of the science fiction novel by Michael Kharitonov."  | [180]
Meoneura mucki Stuke & Barták, 2019 | Fly | Little Muck (German fairy tale) | "The "Kleine Muck" is a participant of Wilhelm Hauff's fairy tale "Die Karavane". The Kleine Muck is a small misshapen figure and outsider as Carnidae are small inconspicuous Diptera which hardly anybody is interested in."  | [181]
Mischocyttarus kallindusfloren Borges & Silveira, 2019 | Wasp | Kalinda, Neverness | "The specific epithet is a reference made to the computational goddess Kalinda of the flowers, created by the mathematician David Zindell in his book Neverness."  | [182]
Forcepsioneura gabriela Pimenta, Pinto & Takiya, 2019 | Damselfly | Gabriela, Gabriela, Clove and Cinnamon | "after the strong female character of the famous novel Gabriela, cravo e canela by Brazilian writer Jorge Amado. The novel is set in the region of the type locality at the beginning of the 20th century, when the southern coast of Bahia prospered from the exploitation of cacao trees."  | [183]
Tegenaria shirin Zamani & Marusik, 2019 | Spider | Shirin, Khosrow and Shirin | A funnel weaver spider from Iran, whose name "refers to the main female character of Khosrow and Shirin, a famous Persian tragic romance by the poet Nizami Ganjavi (1141–1209)."  | [184]
Coptoborus katniss Smith & Cognato, 2021 | Bark beetle | Katniss Everdeen, The Hunger Games | "The apex of the elytra declivity of this species is shaped like an arrowhead, Katniss' weapon of choice."  | [185]
Pseudotanais rapunzelae Błażewicz, Jakiel, Bamber & Bird, 2021 | Tanaid | Rapunzel | "Rapunzel is a princess in a Brothers Grimm fairy tale who, imprisoned in a high tower, offered to let down her long hair so that her lover could climb up to her. This alludes to the unusually long dorsodistal seta on pereopods 5 and 6."  | [186]
†Gallirallus astolfoi Salvador, Anderson & Tennyson, 2021 | Rail | Astolfo | An extinct flightless rail from the island of Rapa Iti, French Polynesia; "The specific epithet honours Astolfo, one of Charlemagne's fictional paladins. In the epic Orlando Furioso, Astolfo becomes trapped on a remote island because of the sorceress Alcina."  | [187]
Caramuruacarus Bassini-Silva & Jacinavicius, 2022 | Chigger | Caramuru | A genus native to Brazil, "named in honour of the epic poem Caramuru that exalts Brazilian lands, combined with the word 'acarus', which means 'mite' in Latin."  | [188]
Helicina marfisae Salvador, Silva & Bichuette, 2023 | Land snail | Marfisa | "After Marfisa, a fictional character in the epics Orlando Innamorato and Orlando Furioso. Marfisa was the Queen of India and one of the fiercest warriors on the land, often compared to a lioness. The name is an allusion to the name of the type locality [Gruta do Leão, Bahia state, Brazil], which translates to Cave of the Lion."  | [189]

## Comics[edit]

### The Adventures of Tintin[edit]

Taxon | Type | Named for | Notes | Ref

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Scelio dupondi Yoder, 2014 | Wasp | Dupont and Dupond, original French names  | Named for the close similarity of the sister species, just as the "two [twins] were always being confused with one another"  | [190]
Scelio janseni Yoder, 2014 | Wasp | Jansen and Janssen, translated Dutch names
Chaleponcus tintin Enghoff, 2014 | Millipede | Tintin | The specific epithet "refers to a cartoon character developed by the Belgian author Hergé because of the (somewhat remote) resemblance of the shape of the gonopod cucullus to Tintin's hairstyle."  | [191]
Neralsia haddocki van Noort, Buffington & Forshage, 2014 | Wasp | Captain Haddock | "The specific epithet [...] is for Captain Haddock, the comic book character by Hergé. The derivation has specific reference to Captain Haddock's consistent state of inebriation and utterance of the phrases "ten thousand thundering typhoons" and "billions of bilious blue blistering barnacles", expletives commiserate with the discovery and generic determination of this novel Afrotropical record in the CAR ethanol samples."  | [192]
Xyalophora tintini van Noort, Buffington & Forshage, 2014 | Wasp | Tintin | "The specific epithet [...] is for Tintin, the comic book character by Hergé, whose adventures in the Congo [where this species is found] have done much to popularise the country in a very controversial manner in parts of the world. The Xyalophora spine may possibly suggest Tintin’s famous tuft of hair."

### Popeye[edit]

Taxon | Type | Named for | Notes | Ref

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Campsicnemus popeye Evenhuis, 2013 | Fly | Popeye | "The specific name derives from the American cartoon character 'Popeye the Sailor Man', who had swollen forearms; and refers to the swollen mid tibia of the males"  | [193]
Masona popeye Quicke & Chaul, 2019 | Wasp | Popeye | "Name refers to the fictional cartoon character 'Popeye the sailorman', created by Elzie Crisler Segar, who had extremely swollen, muscular, fore arms." (similar to the femurs of this insect)  | [194]

### Asterix[edit]

Taxon | Type | Named for | Notes | Ref

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Stichoplastoris asterix (Valerio, 1980) | Tarantula | Asterix |  | [195]
Stichoplastoris obelix (Valerio, 1980) | Tarantula | Obelix |
†Abraracourcix Stroiński & Szwedo, 2011 | Planthopper | Vitalstatistix (Abraracourcix in the original French)  | A fossil genus found in Eocene Oise amber, in Northern France.  | [196]
†Ordralfabetix Szwedo & Jacek, 2011 | Planthopper | Unhygienix (Ordralfabétix in the original French)  | A fossil genus found in Eocene Oise amber, in Northern France.  | [197]
Nemesia asterix Decae & Huber, 2017 | Trapdoor spider | Asterix | "The trivial name asterix is derived from the Greek asteriskos meaning 'small star' and refers to the size and shape of the trapdoor that the species constructs. The spelling of the name is taken from the name of the fictional hero and star in the French comic book Asterix le Gaul 1959, by R. Goscinny & A. Uderzo."  | [198]
†Bela falbalae Ceulemans, Van Dingenen & Landau, 2018 | Sea snail | Panacea (Falbala in the original French)  | A fossil species from the Pliocene of western France, with a slender high spired shell, "Named after Falbala, the beautiful, tall, slender girl in the 'Asterix' comics."  | [199]
†Ondina asterixi Ceulemans, Van Dingenen & Landau, 2018 | Sea snail | Asterix | A fossil species from the Pliocene of western France, "Named after the heroic comic character Asterix, [...] the valiant Gaul who resisted Julius Caesar from an unnamed village set not far from the study area."
†Payraudeautia obelixi Landau, Ceulemans & Van Dingenen, 2018 | Sea snail | Obelix | A fossil species from the Miocene of western France, "Named after the heroic comic character Obelix, [...] the valiant Gaul who resisted Julius Caesar from an unnamed village set not far from the study area. Obelix is a rotund character, reminiscent of the shape of this new species."  | [200]
Trigonopterus asterix Riedel, 2019 | Weevil | Asterix |  | [201]
Trigonopterus idefix Riedel, 2019 | Weevil | Dogmatix (Idéfix in the original French)  |
Trigonopterus obelix Riedel, 2019 | Weevil | Obelix |

### DC Comics[edit]

Taxon | Type | Named for | Notes | Ref

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Otocinclus batmani Lehmann A., 2006 | Catfish | Batman | Named for having a "bat-shaped vertical spot on [its] caudal-fin"  | [202]
Euragallia batmani Rodrigues, Goncalves & Mejdalani, 2012  | Leafhopper | Batman | "The specific epithet, batmani, is a reference to the dorsal region of the aedeagal base in dorsal view, which closely resembles the open wings of a bat, like those of the Batman symbol."  | [203]
†Kalelia Pérez & del Río, 2017 | Clam | Kal-El, the Kryptonian name of Superman |  | [204]
Trissolcus nycteridaner Talamas, 2017 | Wasp | Batman | "Like Batman, species of Trissolcus are dark in color, rarely seen by the public, kill bad guys (stink bugs) for the benefit of humankind, and are at times unfairly vilified on the basis of their name (parasitic wasp). The epithet derives from the Greek words 'nycteris,' meaning 'bat', and 'aner,' meaning 'man'"  | [205]
Anthracites furvuseques Tan, Baroga-Barbecho & Yap, 2018 | Katydid | Batman | "The species name refers to the Dark Knight (in Latin; dark = furvus, knight = eques). This species is named after the fictional superhero character Batman in the Dark Knight Trilogy directed by Christopher Nolan. The species’ black colouration resembles the dark suit of the vigilante while the titillators also shows certain resemblance to the iconic Batman mask and logo in the film."  | [206]
Biconcavus batmani Figuerola, Gordon & Cristobo, 2018 | Bryozoan | Batman | "Alluding to 'Batman', a fictional superhero of comic books and movies, alluding to the distal oral rim bearing lateral stout tubercles which resemble a Batman mask."  | [207]
†Kariridraco dianae Cerqueira et al, 2021 | Pterosaur | Diana Prince, the civilian identity of Wonder Woman |  | [208]
Agamopus joker Costa-Silva & Carvalho & Vaz-De, 2022 | Scarab beetle | The Joker | "The specific name 'joker' [...] is an eponym after the homonymous supervillain from the DC Comics Universe. It is a reference to the sinuous pygidial sulcus of this species, which resembles the shape of a smiley clown's mouth."  | [209]

