Pycnonotus

Genus of birds

Pycnonotus

Olive-winged bulbul (Pycnonotus plumosus)
Scientific classification
Domain:  | Eukaryota
Kingdom:  | Animalia
Phylum:  | Chordata
Class:  | Aves
Order:  | Passeriformes
Infraorder:  | Passerides
Family:  | Pycnonotidae
Genus:  | Pycnonotus
F. Boie, 1826
Type species
Turdus capensis (Cape bulbul)

Linnaeus, 1766

Synonyms

* Brachypus
* Haematornis
* Kelaartia
* Micropus
* Molpastes
* Oreoctistes
* Otocompsa
* Xanthiscus

Yellow-vented bulbul (Pycnonotus goiavier)

Pycnonotus is a genus of frugivorous passerine birds in the bulbul family Pycnonotidae.

## Taxonomy and systematics[edit]

The genus Pycnonotus was introduced by the German zoologist Friedrich Boie in 1826 with the Cape bulbul as the type species.[1] The name of the genus combines the Ancient Greek words puknos "thick" or "compact" and -nōtos "-backed".[2]

The genus contains the following 34 species:[3]

* Cream-vented bulbul (Pycnonotus simplex)
* Olive-winged bulbul (Pycnonotus plumosus)
* Asian red-eyed bulbul (Pycnonotus brunneus)
* Straw-headed bulbul (Pycnonotus zeylanicus)
* Spot-necked bulbul (Pycnonotus tympanistrigus)
* Cream-eyed bulbul (Pycnonotus pseudosimplex)
* Ashy-fronted bulbul (Pycnonotus cinereifrons)
* White-browed bulbul (Pycnonotus luteolus)
* Ayeyarwady bulbul (Pycnonotus blanfordi)
* Streak-eared bulbul (Pycnonotus conradi)
* Stripe-throated bulbul (Pycnonotus finlaysoni)
* Pale-eyed bulbul (Pycnonotus davisoni)
* Flavescent bulbul (Pycnonotus flavescens)
* Aceh bulbul (Pycnonotus snouckaerti)
* Orange-spotted bulbul (Pycnonotus bimaculatus)
* Pale-faced bulbul (Pycnonotus leucops)
* Yellow-throated bulbul (Pycnonotus xantholaemus)
* Yellow-eared bulbul (Pycnonotus penicillatus)
* Brown-breasted bulbul (Pycnonotus xanthorrhous)
* Light-vented bulbul (Pycnonotus sinensis)
* Styan's bulbul (Pycnonotus taivanus)
* Red-whiskered bulbul (Pycnonotus jocosus)
* Yellow-vented bulbul (Pycnonotus goiavier)
* Red-vented bulbul (Pycnonotus cafer)
* Sooty-headed bulbul (Pycnonotus aurigaster)
* White-eared bulbul (Pycnonotus leucotis)
* Himalayan bulbul (Pycnonotus leucogenys)
* White-spectacled bulbul (Pycnonotus xanthopygos)
* African red-eyed bulbul (Pycnonotus nigricans)
* Common bulbul (Pycnonotus barbatus)
* Dodson's bulbul (Pycnonotus dodsoni)
* Somali bulbul (Pycnonotus somaliensis)
* Dark-capped bulbul (Pycnonotus tricolor)
* Cape bulbul (Pycnonotus capensis)

### Former species[edit]

In previous circumscriptions the genus Pycnonotus was considerably larger. Recent taxonomic revisions have seen many species transferred to other genera.

In 2010, eighteen former Pycnonotus species were reclassified into different genera, either directly from Pycnonotus or from the genus Andropadus, to which they had already been transferred by some authorities. These changes were as follows:

