Northern giraffe

Proposed species of giraffe

See Giraffe for details of how this proposed taxonomy fits with the currently accepted taxonomy of giraffes.

Northern giraffe

In Murchison Falls National Park

Conservation status

Vulnerable (IUCN 3.1)[1]

CITES Appendix II (CITES)

Scientific classification
Domain:  | Eukaryota
Kingdom:  | Animalia
Phylum:  | Chordata
Class:  | Mammalia
Order:  | Artiodactyla
Family:  | Giraffidae
Genus:  | Giraffa
Species:  |

G. camelopardalis

Binomial name
Giraffa camelopardalis

Linnaeus, 1758

The northern giraffe (Giraffa camelopardalis), also known as three-horned giraffe,[2] is the type species of giraffe, G. camelopardalis, and is native to North Africa, although alternative taxonomic hypotheses have proposed the northern giraffe as a separate species.[3][1]

Once abundant throughout Africa since the 19th century, the northern giraffe ranged from Senegal, Mali and Nigeria from West Africa to up north in Egypt. [4] The similar West African giraffe lived in Algeria and Morocco in ancient periods until their extinctions due to the Saharan dry climate.[5][6][4]

Giraffes collectively are listed as Vulnerable on the IUCN Red List, as the global population is thought to consist of about 97,000 individuals as of 2016.[1]

## Taxonomy and evolution

[edit]

The current IUCN taxonomic scheme lists one species of giraffe with the name G. camelopardalis and nine subspecies.[1][7] A 2021 whole genome sequencing study suggests the northern giraffe as a separate species, and postulates the existence of three distinct subspecies,[8] and more recently, one extinct subspecies.[9][10][11]

Image | Subspecies | Description | Distribution

|
|
|

| Kordofan giraffe (G. c. antiquorum) | Its spots may be found below the hocks and the insides of the legs. A median lump is present in males. | Southern Chad, the Central African Republic, northern Cameroon, and the northeastern DR Congo.
| Nubian giraffe (G. c. camelopardalis) | It has sharply defined chestnut-coloured spots surrounded by mostly white lines, while undersides lack spotting. Includes the Rothschild's giraffe ecotype | Eastern South Sudan and southwestern Ethiopia, in addition to Kenya and Uganda.
| West African giraffe (G. c. peralta) | This animal has a lighter pelage than other subspecies, with red lobe-shaped blotches that reach below the hocks. | Southwestern Niger
| Senegalese giraffe (G. c. senegalensis)  | It had dark brown patches, with a clear contour. Body was almost uniform in size.  | Extinct; formerly parts of Senegal, The Gambia, Mali, and Mauritania up until the 1970s.

## Description

[edit]

Skull of a northern giraffe

Three-dimensional model of a northern giraffe skeleton

Often mistaken with the southern giraffe, the northern giraffe differs by the shape and size of the two distinctive horn-like protuberances known as ossicones on its forehead; they are longer and larger than those of southern giraffe. Male northern giraffes have a third cylindrical ossicone in the center of the head just above the eyes, ranging from 76 to 127 mm (3 to 5 in) long.[2]

## Distribution and habitat

[edit]

Northern giraffes live in savannahs, shrublands, and woodlands. After numerous local extinctions, Northern giraffes are the least numerous giraffe species, and the most endangered. In East Africa, they are mostly found in Kenya and southwestern Ethiopia, and rarely in northeastern Democratic Republic of the Congo and South Sudan. In Central Africa, there are about 2,000 in the Central African Republic, Chad and Cameroon. Once widespread in West Africa, a few hundred Northern giraffes are confined in the Dosso Reserve of Kouré, Niger. They are isolated in South Sudan, Kenya, Chad and Niger. They commonly live both in and outside of protected areas.[1]

The earliest ranges of the Northern giraffes were in Chad during the late Pliocene. Once abundant in North Africa, they lived in Algeria from the early Pleistocene during the Quaternary period. They lived in Morocco, Libya and Egypt until their extinction there around AD 600, as the drying climate of the Sahara made conditions impossible for giraffes. Giraffe bones and fossils have been found across these countries.[5][6]

## References

[edit]