### Marvel Comics[edit]

Taxon | Type | Named for | Notes | Ref

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†Medusaceratops lokii Ryan, Russell & Hartman, 2010 | Ceratopsian dinosaur | Loki | Loki has a "helmet with [...] two giant hooks that come out of the top"  | [210][211]
Ogdoconta satana Metzler, Knudson & Poole, 2013 | Moth | Satana | "The scientific name satana comes from the Marvel comic book fictional character Satana, a child of Satan and sinister character, who taught black magic. The name refers to the black (often equated with evil) color of the adult moth."  | [212]
†Allodaposuchus hulki Blanco et al., 2015 | Crocodylomorph | Hulk | A fossil primitive crocodile from the Cretaceous of Spain, whose name comes "from the character of Marvel, Hulk; due to the strong muscle attachments of the bones."  | [213]
Bromeloecia wolverinei Yau & Marshall, 2018 | Fly | Wolverine | "This species is named for the group of 3 heavily sclerotized, flattened setae on the ventral surface of the surstylus, which closely resembles the claws and fist of the comic book character 'Wolverine'."  | [214]
Otiothops doctorstrange Cala-Riquelme et al, 2018 | Spider | Doctor Strange | "The specific epithet is a noun in apposition referring to the Marvel comics 'Doctor Strange' created by artist Steve Ditko and writer Stan Lee in 1963."  | [215]
†Thanos simonattoi Delcourt & Iori, 2018 | Theropod dinosaur | Thanos | "Genus name from the Greek thánato, meaning death and from the Marvel's [sic] character Thanos, the Conquer, created by Jim Starlin"  | [216]
Cirrhilabrus wakanda Tea, et al., 2019 | Wrasse | Wakanda | It was given the common name "Vibranium fairy wrasse" as the "purple chain-link scale pattern of the new species is reminiscent of" the fictional metal Vibranium.  | [217]
Discothyrea wakanda Hita Garcia & Lieberman, 2019 | Ant | Wakanda | "The new species is endemic to the Rwenzori Mountains in the Albertine Rift, the location of Wakanda in the Black Panther comics. Wakanda is a peaceful, prosperous country which was never colonized, where ancient cultural traditions coexist with conservation of natural resources and high-technology modernity. Discothyrea wakanda is named in honor of these ideals for Africa and the world."  | [218]
Miroculis wolverine Costa & Almeida & Salles, 2019 | Mayfly | Wolverine | "The specific epithet alludes to Marvel Comics character Wolverine, as the penis of the new species resembles the retractable adamantium claw that emerge from the back of the superhero's hand"  | [219]
Daptolestes bronteflavus Robinson & Yeates, 2020 | Robber fly | Thor | The specific name means "blonde thunder" and was given the common name "Thor's fly."  | [220]
Daptolestes illusiolautus Robinson & Yeates, 2020 | Robber fly | Loki | The specific name means "elegant deception" and was given the common name "Loki's fly."
Daptolestes feminategus Robinson & Yeates, 2020 | Robber fly | Black Widow | The specific name means "woman wearing leather" and was given the common name "Black Widow's fly."
Humorolethalis Robinson & Yeates, 2020 | Robber fly | Deadpool | The newly erected genus means "wet or moist and dead" and its sole member, Humorolethalis sergius, was given the common name "Deadpool's fly" for sharing similar markings to Deadpool's mask.
Habeastrum strangei Simone & Cavallari & Salvador, 2020 | Snail | Doctor Strange | "The specific epithet is given in honor of Dr. Stephen V. Strange, a fictional character from Marvel Comics. Besides being an acknowledgement of one of the greatest Marvel characters, the name also alludes to the weirdness of this dextral diplommatinid species."  | [221]
Gekko hulk Grismer et al., 2022 | Gekko | Hulk | Named for sharing the characteristics of "great physical strength and a very aggressive temperament"  | [222]
Voconia loki Castillo & Rédei & Weirauch, 2022 | True bug | Loki | "Named after the cunning trickster from Norse mythology and from the Marvel Comics’ character, Loki, since this specimen deceived and tricked authors in a previous study (Hwang & Weirauch 2012) who misidentified it as 'Kayanocoris wegneri'."  | [223]
Venomius Rossi, Castanheira, Baptista & Framenau, 2023 | Orb-weaver spider | Venom | "This genus-group name is a reference to the head of the character Venom, with conspicuous black spots, that resembles the abdomen of our species, specifically the male holotype." The only species in the genus, Venomius tomhardyi, is named after Tom Hardy, who plays Venom in the eponymous films.  | [224]

### Peanuts[edit]

Taxon | Type | Named for | Notes | Ref

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Lepidopa luciae Boyko, 2002 | Crab | Lucy Van Pelt | "The specific name 'luciae' is given after the 'Peanuts' character Lucy Van Pelt, as suggested by Jean Schulz, in recognition of Lucy's supremely 'crabby' attitude"  | [225]
Mexicope sushara Bruce, 2004 | Isopod | Pig-Pen | "The epithet combines the Latin words sus (pig) and hara (pen, coop or sty) and alludes to the ability of these preserved specimens to collect adherent detritus; referring to the character 'Pigpen' in the famous comic strip Peanuts, who gathered dirt no matter what."  | [226]
Ceratocorema woodstocki Liang & Hsu, 2017 | Moth | Woodstock | "Stripe patterns of adult body resemble the fictional character 'Woodstock' in Charles M. Schulz's comic strip of 'Peanuts'."  | [227]

### Monica and Friends[edit]

Taxon | Type | Named for | Notes | Ref

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Ochyrocera dorinha Brescovit et al., 2021 | Spider | Doreen/Dorinha | "Noun in apposition is a tribute to the fictional character of the Brazilian "Turma da Mônica" comic books by Maurício de Sousa. Dorinha, created in 2004, is a visually impaired character, in this case blind." This species has no eyes.  | [228]
Ochyrocera magali Brescovit et al., 2021 | Spider | Maggy/Magali | "Noun in apposition is a tribute to the fictional character of the Brazilian “Turma de Mônica” comic books by Maurício de Sousa, based on his daughter Magali. She is 7 years old and her main characteristic is her voracious appetite."
Ochyrocera monica Brescovit et al., 2021 | Spider | Monica | "Noun in apposition is a tribute to the fictional character of the Brazilian 'Turma da Mônica' comic books by Maurício de Sousa."
Ochyrocera rosinha Brescovit et al., 2021 | Spider | Rosinha  | "Noun in apposition is a tribute to the fictional character of the Brazilian “Turma da Mônica” comic books by Maurício de Sousa. She is a country girl, who is always wearing a red dress and a pair of pigtails in her hair."

### Calvin and Hobbes[edit]

Taxon | Type | Named for | Notes | Ref

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Zagrammosoma hobbesi LaSalle, 1989 | Wasp | Hobbes  | "Named after the tiger 'Hobbes' from the Calvin & Hobbes comic, as this species has yellow and black stripes on the face, similar to a tiger."  | [229]
Zagrammosoma calvini Perry, 2021 | Wasp | Calvin  | "The characteristic number of setae present on the head and body of Z. calvini is reminiscent of Calvin's spiked hair."