* one species was transferred to genus Stelgidillas.[4]
* Slender-billed greenbul ( former Pycnonotus gracilirostri or Pycnonotus gracilirostris)[5]
* twelve species were transferred to genus Arizelocichla:
* Shelley's greenbul (former Pycnonotus masukuensis)[6]
* Kakamega greenbul (nominate) (former Pycnonotus kakamegae)[7]
* Cameroon greenbul (former Pycnonotus montanus, as Arizelocichla montana)[8]
* Western greenbul (former Pycnonotus tephrolaemus)[9]
* Olive-breasted greenbul (former Pycnonotus tephrolaema)
* Mountain greenbul (former Pycnonotus nigriceps)[10]
* Uluguru greenbul (former Pycnonotus neumanni)
* Black-browed greenbul (former Pycnonotus fusciceps)
* Yellow-throated greenbul (former Pycnonotus chlorigula)[11]
* Stripe-cheeked greenbul (former Pycnonotus milanjensis)[12]
* Olive-headed greenbul (former Pycnonotus olivaceiceps)[13]
* Stripe-faced greenbul (former Pycnonotus strifacies)
* five species were transferred to genus Eurillas:
* Little greenbul (former Pycnonotus virens)[14]
* Little grey greenbul (former Pycnonotus gracilis)[15]
* Ansorge's greenbul (former Pycnonotus ansorgei)[16]
* Plain greenbul (former Pycnonotus curvirostris)[17]
* Yellow-whiskered greenbul (former Pycnonotus latirostris)[18]

In 2020, a further 17 species were transferred to other genera:[3][19]

* Black-and-white bulbul (former Pycnonotus melanoleucos, moved to the monotypic genus Microtarsus)
* Puff-backed bulbul (former Pycnonotus eutilotus, moved to the monotypic genus Eutilotus)
* Yellow-wattled bulbul (former Pycnonotus urostictus, moved to monotypic genus Poliolophus)
* Two species to genus Alcurus:
* Striated bulbul (former Pycnonotus striatus)
* Spot-necked bulbul (former Pycnonotus tympanistrigus)
* Four species to genus Brachypodius:
* Grey-headed bulbul (former Pycnonotus priocephalus)
* Black-headed bulbul (former Pycnonotus atriceps, as Brachypodius melanocephalos; the species epithet melanocephalos has priority over atriceps[3])
* Andaman bulbul (former Pycnonotus fuscoflavescens)
* Blue-wattled bulbul (former Pycnonotus nieuwenhuisii)
* Three species to genus Ixodia:
* Spectacled bulbul (former Pycnonotus erythropthalmos)
* Grey-bellied bulbul (former Pycnonotus cyaniventris)
* Scaly-breasted bulbul (former Pycnonotus squamatus)
* Five species to genus Rubigula:
* Black-crested bulbul (former Pycnonotus flaviventris)
* Flame-throated bulbul (former Pycnonotus gularis)
* Black-capped bulbul (former Pycnonotus melanicterus)
* Ruby-throated bulbul (former Pycnonotus dispar)
* Bornean bulbul (former Pycnonotus montis)

Other former Pycnonotus species include:

* Sombre greenbul (former Pycnonotus importunus, now Andropadus importunus)[20]
* Yellow-throated leaflove (former Haematornis flavicollis or Pycnonotus flavicollis, now Atimastillas flavicollis)[21][22]
* Hairy-backed bulbul (former Brachypodius criniger, now Tricholestes criniger)[23]
* Bare-faced bulbul (former Pycnonotus hualon, now Nok hualon)[24]
* Cream-striped bulbul (former Pycnonotus leucogrammicus, now Hemixos leucogrammicus)[25]