1. ^ a b c d e Muller, Z.; Bercovitch, F.; Brand, R.; Brown, D.; Brown, M.; Bolger, D.; Carter, K.; Deacon, F.; Doherty, J.B.; Fennessy, J.; Fennessy, S.; Hussein, A.A.; Lee, D.; Marais, A.; Strauss, M.; Tutchings, A.; Wube, T. (2018) [amended version of 2016 assessment]. "Giraffa camelopardalis". IUCN Red List of Threatened Species. 2018: e.T9194A136266699. doi:10.2305/IUCN.UK.2016-3.RLTS.T9194A136266699.en. Retrieved 12 November 2021.
2. ^ a b De Winton, W.E. (1897). "Remarks on the existing forms of Giraffe". Proceedings of the Zoological Society of London (February): 273–285.273-285&rft.date=1897&rft.au=De+Winton,+W.E.&rft_id=https://archive.org/stream/biostor-107888#page/n11/mode/2up&rfr_id=info:sid/en.wikipedia.org:Northern+giraffe" class="Z3988">
3. ^ Petzold, A.; Hassanin, A. (2020). "A comparative approach for species delimitation based on multiple methods of multi-locus DNA sequence analysis: A case study of the genus Giraffa (Mammalia, Cetartiodactyla)". PLOS ONE. 15 (2): e0217956. Bibcode:2020PLoSO..1517956P. doi:10.1371/journal.pone.0217956. PMC 7018015. PMID 32053589.
4. ^ a b Hassanin, A.; Ropiquet, A.; Gourmand, A.-L.; Chardonnet, B.; Rigoulet, J. (2007). "Mitochondrial DNA variability in Giraffa camelopardalis: consequences for taxonomy, phylogeography and conservation of giraffes in West and central Africa". Comptes Rendus Biologies. 330 (3): 265–274\. doi:10.1016/j.crvi.2007.02.008. PMID 17434121.265-274&rft.date=2007&rft_id=info:doi/10.1016/j.crvi.2007.02.008&rft_id=info:pmid/17434121&rft.aulast=Hassanin&rft.aufirst=A.&rft.au=Ropiquet,+A.&rft.au=Gourmand,+A.-L.&rft.au=Chardonnet,+B.&rft.au=Rigoulet,+J.&rft_id=https://doi.org/10.1016%2Fj.crvi.2007.02.008&rfr_id=info:sid/en.wikipedia.org:Northern+giraffe" class="Z3988">
5. ^ a b Dagg, A.I. (2014). Giraffe: Biology, Behaviour and Conservation. Cambridge: Cambridge University Press. ISBN 9781107729445.
6. ^ a b Wendorf, F.; Schild, R. (2013). Holocene Settlement of the Egyptian Sahara: Volume 1: The Archaeology of Nabta Playa. Springer Science & Business Media. ISBN 9781461506539.
7. ^ Bercovitch, F.B.; Berry, P.S.M.; Dagg, A.; Deacon, F.; Doherty, J.B.; Lee, D.E.; Mineur, F.; Muller, Z.; Ogden, R. (2017). "How many species of giraffe are there?". Current Biology. 27 (4): R136 – R137. Bibcode:2017CBio...27.R136B. doi:10.1016/j.cub.2016.12.039. hdl:20.500.11820/4c828f1a-f644-4268-9197-eb50244a1d75. PMID 28222287.R136+-+R137&rft.date=2017&rft_id=info:hdl/20.500.11820/4c828f1a-f644-4268-9197-eb50244a1d75&rft_id=info:pmid/28222287&rft_id=info:doi/10.1016/j.cub.2016.12.039&rft_id=info:bibcode/2017CBio...27.R136B&rft.aulast=Bercovitch&rft.aufirst=F.B.&rft.au=Berry,+P.S.M.&rft.au=Dagg,+A.&rft.au=Deacon,+F.&rft.au=Doherty,+J.B.&rft.au=Lee,+D.E.&rft.au=Mineur,+F.&rft.au=Muller,+Z.&rft.au=Ogden,+R.&rft_id=https://doi.org/10.1016%2Fj.cub.2016.12.039&rfr_id=info:sid/en.wikipedia.org:Northern+giraffe" class="Z3988">
8. ^ Coimbra, R.T.F.; Winter, S.; Kumar, V.; Koepfli, K.-P.; Gooley, R.M.; Dobrynin, P.; Fennessy, J.; Janke, A. (2021). "Whole-genome analysis of giraffe supports four distinct species". Current Biology. 31 (13): 2929–2938.e5. Bibcode:2021CBio...31E2929C. doi:10.1016/j.cub.2021.04.033. PMID 33957077.
9. ^ "Giraffa camelopardalis senegalensis Petzold, Magnant & Hassanin, 2020". www.gbif.org. Retrieved 2023-06-03.
10. ^ Petzold, A.; Magnant, A.-S.; Edderai, D.; Chardonnet, Bertrand; Rigoulet, J.; Saint-Jalme, Michel; Hassanin, Alexandre (2020). "First insights into past biodiversity of giraffes based on mitochondrial sequences from museum specimens". European Journal of Taxonomy (703). doi:10.5852/ejt.2020.703.
11. ^ "Giraffa camelopardalis senegalensis (Senegalese giraffe) - The Recently Extinct Plants and Animals Database". recentlyextinctspecies.com. Retrieved 2023-06-03.