### Other comics[edit]

Taxon | Type | Named for | Notes | Ref

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†Ninjemys oweni Gaffney, 1992 | Turtle | Teenage Mutant Ninja Turtles | "Ninja, in allusion to that totally rad, fearsome foursome epitomizing shelled success; emys, turtle."  | [230]
Agra smurf Erwin, 2000 | Ground beetle | The Smurfs | "The specific epithet, smurf, is just for fun in that the weird head shape of this species reminded me of the Saturday-morning television cartoon characters of that name."  | [23]
Hypocaccus kidpaddlei Gomy, 2007 | Clown beetle | Kid Paddle | The beetle resembles the "blorks", fictional aliens from the Kid Paddle comic  | [231]
Neostygarctus lovedeluxe Fujimoto & Miyazaki, 2013 | Tardigrade | Love Deluxe, JoJo's Bizarre Adventure | "The specific epithet, lovedeluxe, refers to ‘Love Deluxe’, the name of a supernatural power enabling one to have complete control over one's hair, which appeared in JoJo's Bizarre Adventure Part 4: Diamond Is Unbreakable, a famous Japanese manga written and illustrated by Hirohiko Araki. The hairy appearance of the new species appears as if affected by the power of ‘Love Deluxe’."  | [232]
Euathlus condorito Perafán & Pérez-Miles, 2014 | Tarantula | Condorito | A Chilean species whose name is "inspired by the main character of the most popular Chilean comic book of the same name, and one of the most acclaimed comics in Hispanoamerica [sic]. Condorito represents a man–condor, emblematic bird of the Andeans and Chile's national symbol, created in 1949 by Chilean cartoonist 'Pepo'."  | [233]
Taito kakera Kury & Barros, 2014 | Harvestman | Kakera, Inuyasha | "From Japanese kakera = shard, because of the shape of the equuleus, reminiscent of the fragments of the Jewel of Four Souls in the Japanese manga ‘InuYasha, a Feudal Fairy Tale’, written and illustrated by Rumiko Takahashi."  | [234]
Nausicaamantis miyazakii Mériguet, 2018 | Mantis | Nausicaä, Nausicaä of the Valley of the Wind | The genus Nausicaamantis "is dedicated to the manga Nausicaä of the Valley of the Wind (Kaze no tani no Naushika) by Hayao Miyazaki, published between 1982 and 1995 [...] In this ecological fable, the heroine, Nausicaä, a naturalist and benevolent character, explores the relationship between humans and a hostile nature in a universe where humanity is in its twilight. Insects play a central role." The species "is dedicated to the creator of the work Nausicaä of the Valley of the Wind, Hayao Miyazaki. The author, sensitive to our environment, places nature at the centre of several of his works."
This species was described from a single specimen collected in Madagascar in 1906 (112 years before publication) and preserved in the National Museum of Natural History, France; its current conservation status is unknown and it may be extinct.  | [235]
Meoneura joedaltoni Stuke & Barták, 2019 | Fly | Joe Dalton, Lucky Luke | "The species is named after Joe Dalton, the smallest of the four Dalton brothers, who appear in the Lucky Luke comic book series of Maurice de Bevere (Morris) and René Goscinny. Joe is the smallest of the four Dalton brothers as Meoneura joedaltoni is one of the smallest Acalyptratae."  | [181]
Cigaritis syama lamuae Hsu & Liang, 2020 | Butterfly | Lamu, Urusei Yatsura | "The subspecific name lamuae refers to a comic character Lam (Lamu) created by a famous manga artist Rumiko Takahashi. The patterns of wing undersides recall the graphic design of the bikini Lam wears."  | [236]
Luthela kagami Wei & Lin, 2023 | Spider | Hiiragi Kagami, Lucky Star | "The specific epithet is from 'Hiiragi Kagami', a fictional character from the comic 'Lucky Star' (written and illustrated by the Japanese cartoonist Yoshimizu Kagami) with haircut similar to 'Asuka Langley Soryu'; the name refers to the great similarity between these two new species [Luthela kagami and Luthela asuka]."  | [237]
Funny valentine Lin & Li, 2024 | Spider | Funny Valentine, JoJo's Bizarre Adventure |  | [238]
Gyro zeppeli Lin & Li, 2024 | Spider | Gyro Zeppeli, JoJo's Bizarre Adventure |  | [239]

## Films[edit]

### Disney and Pixar[edit]

Taxon | Type | Named for | Notes | Ref

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Ceraeochrysa michaelmuris Adams & Penny, 1987 | Lacewing | Mickey Mouse | "The species name emphasizes the resemblance of the protruding gonapsis lobes to mouse ears"  | [240]
†Celmus michaelmus Adrain & Fortey, 1997 | Trilobite | Mickey Mouse | Its abdominal apex looks like Mickey Mouse  | [241]
†Bambiraptor feinbergi Burnham et al., 2000 | Theropod dinosaur | Bambi | "Bambi: from the now widely used nickname for the holotype, originally coined by the Linster family"  | [242]
Adelopsis dumbo Gnaspini & Peck, 2001 | Beetle | Dumbo | Named "because the beetle's aedeagus, which resembles an elephant proboscis, has at its tip a very large lateral projection resembling an ear"  | [243]
Corinna zecarioca Rodrigues et al., 2014 | Corinnid sac spider | José Carioca, Saludos Amigos | "The specific name is a noun in apposition referring to Walt Disney's character “José Carioca” or, shortly, “Zé Carioca”, created in 1942. The green parrot Zé Carioca is a friend of Donald Duck and the comic books are still popular among Brazilian kids, especially those born in the city of Rio de Janeiro, the Cariocas."  | [244]
Endodrelanva jimini Tan & Kamaruddin, 2016 | Cricket | Jiminy Cricket, Pinocchio | "The species is named after the Disney character, Jiminy cricket, from the book The Adventure of Pinocchio."  | [245]
Siphopteron dumbo Ong & Gosliner, 2017 | Sea slug | Dumbo | The specific name "refers to the similarity of this species to the Disney character, Dumbo the elephant, as it swims through the water."  | [246]
Helobdella buzz Iwama, Nogueira & Gonçalves, 2017 | Leech | Buzz Lightyear, Toy Story | "The protrusion of the skin on the nuchal region of Helobdella buzz n. sp. is a main diagnostic characteristic and its size and location resemble an astronaut helmet, as that of Buzz Lightyear."  | [247]
Epicratinus stitch Gonçalves & Brescovit, 2020 | Spider | Stitch, Lilo & Stitch | "The specific epithet is a noun taken in apposition and is in reference to Stitch, a fictional character in Disney's Lilo & Stitch franchise. The shade of the colors of its ears resembles the spermathecae from ventral view."  | [248]
Maratus nemo Schubert, 2021 | Jumping spider | Nemo, Finding Nemo | "The specific epithet refers to the colouration of the male of this species which resembles that of the character Nemo"  | [249]

### Looney Tunes[edit]

Taxon | Type | Named for | Notes | Ref

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Strotarchus beepbeep Bonaldo, et al., 2012 | Spider | "Beep beep", Road Runner | "The specific name refers to the cartoon character Road Runner, which always manages to fool its enemy Wile E. Coyote in the classic TV series Looney Tunes and Merrie Melodies. The Road Runner character vocalises mostly with a sharp " Beep, Beep " and was inspired upon the bird Geococcyx californianus that is recorded in Arizona, where the species here described is known to occur."  | [250]
Cremnops wyleycoyotius Tucker et al., 2015 | Wasp | Wile E. Coyote | "Named for the collector, J. Wiley, and for the holotype itself, which sneakily (like the well-known canine) remained undescribed until now"  | [251]

### Orson Welles[edit]

Taxon | Type | Named for | Notes | Ref

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Erwiniana rosebudae (Erwin, 1994) | Ground beetle | Rosebud, Citizen Kane | "The species is named for the sled in Orsen Wells' film, Citizen Kane, because the male genitalic apex is gracefully curved as was the front of the sled, yet such gracefulness existed in a crumbling empire as does the beetle in a rapidly disappearing rainforest."  | [252]
Orsonwelles ambersonorum Hormiga, 2002 | Spider | The Magnificent Ambersons |  | [253]
Orsonwelles arcanus Hormiga, 2002 | Spider | Mr. Arkadin | "The species epithet, a Latin noun in apposition meaning 'hidden, concealed', is derived from the Orson Welles film Mr. Arkadin/Confidential Report (1955)."
Orsonwelles calx Hormiga, 2002 | Spider | Harry Lime, The Third Man | "This species is named after Harry Lime, Welles' character in Carol Reed's movie The Third Man (1949). Calx is a Latin noun in apposition meaning 'lime' (that is, the mineral)."
Orsonwelles falstaffius Hormiga, 2002 | Spider | John Falstaff, Chimes at Midnight |
Orsonwelles iudicium Hormiga, 2002 | Spider | The Trial (1962 film) | Iudicium means 'trial' in Latin.
Orsonwelles macbeth Hormiga, 2002 | Spider | Macbeth (1948 film) |
Orsonwelles malus Hormiga, 2002 | Spider | Touch of Evil | "The species epithet, from the Latin adjective meaning 'evil,' is derived from the Orson Welles film Touch of Evil (1958)."
Orsonwelles othello Hormiga, 2002 | Spider | Othello (1951 film) |
Orsonwelles polites Hormiga, 2002 | Spider | Citizen Kane | "The species epithet is derived from the Orson Welles film Citizen Kane (1941). Polites (citizen) is a Greek noun in apposition."
Orsonwelles ventus Hormiga, 2002 | Spider | The Other Side of the Wind | "The species epithet, a Latin noun in apposition meaning 'wind', is derived from Welles' posthumous (and unfinished) film The Other Side of the Wind."