## References[edit]

1. ^ Boie, Friedrich (1826). "Generalübersicht". Isis von Oken (in German). 19. Col 973.
2. ^ Jobling, James A. (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 325\. ISBN 978-1-4081-2501-4.
3. ^ a b c Gill, Frank; Donsker, David; Rasmussen, Pamela, eds. (July 2023). "Bulbuls". IOC World Bird List Version 13.2. International Ornithologists' Union. Retrieved 23 July 2023.
4. ^ "Taxonomy Version 2 « IOC World Bird List". www.worldbirdnames.org. Retrieved 2017-04-02.
5. ^ "Stelgidillas gracilirostris - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-07.
6. ^ "Arizelocichla masukuensis - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-02.
7. ^ "Arizelocichla kakamegae kakamegae - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-02.
8. ^ "Arizelocichla montana - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-03.
9. ^ "Arizelocichla tephrolaema - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-03.
10. ^ "Arizelocichla nigriceps nigriceps - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-07.
11. ^ "Arizelocichla chlorigula - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-06.
12. ^ "Arizelocichla milanjensis - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-06.
13. ^ "Arizelocichla olivaceiceps - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-06.
14. ^ "Eurillas virens [incl. hallae] \- Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-07.
15. ^ "Eurillas gracilis - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-09.
16. ^ "Eurillas ansorgei - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-09.
17. ^ "Eurillas curvirostris - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-09.
18. ^ "Eurillas latirostris - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-10.
19. ^ Gill, F.; Donsker, D.; Rasmussen, P. (eds.). "Taxonomic Updates". IOC World Bird List. International Ornithological Congress. Retrieved 3 August 2020.
20. ^ "Andropadus importunus - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-10.
21. ^ "Atimastillas flavicollis - Avibase". avibase.bsc-eoc.org. Retrieved 2017-04-16.
22. ^ Catalogue of the Birds in the British Museum: Passeriformes, or perching birds. Cichlomorphœ: pt. III-IV, containing the ... family Timeliidœ (babbling-thrushes) by R.B. Sharpe. order of the Trustees. 1881-01-01.
23. ^ "Tricholestes criniger - Avibase". avibase.bsc-eoc.org. Retrieved 2017-06-02.
24. ^ Fuchs, Jérôme; Pasquet, Eric; Stuart, Bryan L.; Woxvold, Iain A.; Duckworth, J. W.; Bowie, Rauri C. K. (2018). "Phylogenetic affinities of the enigmatic Bare-faced Bulbul Pycnonotus hualon with description of a new genus". Ibis. 160 (3): 659–665. doi:10.1111/ibi.12580. ISSN 1474-919X.
25. ^ Shakya, Subir B.; Sheldon, Frederick H. (2017). "The phylogeny of the world's bulbuls (Pycnonotidae) inferred using a supermatrix approach". Ibis. 159 (3): 498–509. doi:10.1111/ibi.12464. ISSN 0019-1019.

## External links[edit]

* Media related to Pycnonotus at Wikimedia Commons

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Genera of passerides and their extinct allies

* Kingdom: Animalia
* Phylum: Chordata
* Class: Aves
* Suborder: Passeri
* Infraorder: Passerida

Passerida

| Chaetopidae?|

* Chaetops

|

Chloropseidae?|

* Chloropsis

Hyliotidae?|

* Hyliota

Irenidae|

* Irena

Paridae|

* Baeolophus
* Cyanistes
* Cephalopyrus
* Lophophanes
* Machlolophus
* Melaniparus
* Melanochlora
* Pardaliparus
* Parus
* Periparus
* Poecile
* Pseudopodoces
* Sittiparus
* Sylviparus

Picathartidae?|

* Picathartes

Promeropidae?|

* Promerops

Remizidae|

* Anthoscopus
* Auriparus
* Remiz

Stenostiridae|

* Chelidorhynx
* Culicicapa
* Elminia
* Stenostira

Muscicapida|

*     * See below ↓

Sylvioidea|

*     * See below ↓

Passeroidea|

*     * See Passeroidea

Muscicapida

| Regulidae|

* Corthylio
* Regulus

|

Bombycilloidea|

| Bombycillidae|

* Bombycilla

|

Dulidae|

* Dulus

Hylocitreidae|

* Hylocitrea

Hypocoliidae|

* Hypocolius

†Mohoidae|

* Chaetoptila
* Moho

Ptiliogonatidae|

* Phainopepla
* Phainoptila
* Ptiliogonys

Certhioidea|

| incertae sedis|

* †Certhiops
* †Kischinskinia

|

Certhiidae|

* Certhia
* Salpornis

Polioptilidae|

* Microbates
* Polioptila
* Ramphocaenus

Sittidae|

* Sitta

Tichodromidae|

* Tichodroma

Troglodytidae|

* Campylorhynchus
* Cantorchilus
* Catherpes
* Cinnycerthia
* Cistothorus
* Cyphorhinus
* Ferminia
* Henicorhina
* Hylorchilus
* Microcerculus
* Odontorchilus
* Pheugopedius
* Salpinctes
* Thryomanes
* Thryophilus
* Thryorchilus
* Thryothorus
* Troglodytes
* Uropsila

