Prehistoric Caucasus

Human habitation of the Caucasus region before written records

The Caucasus region, on the gateway between Southwest Asia and Europe, plays a pivotal role in the peopling of Eurasia, possibly as early as during the Homo erectus expansion to Eurasia, in the Upper Paleolithic peopling of Europe, and again in the re-peopling Mesolithic Europe following the Last Glacial Maximum, and in the expansion associated with the Neolithic Revolution.

## Lower to Middle Paleolithic

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Dmanisi skull 5, found in Dmanisi, Georgia, is among the earliest Homo erectus fossils, dated to 1.8 Ma.

* Azykh Cave has remnants of the pre-Acheulean, estimated at 0.7 Ma.

Further information: Neanderthals in Southwest Asia

* Mousterian
* Mezmaiskaya cave (70–40 ka)

## Upper Paleolithic to Epipaleolithic

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* Dzudzuana cave (30 ka)
* Satsurblia cave (24 ka)
* Damjili Cave
* Dash Salakhly (20 ka)

Further information: Epipalaeolithic (Levant) and Mesolithic Europe

Further information: Caucasian Hunter-Gatherer

* Gobustan National Park (20–5 ka)
* Trialetian (16–8 ka)

## Neolithic to Iron Age

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Neolithic:

* Shulaveri-Shomu culture (8–6 ka)
* Metsamor site (6 ka)

Bronze Age:

* Maykop culture
* Leyla-Tepe culture
* Kura-Araxes culture
* Trialeti culture
* Jar-Burial Culture
* Kurgan culture
* Khojaly–Gadabay culture (c. 1300 – 600 BC)
* Kingdom of Arme-Shupria (c. 1300 – 1190 BC)
* Colchian culture (c. 2700 – 700 BC[1])
* Koban culture (c. 1100 – 400 BC)

The South Caucasus gradually enters the historical period following the Bronze Age collapse, see history of the Caucasus

* Kingdom of Diauehi (12th – 9th century BC)
* Nairi (1114 – 860 BC)
* Kingdom of Urartu (c. 860 – 590 BC)
* Neo-Assyrian Empire (911 – 609 BC)

## Genetic history

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Main article: Caucasus hunter-gatherer

Further information: Peoples of the Caucasus and Y-DNA haplogroups in populations of the Caucasus

See also: Genetic history of Europe

Language groups in the Caucasus have been found to have a close correlation to genetic ancestry.[2]

A genetic study in 2015 by Jones et al. identified a previously unidentified lineage, which was dubbed Caucasian Hunter-Gatherer (CHG).[3] The study detected a split between CHG and so-called "Western European Hunter-Gatherer" (WHG) lineages, about 45,000 years ago, the presumed time of the original peopling of Europe. CHG separated from the "early Anatolian farmers" (EAF) lineage later, at 25,000 years ago, during the Last Glacial Maximum. (CHG was extrapolated from, among other sources, the genomes of two fossils from western Georgia – one about 13,300 years old (Late Upper Paleolithic) and the other 9,700 years (Mesolithic), which were compared to the 13,700 year-old Bichon man genome (found in Switzerland).

A genetic study in 2020 analyzing samples from Klin-Yar communities, including the Koban culture, found that the ancient population had one sample of paternal Haplogroup D-Z27276. Most other paternal lineages in the study were Haplogroup J1 and Haplogroup G-M285.[4]

## See also

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* Prehistoric Georgia
* Prehistoric Armenia
* Prehistoric Azerbaijan
* Scythia
* Peoples of the Caucasus
* Proto-Northwest Caucasian language
* Caucasic languages
* Dené-Caucasian

## References

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1. ^ Antonio Sagona, chapter 10 A World Apart : Colchian Culture. The Archaeology of the Caucasus. From Earliest Settlements to the Iron Age. Cambridge University Press. [1]
2. ^ O.Balanovsky et al., "Parallel Evolution of Genes and Languages in the Caucasus Region", Mol Biol Evol00 (2011), doi:10.1093/molbev/msr126.
3. ^ Jones, Eppie; Gloria, Gonzalez-Fortes; Manica, Andrea; Pinhasi, Ron; Bradley, Dan (2015). "Upper Palaeolithic genomes reveal deep roots of modern Eurasians". Nature Communications. 6: 8912. doi:10.1038/ncomms9912. PMC 4660371. PMID 26567969.
4. ^ Boulygina, Eugenia; Tsygankova, Svetlana; Sharko, Fedor; Slobodova, Natalia; Gruzdeva, Natalia; Rastorguev, Sergey; Belinsky, Andrej; Härke, Heinrich; Kadieva, Anna; Demidenko, Sergej; Shvedchikova, Tatiana (2020-06-01). "Mitochondrial and Y-chromosome diversity of the prehistoric Koban culture of the North Caucasus". Journal of Archaeological Science: Reports. 31: 102357. doi:10.1016/j.jasrep.2020.102357. ISSN 2352-409X. S2CID 218789467.