### Godzilla[edit]

Taxon | Type | Named for | Notes | Ref

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Godzilliidae Schram, Yager & Emerson, 1986 | Crustacean | Godzilla | See for Godzillius, the type genus of the family.  | [254]
Godzillius Yager, 1986 | Crustacean | Godzilla | "A reference to the almost monstrously large size of these animals as adults, the extreme styliform talon on the maxillule, and the grappling-like claws on the maxillae and maxillipedes."  | [254]
Godzillognomus Yager, 1989 | Crustacean | Godzilla | "The name is derived from Godzillius, the largest known remipede and the New Latin word "gnomus", meaning a diminutive fabled being."  | [255]
Pleomothra Yager, 1989 | Crustacean | Mothra | "In keeping with the spirit of the first described godzilliid, the name is derived from the Japanese horror creature Mothra" and the Greek word "pleo", meaning swim."  | [255]
†Gojirasaurus Carpenter, 1997 | Theropod dinosaur | Gojira, the Japanese name of Godzilla  | "Gojira, a large fictional monster of the Japanese cinema, in reference to the large size of this Triassic theropod."  | [256]
†Diplacodon gigan Mihlbachler, 2011 | Odd-toed ungulate | Gigan | "'Gigan' is a fictional giant horned monster first appearing in the 1972 Japanese film 'Godzilla versus Gigan' and other Godzilla films thereafter."  | [257]
Angirasu Hoenemann et al, 2013 | Crustacean | Anguirus | "Angirasu is another of the legendary Japanese Kaiju monsters that battled with Godzilla."  | [258]
Pleomothridae Hoenemann et al, 2013 | Crustacean | Mothra | See for Pleomothra, the type genus of the family.
Kumonga Hoenemann et al, 2013 | Crustacean | Kumonga | "Kumonga, a giant spider, is one of the legendary Japanese Kaiju monsters that battled with Godzilla."
Kumongidae Hoenemann et al, 2013 | Crustacean | Kumonga | See for Kumonga, the type genus of the family.
Amblyrhynchus cristatus godzilla Miralles et al, 2017 | Marine Iguana | Godzilla | "The subspecific epithet refers to the fictional saurian monster from the eponym movie franchise created by Tomoyuki Tanaka (IMDb, 2016). Explicitly mentioned in the title sequence of the 1990s American remake (Emmerich, 1998), the physical appearance and swimming behaviour of marine iguanas were a significant source of inspiration to the creature's designer (Patrick Tatopoulos, personal communication). The name is an invariable noun in apposition."  | [259]
Mecodema godzilla Seldon & Buckley, 2019 | Beetle | Godzilla | "Named for the giant mythical monster of Japan. Godzilla is a non-Latinised word so spelling remains invariant."  | [260]
Microgaster godzilla Fernandez-Triana & Kamino, 2020 | Wasp | Godzilla | "The wasp's parasitization behaviour bears some loose resemblance to the kaiju character, in the sense that the wasp (after diving underwater to search for its host, a moth caterpillar) suddenly emerges from the water (to parasitize the host), similar to how Godzilla suddenly emerges from the water in the movies."  | [261]
Scytodes kumonga Zamani & Marusik, 2020 | Spider | Kumonga | "The specific epithet is a noun in apposition and refers to a fictional, mutated, enormous “spitting” spider first appearing in Toho's 1967 movie Son of Godzilla."  | [262]
Agroeca angirasu Zamani & Marusik, 2021 | Spider | Anguirus | "The new species is named after Anguirus (Hepburn: Angirasu), a fictional monster which first appeared in Godzilla Raids Again (1955) and has a covering of spikes over his carapace, referring to the distinct rows of six pairs of long spines on the tibiae I and II of the holotype specimen."  | [263]
Ramisyllis kingghidorahi Aguado, Ponz-Segrelles, Glasby, Ribeiro, Jimi & Miura, 2022 | Bristle worm | King Ghidorah | "The name refers to King Ghidorah, the three-headed and two-tailed monster enemy of Godzilla. Both characters were created by Tomoyuki Tanaka based on Japanese mythology and folklore. King Ghidorah is a branching fictitious animal that can regenerate its lost ends."  | [264]
†Mutotylaspis tripudium René H.B. Fraaije, Eduard V. Mychko, Leonid S. Barsukov, John W.M. Jagt., 2023 | Crustacean | MUTO | "A combination of ‘Muto’, a giant parasitic monster (daikaiju) from the Godzilla universe that walks on four legs, and the generic name Tylaspis"  | [265]

### Star Wars[edit]

Main article: List of organisms named after the Star Wars series

### Alien[edit]

Taxon | Type | Named for | Notes | Ref

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†Xenomorphia Krogmann et al., 2018 | Wasp | Xenomorph | "The genus name refers to the endoparasitoid Xenomorph creature featured in the “Alien” media franchise."  | [266]
Dolichogenidea xenomorph Fagan-Jeffries & Austin, 2018 | Wasp | Xenomorph | "This species is named for the fictional creature from the movie franchise ‘Alien’, which reportedly was inspired by the lifecycle of parasitic wasps."  | [267]
Coptoborus newt Smith & Cognato, 2021 | Bark beetle | Rebecca "Newt" Jorden |  | [185]
Coptoborus ripley Smith & Cognato, 2021 | Bark beetle | Ellen Ripley | "This species is glabrous and reminiscent of Ripley's shaved head in 'Alien 3'"
Coptoborus vasquez Smith & Cognato, 2021 | Bark beetle | Private Vasquez |

### Terminator[edit]

Taxon | Type | Named for | Notes | Ref

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Hortipes terminator Bosselaers & Jocqué, 2000 | Spider | The Terminator | The male's pedipalps resemble a "futuristic gun"  | [268]
Coptoborus sarahconnor Smith & Cognato, 2021 | Bark beetle | Sarah Connor | "The vermiculate elytral declivity gives the species a rough appearance like the character it recognizes."  | [185]

### Crocodile Dundee[edit]

Taxon | Type | Named for | Notes | Ref

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Pristomerus dundeei Klopfstein, 2016 | Wasp | Michael "Crocodile" Dundee, Crocodile Dundee | An Australian species whose "name refers to one of the most famous Australians, the character "Crocodile Dundee" from the 1986 comedy movie set in the Australian outback."  | [269]
Qrocodiledundee outbackense Fernandez-Triana & Boudreault, 2018 | Wasp | Crocodile Dundee | "Named after the iconic Australian movie "Crocodile Dundee", one of the favorite movies of the first author [and] after the Outback, the vast and remote interior of Australia where the holotype specimen was collected."  | [270]

### Predator[edit]

Taxon | Type | Named for | Notes | Ref

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Predatoroonops Brescovit, Rheims & Ott, 2012 | Goblin spider | Predator | A genus of goblin spiders native to Brazil; "The name refers the fact that all species show the frontal area of the male chelicerae with modified structures that resemble the face of the Predator character"  | [271]
Predatoroonops anna Brescovit, Rheims & Bonaldo, 2012 | Goblin spider | Anna | "refers to the character Anna, played by Elpidia Carrillo"
Predatoroonops billy Brescovit, Rheims & Ott, 2012 | Goblin spider | Billy | "for Billy Sole, played by Sonny Landham."
Predatoroonops blain Brescovit, Rheims & Ott, 2012 | Goblin spider | Blain | "refers to the character Blain Cooper, played by Jesse Ventura"
Predatoroonops chicano Brescovit, Rheims & Santos, 2012 | Goblin spider | Poncho | "a second nickname for the character Jorge "Poncho" Ramirez, played by Richard Chaves"
Predatoroonops dillon Brescovit, Rheims & Bonaldo, 2012 | Goblin spider | Dillon | "refers to the character George [sic; actually Alan] Dillon, a former teammate of Dutch and current CIA agent, played by Carl Weathers"
Predatoroonops dutch Brescovit, Rheims & Bonaldo, 2012 | Goblin spider | Dutch | "refers to the main character in the movie Predator, Major Alan "Dutch" Schaefer, played by Arnold Schwarzenegger."
Predatoroonops maceliot Brescovit, Rheims & Ott, 2012 | Goblin spider | Mac | "refers to the character Mac Eliot, played by Bill Duke"
Predatoroonops olddemon Brescovit, Rheims & Santos, 2012 | Goblin spider | Predator (fictional species) | "a popular name used by natives of the fictional nation of Val Verde for the Predators"
Predatoroonops phillips Brescovit, Rheims & Santos, 2012 | Goblin spider | General Phillips | "refers to the character General Homer Phillips, played by Robert G. Armstrong"
Predatoroonops poncho Brescovit, Rheims & Ott, 2012 | Goblin spider | Poncho | "for Jorge "Poncho" Ramirez, played by Richard Chaves"
Predatoroonops rickhawkins Brescovit, Rheims & Bonaldo, 2012 | Goblin spider | Hawkins | "refers to the character Rick Hawkins, played by Shane Black"
Predatoroonops vallarta Brescovit, Rheims & Bonaldo, 2012 | Goblin spider | Puerto Vallarta | "refers to Puerto Vallarta, Mexico, one of the filming locations for the movie Predator; the set there is now a tourist attraction." (the species itself is not native to Mexico but Rio de Janeiro state, Brazil)
Predatoroonops valverde Brescovit, Rheims & Ott, 2012 | Goblin spider | Val Verde (fictional country) | "The specific name refers to the South/Central American country Val Verde, a fictional country created for Predator [sic] by Hollywood filmmakers, to avoid getting into legal or diplomatic disputes." (In actuality, Val Verde was created for a previous Schwarzenegger film, Commando, and its status as the location of the events of Predator is a fan theory, supported by Commando screenwriter Steven E. de Souza; other media of the Predator franchise place the action of the original in Guatemala or Colombia.)
Predatoroonops yautja Brescovit, Rheims & Santos, 2012 | Goblin spider | Predator (fictional species) | "refers to the fictional name by which Predators are known on their planet."