Muscicapoidea|

| Buphagidae|

* Buphagus

|

Cinclidae|

* Cinclus

Elachuridae|

* Elachura

Mimidae|

* Allenia
* Cinclocerthia
* Dumetella
* Margarops
* Melanotis
* Mimus
* Oreoscoptes
* Ramphocinclus
* Toxostoma

Muscicapidae|

| Erithacinae|

* Chamaetylas
* Cichladusa
* Cossypha
* Cossyphicula
* Dessonornis
* Erithacus
* Pogonocichla
* Sheppardia
* Stiphrornis
* Swynnertonia
* Xenocopsychus

|

Muscicapinae|

| Copsychini|

* Alethe
* Cercotrichas
* Copsychus

|

Muscicapini|

* Agricola
* Bradornis
* Empidornis
* Fraseria
* Humblotia
* Melaenornis
* Muscicapa
* Namibornis
* Sigelus

Niltavinae|

* Anthipes
* Cyanoptila
* Cyornis
* Eumyias
* Leucoptilon
* Niltava
* Sholicola

Saxicolinae|

* Brachypteryx
* Calliope
* Campicoloides
* Cinclidium
* Emarginata
* Enicurus
* Ficedula
* Heinrichia
* Heteroxenicus
* Irania
* Larvivora
* Leonardina
* Luscinia
* Monticola
* Myiomela
* Myophonus
* Myrmecocichla
* Oenanthe
* Phoenicurus
* Pinarochroa
* Saxicola
* Tarsiger
* Thamnolaea
* Vauriella

Sturnidae|

* Acridotheres
* Agropsar
* Ampeliceps
* Aplonis
* Arizelopsar
* Basilornis
* Cinnyricinclus
* Creatophora
* †Cryptopsar
* Enodes
* †Fregilupus
* Goodfellowia
* Gracula
* Gracupica
* Grafisia
* Hartlaubius
* Hylopsar
* Lamprotornis
* Leucopsar
* Mino (bird)
* †Necropsar
* Neocichla
* Notopholia
* Onychognathus
* Pastor
* Pholia
* Poeoptera
* Rhabdornis
* Sarcops
* Saroglossa
* Scissirostrum
* Speculipastor
* Spodiopsar
* Streptocitta
* Sturnia
* Sturnornis
* Sturnus

Turdidae|

* †Meridiocichla
* Pinarornis

Myadestinae|

* Grandala
* Myadestes
* Neocossyphus
* Sialia
* Stizorhina

Turdinae|

* Catharus
* Chlamydochaera
* Cichlopsis
* Cochoa
* Entomodestes
* Geokichla
* Hylocichla
* Ridgwayia
* Ixoreus
* Turdus
* Zoothera

Sylvioidea

| Acrocephalidae|

* Acrocephalus
* Calamonastides
* Chloropeta
* Hippolais
* Iduna
* Nesillas

|

Aegithalidae|

* Aegithalos
* Leptopoecile
* Psaltriparus

Alaudidae|

| Alaudinae|

* Alauda
* Alaudala
* Calandrella
* Chersophilus
* Eremalauda
* Eremophila
* Galerida
* Lullula
* Melanocorypha
* Spizocorys

|

Certhilaudinae|

* Alaemon
* Ammomanes
* Ammomanopsis
* Certhilauda
* Chersomanes
* Eremopterix
* Pinarocorys
* Ramphocoris

Mirafrinae|

* Calendulauda
* Heteromirafra
* Mirafra

Alcippeidae|

* Alcippe

Bernieridae|

* Bernieria
* Crossleyia
* Cryptosylvicola
* Hartertula
* Oxylabes
* Randia
* Thamnornis
* Xanthomixis

Cettiidae|

* Abroscopus
* Cettia
* Hemitesia
* Horornis
* Phyllergates
* Tesia
* Tickellia
* Urosphena

Cisticolidae|

* Apalis
* Artisornis
* Bathmocercus
* Calamonastes
* Camaroptera
* Cisticola
* Drymocichla
* Eminia
* Eremomela
* Euryptila
* Hypergerus
* Incana
* Malcorus
* Micromacronus
* Neomixis
* Oreolais
* Oreophilais
* Orthotomus
* Phragmacia
* Phyllolais
* Poliolais
* Prinia
* Scepomycter
* Schistolais
* Spiloptila
* Urolais