## External links

[edit]

Northern giraffe at Wikipedia's sister projects

* Definitions from Wiktionary
* Media from Commons
* Texts from Wikisource
* Taxa from Wikispecies

* ARKive – images and movies of the giraffe (Giraffa camelopardalis).
* Giraffe, African Wildlife Foundation
* Giraffe

* v
* t
* e

Extant Artiodactyla species

* Kingdom: Animalia
* Phylum: Chordata
* Class: Mammalia
* Infraclass: Eutheria
* Superorder: Laurasiatheria

Suborder Ruminantia

| Antilocapridae|

| Antilocapra|

* Pronghorn (A. americana)

|

Giraffidae|

| Okapia|

* Okapi (O. johnstoni)

|

Giraffa|

* Northern giraffe (G. camelopardalis)
* Southern giraffe (G. giraffa)
* Reticulated giraffe (G. reticulata)
* Masai giraffe (G. tippelskirchi)

Moschidae|

| Moschus|

* Anhui musk deer (M. anhuiensis)
* Dwarf musk deer (M. berezovskii)
* Alpine musk deer (M. chrysogaster)
* Kashmir musk deer (M. cupreus)
* Black musk deer (M. fuscus)
* Himalayan musk deer (M. leucogaster)
* Siberian musk deer (M. moschiferus)

|

Tragulidae|

| Hyemoschus|

* Water chevrotain (H. aquaticus)

|

Moschiola|

* Indian spotted chevrotain (M. indica)
* Yellow-striped chevrotain (M. kathygre)
* Sri Lankan spotted chevrotain (M. meminna)

Tragulus|

* Java mouse-deer (T. javanicus)
* Lesser mouse-deer (T. kanchil)
* Greater mouse-deer (T. napu)
* Philippine mouse-deer (T. nigricans)
* Vietnam mouse-deer (T. versicolor)
* Williamson's mouse-deer (T. williamsoni)

Cervidae|

Large family listed below

Bovidae|

Large family listed below

Family Cervidae

| Cervinae|

| Muntiacus|

* Bornean yellow muntjac (M. atherodes)
* Hairy-fronted muntjac (M. crinifrons)
* Fea's muntjac (M. feae)
* Gongshan muntjac (M. gongshanensis)
* Sumatran muntjac (M. montanus)
* Southern red muntjac (M. muntjak)
* Pu Hoat muntjac (M. puhoatensis)
* Leaf muntjac (M. putaoensis)
* Reeves's muntjac (M. reevesi)
* Roosevelt's muntjac (M. rooseveltorum)
* Truong Son muntjac (M. truongsonensis)
* Northern red muntjac (M. vaginalis)
* Giant muntjac (M. vuquangensis)

|

Elaphodus|

* Tufted deer (E. cephalophus)

Dama|

* European fallow deer (D. dama)
* Persian fallow deer (D. mesopotamica)

Axis|

* Chital (A. axis)
* Calamian deer (A. calamianensis)
* Bawean deer (A. kuhlii)
* Hog deer (A. porcinus)