### The Fifth Element[edit]

Taxon | Type | Named for | Notes | Ref

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Agra lilu Erwin, 2000 | Ground beetle | Leeloo | Named after Leeloo for being "red-headed"  | [272]
Hirudicryptus quintumelementum Korsós et al., 2008 | Millipede | The Fifth Element | "The species is named as the fifth member of the millipede order Siphonocryptida; but also in honour of the alien custodians (called ‘‘mondoshawans’’) [...] whom the head and the collum of the new species (and actually all members of the order) resemble superficially"  | [273]
Coptoborus leeloo Smith & Cognato, 2021 | Bark beetle | Leeloo |  | [185]

### The Big Lebowski[edit]

Taxon | Type | Named for | Notes | Ref

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Anelosimus biglebowski Agnarsson, 2006 | Spider | The Big Lebowski |  | [274]
Anelosimus dude Agnarsson, 2006 | Spider | Jeffrey "The Dude" Lebowski, The Big Lebowski |

### Avatar[edit]

Taxon | Type | Named for | Notes | Ref

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†Ikrandraco avatar Wang et al., 2014 | Pterosaur | Ikran  | "Ikran, from the fictional flying creature portrayed in the movie Avatar that shows a well developed dentary crest and draco, from the Latin meaning dragon."  | [275]
Arctesthes avatar Patrick, Patrick & Hoare, 2019 | Moth | Avatar | "The name refers to the James Cameron movie Avatar; like the indigenous people and fauna of that film, the moth is vulnerable to habitat change or destruction in its very limited area of occurrence."  | [276]
†Oligomonoctenus neytiriae Nel et al, 2023 | Sawfly | Neytiri | "Named after Neytiri, princess of the Na'vi people from Pandora planet in the Avatar animation movie (James Cameron, 2009 and 2022), a blue and bioluminescent skin people who defend nature and life in all its forms."  | [277]
†Torukjara Pêgas, 2024 | Pterosaur | Toruk |  | [278]

### Other films[edit]

Taxon | Type | Named for | Notes | Ref

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†Chloridops regiskongi James & Olson, 1991 | Finch | King Kong, King Kong | "Olson was once quoted as saying this species was 'a giant, gargantuan, a King Kong finch', an appellation that would never have occurred to him, this being a typical example of the liberties taken with quotation marks by the print media"  | [279]
Eubetia boop Brown, 1998 | Moth | Betty Boop |  | [280]
†Ozraptor subotaii Long & Molnar, 1998 | Theropod dinosaur | Subotai, Conan the Barbarian | "After the fictional character Subotai, a swift running thief from the film "Conan the Barberian" [...], based on the Robert E. Howard books."  | [281]
†Sinemys gamera Brinkman & Peng, 1993 | Turtle | Gamera | The fossil has wing-like projections from its shell.  | [282]
Shrekin Britto & Navia, 2007 | Mite | Shrek | Named "because of the resemblance of the long, laterodorsal scapular tubercules to the long stalked ears of this character"  | [283]
Teratohyla amelie (Cisneros-Heredia & Meza-Ramos 2007) | Glass frog | Amélie Poulain, Amélie | Originally described as Cochranella amelie and subsequently transferred to genus Teratohyla.
"The specific name [...] of this new species of Glassfrog is for Amelie, protagonist of the extraordinary movie Le Fabuleux Destin d'Amélie Poulain; a movie where little details play an important role in the achievement of joie de vivre; like the important role that Glassfrogs and all amphibians and reptiles play in the health of our planet."  | [284]
[174]
†Gamerabaena Lyson & Joyce, 2010 | Turtle | Gamera | Name for the "fire-breathing turtle from the 1965 movie Gamera, in allusion to his fire breathing capabilities and the Hell Creek Formation"  | [256]
Eoperipatus totoro Oliveira et al., 2013 | Velvet worm | Totoro, My Neighbour Totoro | Named after Totoro, who "uses a many-legged animal [Catbus] as a vehicle"  | [285]
Cystomastacoides kiddo Quicke & Butcher, 2013 | Wasp | Beatrix Kiddo, Kill Bill | "Named after the character Beatrice Kiddo in the Quentin Tarantino ‘Kill Bill’ films because of the deadly biology to the host."  | [286]
Axima sidi Arias-Penna et al., 2014 | Wasp | Sid, Ice Age | "The name is based on facial resemblance between these two, which is mainly caused by shared bulbous eyes, and the characteristic anteroventral orientation of accompanying structures"  | [287]
Prochyliza georgekaplani Martín-Vega, 2014 | Fly | George Kaplan, North by Northwest | "The specific epithet [...] makes reference to George Kaplan, the nonexistent spy from Alfred Hitchcock's 1959 film North by Northwest for whom the main character is mistaken. Like in that celebrated film, P. georgekaplani had been misidentified as a "nonexistent" species (i.e. an unvalid [sic] name), at least in central Spain"  | [288]
†Zuul crurivastator Arbour & Evans, 2017 | Ankylosaurid dinosaur | Zuul, Ghostbusters |  | [289]
†Prolatcyclus kindzadza Mychko et al., 2019 | Crustacean | Kin-dza-dza! | A fossil cycloid from the Carboniferous of Orenburg Oblast, Russia.  | [290]
Coptoborus brigman Smith & Cognato, 2021 | Bark beetle | Dr. Lindsey Brigman, The Abyss |  | [185]
Coptoborus furiosa Smith & Cognato, 2021 | Bark beetle | Furiosa, Mad Max: Fury Road | "The 'spiny' elytra give the species a fierce appearance."
Coptoborus trinity Smith & Cognato, 2021 | Bark beetle | Trinity, The Matrix | "Three types of setae (trifid, scale-like and bristle-like) help diagnose this species."
Coptoborus vrataski Smith & Cognato, 2021 | Bark beetle | Rita Vrataski, Edge of Tomorrow | "The granulate elytral gives the species an armored appearance reminiscent of the character's combat jacket."
†Anisotremus rambo Lin & Wolf, 2022 | Fish | Rambo: First Blood Part II | A fossil grunt from the Eocene of Texas, USA, "Named after the famous movie 'Rambo', which alludes to the unrestrained, wild, but valiant appearance of the otoliths."  | [291]
Proctoporus optimus Mamani, Cruz, Mallqui, & Catenazzi, 2022 | Lizard | Optimus Prime, Transformers film series | "patronymic for Optimus Prime, leader of the Autobots in the science fiction movie Transformers, in recognition of the seventh film that was filmed in Machu Picchu [the type locality]: Transformers: Rise of the Beasts."  | [292]
Paravima totoro García & Villarreal, 2023 | Harvestman | Totoro, My Neighbour Totoro | "For us, the paramedian armature of the new species resembles the ears of the charismatic Totoro. We take advantage of exalting the excellent work of Studio Ghibli with this tribute."  | [293]