Donacobiidae|

* Donacobius

Erythrocercidae|

* Erythrocercus

Hirundinidae|

* Alopochelidon
* Atronanus
* Atticora
* Cecropis
* Cheramoeca
* Delichon
* Hirundo
* Neophedina
* Orochelidon
* Petrochelidon
* Phedina
* Phedinopsis
* Progne
* Psalidoprocne
* Pseudhirundo
* Pseudochelidon
* Ptyonoprogne
* Pygochelidon
* Riparia
* Stelgidopteryx
* Tachycineta

Hyliidae|

* Hylia
* Pholidornis

Leiothrichidae|

* Actinodura
* Argya
* Cutia
* Garrulax
* Grammatoptila
* Heterophasia
* Ianthocincla
* Laniellus
* Leioptila
* Leiothrix
* Liocichla
* Minla
* Montecincla
* Pterorhinus
* Trochalopteron
* Turdoides

Locustellidae|

* Bradypterus
* Catriscus
* Cincloramphus
* Elaphrornis
* Helopsaltes
* Locustella
* Malia
* Megalurus
* Poodytes
* Robsonius
* Schoenicola

Macrosphenidae|

* Achaetops
* Cryptillas
* Macrosphenus
* Melocichla
* Sphenoeacus
* Sylvietta

Nicatoridae|

* Nicator

Panuridae|

* Panurus

Paradoxornithidae|

* Chamaea
* Chrysomma
* Fulvetta
* Lioparus
* Moupinia
* Myzornis
* Paradoxornis
* Rhopophilus
* Suthora

Pellorneidae|

* Gampsorhynchus
* Graminicola
* Gypsophila
* Illadopsis
* Kenopia
* Laticilla
* Malacocincla
* Malacopteron
* Napothera
* Pellorneum
* Ptilocichla
* Schoeniparus
* Turdinus

Phylloscopidae|

* Phylloscopus

Pnoepygidae|

* Pnoepyga

Pycnonotidae|

* Acritillas
* Alcurus
* Alophoixus
* Altimastillas
* Andropadus
* Arizelocichla
* Baeopogon
* Bleda
* Brachypodius
* Calyptocichla
* Chlorocichla
* Criniger
* Euptilotus
* Eurillas
* Hemixos
* Hypsipetes
* Iole
* Ixodia
* Ixonotus
* Ixos
* Microtarsus
* Neolestes
* Nok
* Phyllastrephus
* Poliolophus
* Pycnonotus
* Rubigula
* Setornis
* Spizixos
* Stelgidillas
* Thescelocichla
* Tricholestes

Scotocercidae|

* Scotocerca

Sylviidae|

* Curruca
* Sylvia

Timaliidae|

* Cyanoderma
* Dumetia
* Erythrogenys
* Macronus
* Melanocichla
* Mixornis
* Pomatorhinus
* Spelaeornis
* Stachyris
* Timalia

Zosteropidae|

* Apalopteron
* Cleptornis
* Dasycrotapha
* Heleia
* Megazosterops
* Parayuhina
* Rukia
* Staphida
* Sterrhoptilus
* Tephrozosterops
* Yuhina
* Zosterops
* Zosterornis

Taxon identifiers

Pycnonotus|

* Wikidata: Q940685
* Wikispecies: Pycnonotus
* AFD: Pycnonotus
* Avibase: 627F72C637213318
* BioLib: 30880
* BOLD: 95548
* CoL: 75Q7
* EPPO: 1PYKNG
* Fauna Europaea: 97378
* Fauna Europaea (new): 6055f2bc-bdc6-48c7-8e4b-4e4f5de21d5f
* GBIF: 2486114
* iNaturalist: 14587
* IRMNG: 1391209
* ITIS: 178505
* NBN: NHMSYS0000533735
* NCBI: 125282
* NZOR: 1740b8e4-e7d2-4c24-bee2-3cceb7a95cf0
* Open Tree of Life: 92678
* Plazi: 997D6F12-68EE-0D3A-B3AD-C30BA3D1884D
* WoRMS: 204987
* ZooBank: 4D8A16BA-20F0-49F3-869C-B5FFD605034A

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