Rucervus|

* Barasingha (R. duvaucelii)
* Eld's deer (R. eldii)

Elaphurus|

* Père David's deer (E. davidianus)

Rusa|

* Visayan spotted deer (R. alfredi)
* Philippine sambar (R. mariannus)
* Rusa deer (R. timorensis)
* Sambar (R. unicolor)

Cervus|

* Thorold's deer (C. albirostris)
* Red deer (C. elaphus)
* Elk (C. canadensis)
* Central Asian red deer (C. hanglu)
* Sika deer (C. nippon)

Capreolinae|

| Alces|

* Moose (A. alces)

|

Hydropotes|

* Water deer (H. inermis)

Capreolus|

* European roe deer (C. capreolus)
* Siberian roe deer (C. pygargus)

Rangifer|

* Reindeer (R. tarandus)

Hippocamelus|

* Taruca (H. antisensis)
* South Andean deer (H. bisulcus)

Mazama|

* Red brocket (M. americana)
* Small red brocket (M. bororo)
* Merida brocket (M. bricenii)
* Dwarf brocket (M. chunyi)
* Gray brocket (M. gouazoubira)
* Pygmy brocket (M. nana)
* Amazonian brown brocket (M. nemorivaga)
* Little red brocket (M. rufina)
* Central American red brocket (M. temama)

Ozotoceros|

* Pampas deer (O. bezoarticus)

Blastocerus|

* Marsh deer (B. dichotomus)

Pudu|

* Northern pudu (P. mephistophiles)?
* Southern pudu (P. pudu)

Pudella?|

* Peruvian Yungas pudu (P. carlae)
* Northern pudu (P. mephistophiles)

Odocoileus|

* Mule deer (O. hemionus)
* Yucatan brown brocket (O. pandora)
* White-tailed deer (O. virginianus)

Family Bovidae

| Hippotraginae|

| Hippotragus|

* Roan antelope (H. equinus)
* Sable antelope (H. niger)

|

Oryx|

* East African oryx (O. beisa)
* Scimitar oryx (O. dammah)
* Gemsbok (O. gazella)
* Arabian oryx (O. leucoryx)

Addax|

* Addax (A. nasomaculatus)

Reduncinae|

| Kobus|

* Waterbuck (K. ellipsiprymnus)
* Kob (K. kob)
* Lechwe (K. leche)
* Nile lechwe (K. megaceros)
* Puku (K. vardonii)

|

Redunca|

* Southern reedbuck (R. arundinum)
* Mountain reedbuck (R. fulvorufula)
* Bohor reedbuck (R. redunca)

Aepycerotinae|

| Aepyceros|

* Impala (A. melampus)

|

Peleinae|

| Pelea|

* Grey rhebok (P. capreolus)

|

Alcelaphinae|

| Beatragus|

* Hirola (B. hunteri)

|

Damaliscus|

* Common tsessebe (D. lunatus)
* Bontebok (D. pygargus)

Alcelaphus|

* Hartebeest (A. buselaphus)

Connochaetes|

* Black wildebeest (C. gnou)
* Blue wildebeest (C. taurinus)

Pantholopinae|

| Pantholops|

* Tibetan antelope (P. hodgsonii)

|

Caprinae|

Large subfamily listed below

Bovinae|

Large subfamily listed below

Antilopinae|

Large subfamily listed below

Family Bovidae (subfamily Caprinae)

| Ammotragus|

* Barbary sheep (A. lervia)

|

Arabitragus|

* Arabian tahr (A. jayakari)

Budorcas|

* Takin (B. taxicolor)

Capra|

* Wild goat (C. aegagrus)
* West Caucasian tur (C. caucasia)
* East Caucasian tur (C. cylindricornis)
* Markhor (C. falconeri)
* Domestic goat (C. hircus)
* Alpine ibex (C. ibex)
* Nubian ibex (C. nubiana)
* Iberian ibex (C. pyrenaica)
* Siberian ibex (C. sibirica)
* Walia ibex (C. walie)

Capricornis|

* Japanese serow (C. crispus)
* Red serow (C. rubidus)
* Mainland serow (C. sumatraensis)
* Taiwan serow (C. swinhoei)