## Television[edit]

### Doctor Who[edit]

Taxon | Type | Named for | Notes | Ref

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†Yochelcionella daleki Runnegar & Jell, 1976) | Mollusc | Daleks | A fossil Helcionellid from the Cambrian of Australia.  | [294]
†Mestoronema Wagner, 2002 | Mollusc | Mestor, The Twin Dilemma | Named after "the ruler of the intelligent evil gastropods from the world's longest running science fiction serial, Doctor Who."  | [295]
Tetramorium dalek Hita Garcia & Fisher, 2014 | Ant | Daleks | "During different stages of the revision we considered placing the material listed here as Tetramorium dalek in at least three to four different groups, which caused a significant amount of nuisance, especially to the first author. Naming this species after an evil, extra-terrestrial, and often annoying race was a logical consequence."  | [153]
Synchiropus sycorax Tea & Gill, 2016 | Dragonet fish | Sycorax | "The species is named after the red-robed and caped Sycorax warriors from the BBC sci-fi series Dr. Who, in showing similarities in both coloration and grandiloquence of their garb."  | [296]
Cyclocardia dalek Pérez & Del Río, 2017 | Bivalve | Daleks | The Daleks are "characterized by an armour with prominent circles, similar to the nodular external sculpture of this species".  | [297]
Choeras zygon Fagan-Jeffries & Austin, 2019 | Wasp | Zygons | "The shape-shifting nature of this fictional race mirrors the large morphological variability within C. zygon [...] The Zygon in Doctor Who also consume their 'host', a trait particularly relevant to endoparasitoid wasps."  | [298]
†Ophiotardis Thuy & Numberger-Thuy, 2021 | Brittle star | TARDIS | "Genus name formed as a combination of óphis, Greek for serpent, a commonly used prefix in ophiuroid genus names, and Tardis, acronym for 'Time And Relative Dimension In Space'"  | [299]
Dalek nationi Noyes, 2023 | Wasp | Daleks, Terry Nation | "This species is named in honour of Terry Nation, creator of the Daleks, an alien species that has terrified children for the past 60 years." John S. Noyes, who described the genus, said, "I thought it was a good name for a genus and a bit of fun having been a big fan of Doctor Who in my early years."  | [300][301]

### Star Trek[edit]

Taxon | Type | Named for | Notes | Ref

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Conus tribblei Walls, 1977 | Sea snail | Tribbles | Named a pet cat that was named Tribbles after the Star Trek alien species  | [302]
Agra dax Erwin, 2000 | Ground beetle | Jadzia Dax | Also dedicated to Terry Farrell | [303]
†Boeckaspis geordii Karim, 2008 | Trilobite | Geordi La Forge | "After the television character Geordi La Forge who wears a similar eye ridge like visor."  | [304]
†Annuntidiogenes worfi Fraaije, 2009 | Hermit crab | Worf | Named "in reference to the wrinkled ornament of the anterior gastric region"  | [305]
Paridris gorn Talamas & Masner, 2012 | Wasp | Gorn | "This species is named after a reptilian alien race from the original Star Trek television series for the similar appearance of their compound eyes. The epithet is treated as a noun in apposition."  | [306]
Ledella spocki Viegas, Benaim & Absalão, 2014 | Mussel | Spock | Named because the species' "valves resembles the shape of the pointed ear of the Vulcans"  | [307]
Bolianus Karner, Salvato & Uliana, 2015 | Beetle | Bolian | "The peculiar median groove along the head prompted us to derive the generic name from the 'Bolians', a fictitious species from the universe of the science fiction series 'Star Trek', characterized by a vertical suture running along the midline of head and face."  | [308]
Synopeas klingunculum Awad, 2021 | Wasp | Klingons | "The epithet 'Synopeas klingunculum' means 'little Klingon' and refers to the rugose head sculpture, which resembles that of the fictional alien race from 'Star Trek'."  | [309]
Phanuromyia odo Nesheim, 2017 | Wasp | Odo | Named after Odo "because this species has variable morphology"  | [71]
Spockia Roca-Cusachs et al., 2019 | Stink bug | Spock | "[Spock] is a Vulcan/Human hybrid, this new genus shares with the commander Spock the fact that as it shares characters from genus Cazira and Blachia."  | [310]
Coptoborus hansen Smith & Cognato, 2021 | Bark beetle | Seven of Nine, born Annika Hansen  |  | [185]
Coptoborus janeway Smith & Cognato, 2021 | Bark beetle | Kathryn Janeway |
Coptoborus uhura Smith & Cognato, 2021 | Bark beetle | Nyota Uhura | "This species is reddish and reminiscent of the uniform Uhura wore"
Coptoborus yar Smith & Cognato, 2021 | Bark beetle | Tasha Yar |
Roddenberryus kirk Sánchez-Ruiz & Bonaldo, 2023 | Spider | James T. Kirk | Genus Roddenberryus was named after Star Trek creator Gene Roddenberry, and three newly discovered species in the genus were named after characters of the original series.  | [311]
Roddenberryus mccoy Sánchez-Ruiz & Bonaldo, 2023 | Spider | Leonard McCoy
Roddenberryus spock Sánchez-Ruiz & Bonaldo, 2023 | Spider | Spock

### Sesame Street and The Muppets[edit]

Taxon | Type | Named for | Notes | Ref

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†Geragnostus waldorfstatleri Turvey, 2005 | Trilobite | Statler and Waldorf | Named after "the resemblance of the pygidial axis to the heads of Waldorf and Statler"  | [312]
Parabradya samsoni Seifried, et al., 2007 | Copepod | Samson, Sesamstraße | "This species is named after the German character 'Samson' of the TV show 'Sesame Street' because of its big size and the unique ornamentation of body and setae."  | [313]
Stelis oscargrouchii Karremans (2015) | Orchid | Oscar the Grouch | "The name honors Oscar Grouch, of whom I am reminded by this extraordinary flower"  | [314]
†Hensonbatrachus kermiti Gardner & Brinkman, 2015 | Frog | Jim Henson, Kermit the Frog |  | [315]
Ariadna gonzo Marsh, Stevens & Framenau, 2022 | Tube-dwelling spider | Gonzo | "The specific epithet is in reference to the curved and hooked embolus of the pedipalp, which resembles the nose of the Muppet character Gonzo."  | [316]
†Kermitops So, Pardo & Mann, 2024 | Amphibian | Kermit the Frog | "Generic epithet is derived from a combination of 'Kermit' the famous lissamphibian and beloved Muppets' character created and originally performed by Jim Henson, and the Greek suffix '-ops', meaning face."  | [317]

### Dungeons & Dragons[edit]

Taxon | Type | Named for | Notes | Ref

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Regalana bobbyi Domahovski, Gonçalves & Cavichioli, 2014 | True bug | Bobby, the Barbarian  | "The new species name alludes to Bobby, the barbarian, one of the protagonists of the TV series Dungeons & Dragons."  | [318]
Regalana dianae Domahovski, Gonçalves & Cavichioli, 2014 | True bug | Diana, the acrobat  |
Regalana ericki Domahovski, Gonçalves & Cavichioli, 2014 | True bug | Eric, the cavalier  |
Regalana hanki Domahovski, Gonçalves & Cavichioli, 2014 | True bug | Hank, the ranger  |
Regalana prestoi Domahovski, Gonçalves & Cavichioli, 2014 | True bug | Presto, the magician  |
Alpaida venger Castanheira & Baptista, 2015 | Orb-weaver spider | Venger  | "The specific name refers to the antagonist character "Venger", from the animated television series named and inspired on the game Dungeons & Dragons. The character has a single horn on the side of the head, resembling the long paramedian apophysis of the male palp."  | [319]

### SpongeBob SquarePants[edit]

Taxon | Type | Named for | Notes | Ref

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Spongiforma squarepantsii Desjardin, Peay & T.D.Bruns, 2011 | Fungus | SpongeBob SquarePants | "Named in honor of the famed cartoon character SpongeBob SquarePants, whose shape shares a strong resemblance to the new fungus"  | [320]
Hemirhamphiculus krabsi Kritsky, 2017 | Monogenean flatworm  | Mr. Krabs | "The specific name (krabsi) was chosen because of the similar body shape of the species to that of Eugene H. Krabs (Mr. Krabs), a cartoon character in the children's animated television series SpongeBob SquarePants."  | [321]
Astrolirus patricki Zhang et al, 2020 | Starfish | Patrick Star | "Since all specimens of the new species were observed in situ living on sponges, it was name[d] by Patrick to reflect this curious relationship."  | [322]
Xizangiana plankton Li & Zhang, 2022 | Spider | Plankton | "The species is named after Sheldon J. Plankton, the main character in SpongeBob SquarePants, as the scape and anterior fold of epigyne of this new species resemble the body and flagellum of Plankton; noun (name) in apposition." Note that Plankton is the antagonist, not the main character.  | [323]

### Battlestar Galactica[edit]

Taxon | Type | Named for | Notes | Ref

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Eucteniza caprica Bond & Godwin, 2013 | Spider | Caprica-Six | Named "in reference to the humanoid cylon model Caprica 6, portrayed by Tricia Helfer in the remake of the science fiction series Battlestar Galactica."  | [324]
Coptoborus starbuck Smith & Cognato, 2021 | Bark beetle | Kara "Starbuck" Thrace | "The vermiculate elytral declivity gives the species a tough persona like the character it recognizes."  | [185]

### The Big Bang Theory[edit]

Taxon | Type | Named for | Notes | Ref

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Euglossa bazinga Nemésio & Ferrari, 2012 | Orchid bee | "Bazinga"  | "Euglossa bazinga sp. n. has tricked us for some time due to its similarity to E. ignita, which eventually led us to use 'bazinga'"  | [325]
Bazinga Gershwin & Davie, 2013 | Jellyfish | "Bazinga"  | "[B]azinga is a slang term in present popular culture, meaning 'fooled you!' [...] the type species, B. rieki, is so small that it has probably been overlooked in the past as a juvenile of a larger species."  | [326]
Kalcerrytus leonardi Bustamante, & Ruiz, 2016 | Jumping spider | Leonard Hofstadter | "The specific name is dedicated to the fiction character of “The Big Bang Theory” TV show, Dr. Leonard Hofstadter."  | [327]

### Other television series[edit]

Taxon | Type | Named for | Notes | Ref

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Savignia naniplopi Bosselaers & Henderickx, 2002 | Spider | Kabouter Plop | "The species is named after the gnome (Latin 'nanus') Plop, a popular character from children's stories whose cap is similar in shape to the male cephalic snout of the present species."  | [328]
Alphomelon simpsonorum Deans, 2003 | Wasp | Simpson family, The Simpsons | "named in honor of television's Simpson family for helping the author maintain a positive attitude throughout his educational endeavors."  | [329]
Acmopolynema isaura Triapitsyn & Berezovskiy, 2007 | Fairyfly | Escrava Isaura (1976 TV series) |  | [163]
Boccacciomymar (Prosto) maria Triapitsyn & Berezovskiy, 2007 | Fairyfly | Simplemente María (1989 TV series) | "The specific name (a noun in apposition) is a common feminine name; this species is named so for no particular reason other that it makes a good combination with the subgeneric name ("Prosto Maria" was a popular soap opera serial shown on Russian television in the 1990s)."
†Yochelcionella snorkorum Vendrasco et al., 2010 | Mollusc | Snorks | Species of the fossil genus Yochelcionella, from the Cambrian of Australia, were characterized by a snorkel connected to their shell; this one is "Named after the Snorks, fictional creatures in an animated television series characterized by a prominent snorkel extending from the head."  | [330]
Odontacolus zimi Valerio & Austin, 2013 | Wasp | Zim, Invader Zim | "This species is named after the anime character 'Invader Zim', in reference to the invasion of the spider egg sacs that occurs when Odontacolus oviposit."  | [331]
Lycocerus evangelium Hsiao & Okushima, 2016 | Soldier beetle | Neon Genesis Evangelion | "The specific epithet is derived from the Latin evangelium ('good news'), referring to [the] fact that its discovery was good news for the team; the specific name is also in memory of the anime 'Neon Genesis Evangelion', which is one of the greatest animes [sic] in Japanese history and had a strong impact on Japanese popular culture"  | [332]
Elthusa xena van der Wal, 2019 | Isopod | Xena, Xena: Warrior Princess | "This species is named after Xena, the warrior princess, in reference to the strong nature of the female cymothoid isopod."  | [333]
Prosopanche demogorgoni Funez, Ribeiro-Nardes, Kossmann et al.., 2019 | Flowering plant | Demogorgon, Stranger Things | "Demogorgon is a fictional monster whose mouth resembles the P. demogorgoni flower."  | [334]
Salticus lucasi Zamani, Hosseini & Moradmand, 2020 | Jumping spider | Lucas the Spider | "The species is named after Lucas the Spider, an animated character created by animator Josh Slice on the basis of the anatomy of jumping spiders, in recognition of the role that it played in "curing" many arachnophobes around the world".  | [335]
Coptoborus scully Smith & Cognato, 2021 | Bark beetle | Dana Scully, The X-Files | The authors also highlighted the "Scully Effect" | [185]
Spiralix heisenbergi Quiñonero-Salgado, Alonso & Rolán, 2021 | Freshwater snail | Walter White, alias Heisenberg, Breaking Bad | "The name is derived from the character Walter White, known as 'Heisenberg', in the TV series Breaking Bad."  | [336]
Luthela asuka Wei & Lin, 2023 | Spider | Asuka Langley Soryu, Neon Genesis Evangelion | "The specific epithet is from 'Asuka Langley Soryu', a fictional character wearing a red combat suit from the animation 'Evangelion' (by the Japanese creator Hideaki Anno), refers to the body color."  | [237]

## Games[edit]

### Galaga[edit]

Taxon | Type | Named for | Notes  |

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†Galagadon nordquistae Gates et al., 2019 | Carpet shark | Galaga | "[N]amed for the shape of the teeth, which when seen in different views resemble the spaceships in the arcade game 'Galaga'"  | [337]
Taito galaga Kury & Barros, 2014 | Harvestman | Galaga | Named in reference to "the shape of the alien insectoids which resemble the butterfly-shaped equuleus of this species".  | [234]

### Super Mario[edit]

Taxon | Type | Named for | Notes | Ref

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Rotundicardia mariobrosorum Pé & del Río, 2017 | Bivalve | Mario and Luigi, the "Mario Bros."  | "The specific epithet honors Mario and Luigi, the Mario Bros. brothers, main characters from the popular videogame Mario Bros., in which they collect mushrooms, and it is a reference to the 'funginate' nodes of the radial ribs in this species."  | [204]
Maraenobiotus supermario Novikov & Sharafutdinova, 2020 | Copepod | Mario | "This species is named after the character of the video game Super Mario, who, like our species, goes often underground and wears a funny mustache (mandibular palp)."  | [338]

### The Legend of Zelda[edit]

Taxon | Type | Named for | Notes | Ref

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Epicratinus zelda Gonçalves & Brescovit, 2020 | Spider | Princess Zelda | "Princess Zelda is the titular character in Nintendo's The Legend of Zelda video game series. [...] The shape of the epigynum remembers the Hyrule's symbol."  | [248]
Lanayrella Salvador & Cunha, 2020 | Gastropod | Lanayru Sea  | "Named after the Lanayru Sea from the game 'The Legend of Zelda: Breath of the Wild'"  | [339]
Orcevia yahaha Yu, Maddison & Zhang, 2023 | Jumping spider | Koroks  | "The specific epithet is a noun in apposition from 'Yahaha' (also known as 'Korok'), a very cute pixie in the game The Legend of Zelda: Breath of the Wild. Yahaha often hide in tree trunks, bushlands, or rocks, and if you find them, they will say 'Yahaha!' and share with you some 'fruits' as a gift. Collecting Orcevia specimens is very like searching for Yahaha."  | [340]
Orcevia bokoblin Yu, Maddison & Zhang, 2023 | Jumping spider | Bokoblins  | "The specific epithet is from 'Bokoblin', a small piggy monster in the game The Legend of Zelda: Breath of the Wild, developed and published by Nintendo. In the Master Mode of this game, Bokoblin commonly has either silver or gold coloration, corresponding to two color-forms observed in the males of the new species."  | [340]

### Street Fighter[edit]

Taxon | Type | Named for | Notes | Ref

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Epicratinus ehonda Gonçalves & Brescovit, 2020 | Spider | E. Honda | "The specific epithet is a noun taken in apposition and is in reference to Edmond Honda, a fictional character in Street Fighter series game from CAPCOM. E. Honda is a Japanese sumo wrestler, and the epigynum resembles two sumo wrestlers in fighting position."  | [248]
Epicratinus zangief Gonçalves & Brescovit, 2020 | Spider | Zangief | "The specific epithet is a noun taken in apposition and is in reference to Zangief, a fictional character in Street Fighter series game from CAPCOM. Zangief is a soviet strongman, and the RTA resembles a sickle, present in old USSR flag."