Hemitragus|

* Himalayan tahr (H. jemlahicus)

Naemorhedus|

* Red goral (N. baileyi)
* Long-tailed goral (N. caudatus)
* Himalayan goral (N. goral)
* Chinese goral (N. griseus)

Oreamnos|

* Mountain goat (O. americanus)

Ovibos|

* Muskox (O. moschatus)

Nilgiritragus|

* Nilgiri tahr (N. hylocrius)

Ovis|

* Argali (O. ammon)
* Domestic sheep (O. aries)
* Bighorn sheep (O. canadensis)
* Dall sheep (O. dalli)
* Mouflon (O. gmelini)
* Snow sheep (O. nivicola)
* Urial (O. vignei)

Pseudois|

* Bharal (P. nayaur)

Rupicapra|

* Pyrenean chamois (R. pyrenaica)
* Chamois (R. rupicapra)

Family Bovidae (subfamily Bovinae)

| Boselaphini|

| Tetracerus|

* Four-horned antelope (T. quadricornis)

|

Boselaphus|

* Nilgai (B. tragocamelus)

Bovini|

| Bubalus|

* Wild water buffalo (B. arnee)
* Domestic water buffalo (B. bubalis)
* Lowland anoa (B. depressicornis)
* Tamaraw (B. mindorensis)
* Mountain anoa (B. quarlesi)

|

Bos|

* American bison (B. bison)
* European bison (B. bonasus)
* Bali cattle (B. domesticus)
* Gayal (B. frontalis)
* Gaur (B. gaurus)
* Domestic yak (B. grunniens)
* Zebu (B. indicus)
* Banteng (B. javanicus)
* Wild yak (B. mutus)
* Cattle (B. taurus)

Pseudoryx|

* Saola (P. nghetinhensis)

Syncerus|

* African buffalo (S. caffer)

Tragelaphini|

| Tragelaphus
(including kudus)|

* Nyala (T. angasii)
* Mountain nyala (T. buxtoni)
* Bongo (T. eurycerus)
* Lesser kudu (T. imberbis)
* Harnessed bushbuck (T. scriptus)
* Sitatunga (T. spekeii)
* Greater kudu (T. strepsiceros)
* Cape bushbuck (T. sylvaticus)

|

Taurotragus|

* Giant eland (T. derbianus)
* Common eland (T. oryx)

Family Bovidae (subfamily Antilopinae)

| Antilopini|

| Ammodorcas|

* Dibatag (A. clarkei)

|

Antidorcas|

* Springbok (A. marsupialis)

Antilope|

* Blackbuck (A. cervicapra)

Eudorcas|

* Mongalla gazelle (E. albonotata)
* Red-fronted gazelle (E. rufifrons)
* Thomson's gazelle (E. thomsonii)
* Heuglin's gazelle (E. tilonura)

Gazella|

* Chinkara (G. bennettii)
* Cuvier's gazelle (G. cuvieri)
* Dorcas gazelle (G. dorcas)
* Erlanger's gazelle (G. erlangeri)
* Mountain gazelle (G. gazella)
* Rhim gazelle (G. leptoceros)
* Speke's gazelle (G. spekei)
* Goitered gazelle (G. subgutturosa)

Litocranius|

* Gerenuk (L. walleri)

Nanger|

* Dama gazelle (N. dama)
* Grant's gazelle (N. granti)
* Bright's gazelle (N. notatus)
* Peter's gazelle (N. petersii)
* Soemmerring's gazelle (N. soemmerringii)

Procapra|

* Mongolian gazelle (P. gutturosa)
* Goa (P. picticaudata)
* Przewalski's gazelle (P. przewalskii)

Saigini|

| Saiga|

* Saiga antelope (S. tatarica)

|

Neotragini|

| Dorcatragus|

* Beira (D. megalotis)

|

Madoqua|

* Günther's dik-dik (M. guentheri)
* Kirk's dik-dik (M. kirkii)
* Silver dik-dik (M. piacentinii)
* Salt's dik-dik (M. saltiana)

Neotragus|

* Royal antelope (N. pygmaeus)

Nesotragus|

* Bates' pygmy antelope (N. batesi)
* Suni (N. moschatus)

Oreotragus|

* Klipspringer (O. oreotragus)

Ourebia|

* Oribi (O. ourebi)

Raphicerus|

* Steenbok (R. campestris)
* Cape grysbok (R. melanotis)
* Sharpe's grysbok (R. sharpei)

Cephalophini|

| Cephalophus|

* Aders's duiker (C. adersi)
* Brooke's duiker (C. brookei)
* Peters' duiker (C. callipygus)
* White-legged duiker (C. crusalbum)
* Bay duiker (C. dorsalis)
* Harvey's duiker (C. harveyi)
* Jentink's duiker (C. jentinki)
* White-bellied duiker (C. leucogaster)
* Red forest duiker (C. natalensis)
* Black duiker (C. niger)
* Black-fronted duiker (C. nigrifrons)
* Ogilby's duiker (C. ogilbyi)
* Ruwenzori duiker (C. rubidis)
* Red-flanked duiker (C. rufilatus)
* Yellow-backed duiker (C. silvicultor)
* Abbott's duiker (C. spadix)
* Weyns's duiker (C. weynsi)
* Zebra duiker (C. zebra)

|

Philantomba|

* Blue duiker (P. monticola)
* Maxwell's duiker (P. maxwellii)
* Walter's duiker (P. walteri)

Sylvicapra|

* Common duiker (S. grimmia)

Suborder Suina

| Suidae|

| Babyrousa|

* Buru babirusa (B. babyrussa)
* North Sulawesi babirusa (B. celebensis)
* Togian babirusa (B. togeanensis)

|

Hylochoerus|

* Giant forest hog (H. meinertzhageni)

Phacochoerus|

* Desert warthog (P. aethiopicus)
* Common warthog (P. africanus)

Porcula|

* Pygmy hog (P. salvania)

Potamochoerus|

* Bushpig (P. larvatus)
* Red river hog (P. porcus)

Sus|

* Palawan bearded pig (S. ahoenobarbus)
* Bornean bearded pig (S. barbatus)
* Visayan warty pig (S. cebifrons)
* Celebes warty pig (S. celebensis)
* Domestic pig (S. domesticus)
* Flores warty pig (S. heureni)
* Oliver's warty pig (S. oliveri)
* Philippine warty pig (S. philippensis)
* Wild boar (S. scrofa)
* Timor warty pig (S. timoriensis)
* Javan warty pig (S. verrucosus)

Tayassuidae|

| Tayassu|

* White-lipped peccary (T. pecari)

|

Catagonus|

* Chacoan peccary (C. wagneri)

Dicotyles|

* Collared peccary (D. tajacu)

Suborder Tylopoda

| Camelidae|

| Lama|

* Llama (L. glama)
* Guanaco (L. guanicoe)
* Alpaca (L. pacos)
* Vicuña (L. vicugna)

|

Camelus|

* Domestic Bactrian camel (C. bactrianus)
* Dromedary/Arabian camel (C. dromedarius)
* Wild Bactrian camel (C. ferus)

Suborder Whippomorpha

| Hippopotamidae|

| Hippopotamus|

* Hippopotamus (H. amphibius)

|

Choeropsis|

* Pygmy hippopotamus (C. liberiensis)

Cetacea|

* see Cetacea

Taxon identifiers

Giraffa camelopardalis|

* Wikidata: Q15083
* Wikispecies: Giraffa camelopardalis
* ADW: Giraffa_camelopardalis
* BOLD: 73517
* CoL: 3G3KN
* CMS: giraffa-camelopardalis
* EoL: 308378
* EPPO: GIRFCA
* GBIF: 2441205
* iNaturalist: 42157
* IRMNG: 10874588
* ITIS: 625036
* IUCN: 9194
* MDD: 1006340
* MSW: 14200476
* NCBI: 9894
* Observation.org: 81428
* Open Tree of Life: 768674
* Paleobiology Database: 133600
* Species+: 98186
* Xeno-canto: Giraffa-camelopardalis

Cervus camelopardalis|

* Wikidata: Q110259076
* GBIF: 7529494

Authority control databases: National |

* United States
* France
* BnF data
* Japan
* Israel

|

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