### Pokémon[edit]

Taxon | Type | Named for | Notes | Ref

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Stentorceps weedlei Neilson & Buffington, 2011 | Wasp | Weedle | "Weedle shares the distinguishing character of S. weedlei, a spine in the middle of its head."  | [341]
†Aerodactylus Vidovic & Martill, 2014 | Pterosaur | Aerodactyl | "The name derives from the Nintendo Pokémon Aerodactyl, a fantasy creature made up of a combination of different pterosaurian features"  | [342]
Chilicola charizard Monckton, 2016 | Bee | Charizard | "The specific epithet is in homage to the fictional monster which this species resembles"  | [343]
†Bulbasaurus phylloxyron Kammerer & Smith, 2017 | Dicynodont | Bulbasaur | "Bulbasaurus (bulb lizard) refers to the bulbous nasal boss and the species phylloxyron (leaf razor) refers to the sharpened beak for slicing through the plant material it ate [...] similarities between this species and certain other squat, tusked quadrupeds may not be entirely coincidental."  | [344][345]
Dicranocentrus pikachu Xisto & Cleide de Mendonça, 2017 | Springtail | Pikachu |  | [346]
Parapharyngodon politoedi Santos et al., 2019 | Roundworm | Politoed | A parasite of the Manaus slender-legged tree frog; "The species epithet is derived from the fictional character named 'Politoed', a frog-type Pokémon from the Pokémon Universe."  | [347]
Binburrum articuno Hsiao & Pollock, 2020 | Fire-coloured beetle | Articuno |  | [348]
Binburrum zapdos Hsiao & Pollock, 2020 | Fire-coloured beetle | Zapdos |
Binburrum moltres Hsiao & Pollock, 2020 | Fire-coloured beetle | Moltres |
Epicratinus pikachu Gonçalves & Brescovit, 2020 | Spider | Pikachu | "The female epigynum resembles the face of Pikachu."  | [248]
Rathalos treecko Lin & Li, 2021 | Spider | Treecko | "The species is named after Treecko, a fictional character from Pokémon Emerald that lives in the forest, as does this new species"  | [349][350]
Anyphaena grovyle Lin & Li, 2021 | Spider | Grovyle | "The species is named after Grovyle, a fictional character from Pokémon Emerald that lives in the forest, as does the new species"
Anyphaena sceptile Lin & Li, 2021 | Spider | Sceptile | "The species is named after Sceptile, a fictional character from Pokémon Emerald who lives in the forest, as does this new species"
Alistra pikachu Lin & Li, 2021 | Spider | Pikachu | "The species is named after Pikachu, a fictional character from Pokémon Yellow, as the habitus color of this new species is yellow"
Hiperantha pikachu Pineda & Barros, 2021 | Beetle | Pikachu | "The specific name is a homage to Pikachu, a fictional monster which this species resembles in its yellow elytra with a black apical band (like the ears of Pikachu)."  | [351]
Nocticola pheromosa Lucañas & Foo, 2023 | Cockroach | Pheromosa | "There are some similarities between Pheromosa and the delicate cockroach [...] found, such as having a long antenna, wings that mimic a hood and long slender legs."  | [352][353]
Phrynarachne dreepy Lin & S. Li, 2022 | Spider | Dreepy | "The species is named after Dreepy, a fictional character from Pokémon Sword and Shield, who has a triangular head that is reminiscent of the opisthosoma of the new species."  | [354]

### BioShock[edit]

Taxon | Type | Named for | Notes | Ref

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Rapturella ryani Salvador & Cunha, 2016 | Gastropod | Rapture and Andrew Ryan | "In honour of Andrew Ryan, the founder of the deep-sea city Rapture, from the science-fiction video game series BioShock."  | [355]
Rapturella atlas Cunha & Simone, 2018 | Gastropod | Rapture and Frank "Atlas" Fontaine | "In honors of Atlas, the nick name of Frank Fontaine, the character of the science-fiction video game series BioShock: an allusion to the hidden identity of the species."  | [356]

### Other games[edit]

Taxon | Type | Named for | Notes | Ref

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†Cortana Salvador & Simone, 2013 | Gastropod | Cortana, Halo | "The name was taken from a character of the science fiction franchise 'Halo', and alludes to the convoluted markings on the shell surface of the holotype of Cortana carvalhoi"  | [357]
†Crash bandicoot Travouillon et al., 2014 | Bandicoot | Crash Bandicoot, Crash Bandicoot | Named after Crash Bandicoot to allude to the "inference that this was the start of a new radiation of more modern bandicoots that 'crashed' through to dominate younger, drier ecosystems of Australia."  | [358]
Halystina umberlee Salvador, Cavallari & Simone, 2014 | Gastropod | Umberlee, Dungeons & Dragons Forgotten Realms | "[N]amed after Umberlee, a fictional goddess of the deep sea from the Faerûnian pantheon of the Forgotten Realms campaign setting of the Dungeons & Dragons role-playing game."  | [359]
Taito spaceinvaders Kury & Barros, 2014 | Harvestman | Space Invaders | "Space Invaders (Japanese Supûsu Inbêdâ) is an arcade video game manufactured and sold by Taito and very successful and popular worldwide in the 1980s."  | [234]
Neobuthus factorio Kovařík, Lowe, Awale, Elmi, & Hurre, 2018 | Scorpion | Factorio | Named after the video game Factorio, which was created by Michal Kovařík, the son of one of the researches who described the species, František Kovařík.  | [360]
Demyrsus digmon Hsiao & Oberprieler, 2020 | Weevil | Digmon, Digimon Adventure 02 | Named after the insectoid Digmon, "who possesses the great power of drilling and manipulating the earth, in reference to the habit of this species, which can bore into hard trunk of cycads."  | [361]
Epicratinus mauru Gonçalves & Brescovit, 2020 | Spider | Mauru, Waku Waku 7 | "The specific epithet is a noun taken in apposition and is in reference to Mauru, a fictional character in Waku Waku Seven, game from SUNSOFT for Neo Geo, which is the non-threatening guardian of Lost Forest."  | [248]
Epicratinus omegarugal Gonçalves & Brescovit, 2020 | Spider | Omega Rugal, The King of Fighters | "The specific epithet is a noun taken in apposition and is in reference to Omega Rugal, a fictional character in The King of Fighters series game from SNK, which is the boss on the first game and a recurrent character on this series, the epigynum looks like an Omega letter, from Greek alphabet."
Euconnus hosakae Hoshina, Fukutomi, & Watanabe, 2020 | Rove beetle | Miyuki Hosaka, Sentimental Graffiti | The type specimen and Miyuki Hosaka are both from Kanazawa.  | [362]
Emphysemastix frampt Olsen & Enghoff, 2020 | Millipede | Kingseeker Frampt, Dark Souls | Named after Kingseeker Frampt from the 2011 video game Dark Souls due to the gonopods' resemblance to the character  | [363]
Abaddon despoliator Derkarabetian, 2021 | Harvestman | Abaddon the Despoiler, Warhammer 40,000 | Named after Abaddon the Despoiler, who is "typically portrayed adorned with spikes and various sharp things"  | [364]

## Other media[edit]

Taxon | Type | Named for | Notes | Ref

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Orsonwelles bellum Hormiga, 2002 | Spider | The War of the Worlds (1938 radio drama) | "This species, collected below the radio tower on Mount Kahili, is named after Orson Welles' 1938 radio broadcast of H. G. Wells' War of the Worlds. Bellum (war) is a Latin noun in apposition."  | [253]

## See also[edit]

* List of unusual biological names
* List of organisms named after famous people

## References[edit]

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2. ^ Davis, Lauren (August 20, 2009). "30 Real Animals with Science Fiction Names". Gizmodo. Retrieved 2023-08-05.
3. ^ Sullivan, Jay. "Ten animals named after mythological creatures". Natural History Museum.
4. ^ Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. pp. 49–50. ISBN 978-1-4081-2501-4.
5. ^ Beolens, B.; Watkins, M.; Grayson, M. (2009). The Eponym Dictionary of Mammals. Johns Hopkins University Press. p. 296\. ISBN 978-0-8018-9533-3.
6. ^ Dunn, Emmett R. (1921). "Two new Central American salamanders". Proceedings of the Biological Society of Washington. 34: 143–146. Archived from the original on 2023-03-09. Retrieved 2022-05-10.
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8. ^ Linnavuori, R. E. (1997). "Taxonomic Studies on the Miridae (Heteroptera) of Yemen and Iran" (PDF). Acta Universitatis Carolinae. Biologica. 40: 301–320. Archived from the original (PDF) on 2020-08-05. Retrieved 2022-11-30.
9. ^ a b c Jakiel, A.; Palero, F.; Błażewicz, M. (2019). "Deep ocean seascape and Pseudotanaidae (Crustacea: Tanaidacea) diversity at the Clarion-Clipperton Fracture Zone". Sci Rep. 9 (1): 17305. Bibcode:2019NatSR...917305J. doi:10.1038/s41598-019-51434-z. PMC 6872736. PMID 31754124.
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11. ^ Liu, Bo; Zhang, Feng (2024-01-18). "Hotwheels gen. nov., a new ground spider genus (Araneae, Gnaphosidae) from southwest China". ZooKeys (1189): 337–347. doi:10.3897/zookeys.1189.115996. PMC 10836147. PMID 38314110.
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16. ^ Knapp, Sandra; Chase, Mark W. (2021). Extraordinary orchids. Chicago: The University of Chicago Press. p. 22\. ISBN 9780226779706.
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24. ^ Portman, Zachary M.; Neff, John L.; Griswold, Terry (2016-12-23). "Taxonomic revision of Perdita subgenus Heteroperdita Timberlake (Hymenoptera: Andrenidae), with descriptions of two ant-like males". Zootaxa. 4214 (1): zootaxa.4214.1.1. doi:10.11646/zootaxa.4214.1.1. ISSN 1175-5334. PMID 28006789. Archived from the original on 2022-06-17. Retrieved 2022-05-26.
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