Sri Lankan leopard

Leopard subspecies

Sri Lankan leopard

Sri Lankan leopard in Yala National Park

Conservation status

Vulnerable  (IUCN 3.1)[1]

Scientific classification
Kingdom:  | Animalia
Phylum:  | Chordata
Class:  | Mammalia
Order:  | Carnivora
Suborder:  | Feliformia
Family:  | Felidae
Subfamily:  | Pantherinae
Genus:  | Panthera
Species:  |

P. pardus

Subspecies:  |

P. p. kotiya

Trinomial name
Panthera pardus kotiya

Deraniyagala, 1956

The Sri Lankan leopard (Panthera pardus kotiya) is a leopard subspecies native to Sri Lanka. It was first described in 1956 by Sri Lankan zoologist Paules Edward Pieris Deraniyagala.[2]

Since 2020, the Sri Lankan leopard has been listed as Vulnerable on the IUCN Red List, as the population is estimated at less than 800 mature individuals, and is probably declining.[1]

## Characteristics

[edit]

Female in Yala National Park

The Sri Lankan leopard has a tawny or rusty yellow coat with dark spots and close-set rosettes. Seven females measured in the early 20th century averaged a weight of 64 lb (29 kg) and had a mean head-to-body-length of 1.04 m (3 ft 5 in) with a 77.5 cm (2 ft 6.5 in) long tail, the largest being 1.14 m (3 ft 9 in) with a 84 cm (2 ft 9 in) long tail; 11 males averaged 56 kg (124 lb), the largest being 77 kg (170 lb), and measured 1.27 m (4 ft 2 in) with a 86 cm (2 ft 10 in) long tail, the largest being 1.42 m (4 ft 8 in) with a 97 cm (3 ft 2 in) long tail.[3] The Sri Lankan leopard has possibly evolved to become a rather large leopard subspecies, because it is the apex predator in the country. Large males have been suggested to reach almost 100 kg (220 lb), but evidence for this is lacking.[4]

Melanistic leopards are rare. Few records exist, including from Mawuldeniya, Pitadeniya, and Nallathanniya.[5] In October 2019, the Department of Wildlife Conservation recorded live footage of a melanistic individual for the first time, a male.[6]

## Distribution and habitat

[edit]

The Sri Lankan leopard is still found in all habitats throughout the island in both protected and unprotected areas.[7] These habitat types can be broadly categorised into:[8]

* Arid zone with <1,000 mm (39 in) rainfall
* Dry zone with 1,000–2,000 mm (39–79 in) rainfall
* Wet zone with >2,000 mm (79 in) rainfall

In Sri Lanka's central hills, leopards have been recorded in forest patches, tea estates, grasslands, home gardens, and pine and eucalyptus plantations.[9]

## Ecology and behaviour

[edit]

Leopard attacking an Asian water buffalo calf

In Yala National Park, the Sri Lankan leopard as in other locations, is a solitary hunter, with the exception of females with young. Male's ranges typically overlap the smaller ranges of several females, as well as portions of the ranges of neighboring males, although exclusive core areas are apparent. They are more active and prefer hunting at night, but are also somewhat active during dawn, dusk, and daytime hours. They do not usually store their kills in trees, perhaps because similarly-sized or larger carnivores are absent in Sri Lanka. In 2001 to 2002, adult resident leopard density was estimated at 12.1 mature individuals and 21.7 individuals of all ages per 100 km2 (39 sq mi) in Block I of Yala National Park.[10]

The Sri Lankan leopard hunts by silently stalking its prey, until it is within striking distance, when it unleashes a burst of speed to quickly pursue and pounce on its victim. The prey is usually dispatched with a single bite to the throat. Like most cats, it is pragmatic in its choice of diet, which can include small mammals, birds, and reptiles, as well as larger animals.[1] Sri Lankan axis deer make up the majority of its diet in the dry zone.[10] The animal also preys on sambar, barking deer, wild boar, and monkeys.[11]

No birth season or peak is apparent, with births occur across the year.[10] Its lifespan ranges from 12 to 15 years in the wild, and up to 22 years in captivity.[12]

The leopard is sympatric with the Sri Lankan sloth bear.[13][14]

## Threats

[edit]

The survival of the Sri Lankan leopard is primarily threatened by increasing habitat loss and fragmentation, together with an increasing risk of human-induced mortality. Leopards are killed by people either accidentally in wire snares set for other species, or as retaliation after livestock depredation (usually through poisoning the livestock carcass). They are also occasionally shot. Since 2010, over 90 leopards are known to have been killed by people in Sri Lanka.[1]

Three individuals were killed by snare traps in the Sinharaja conservation area, one of which is stuffed and displayed at the Giritale Wildlife Museum.[15] In May 2020, the melanistic leopard filmed in 2019 was found caught in a snare at the Lakshapana Estate in Nallathanniya, Hatton. Later, it was transported to Elephant Transit Home in Udawalawa for treatment, where it died. The snare had heavily injured its neck.[16][17] Even in large, contiguous protected areas, human encroachment in the border areas is impacting leopard distribution and reducing the effective size of these protected areas.[18]

## Conservation

[edit]

Ongoing research into the Sri Lankan leopard is needed to ensure that conservation measures are targeted and effective. The Leopard Project under the Wilderness and Wildlife Conservation Trust (WWCT), founded by Anjali Watson,[19] and Dr. Andrew Kittle is working closely with the Government of Sri Lanka to ensure this occurs. The Sri Lanka Wildlife Conservation Society will also undertake some studies. The WWCT is engaged throughout the island with targeted work ongoing in the central hills region where fragmentation of the leopard's habitat is rapidly occurring.[20]

### In captivity

[edit]

As of December 2011, 75 captive Sri Lankan leopards were in zoos worldwide. Within the European Endangered Species Programme, 27 male, 29 female and 8 unsexed individuals are kept.[21]

## See also

[edit]

* Cats portal

* Leopard subspecies
* Chinese leopard
* Zanzibar leopard

## References

[edit]

1. ^ a b c d Kittle, A.M. & Watson, A. (2020). "Panthera pardus ssp.kotiya". IUCN Red List of Threatened Species. 2020: e.T15959A50660847.
2. ^ Deraniyagala, P. E. P. (1956). "The Ceylon leopard, a distinct subspecies". Spolia Zeylanica. 28: 115–116.
3. ^ Pocock, R.I. (1939). "Panthera pardus (Linnaeus). The Leopard or Panther". The Fauna of British India, including Ceylon and Burma. Vol. Mammalia. London: Taylor and Francis. pp. 222–231.
4. ^ "Yala's giant leopards". BBC Earth. 2014. Retrieved 2018-01-16.
5. ^ "Black Panther rediscovered in SLA". Daily News. Retrieved 2020-05-31.
6. ^ "Rare Black Panther Rediscovered In Sri Lanka". iflscience. 28 January 2020. Retrieved 2020-05-31.
7. ^ Kittle, A.M.; Watson, A.; Cushman, S.A.; Macdonald, D.W. (2018). "Forest cover and level of protection influence the island-wide distribution of an apex carnivore and umbrella species, the Sri Lankan leopard (Panthera pardus kotiya)". Biodiversity & Conservation. 27 (1): 235–263\. Bibcode:2018BiCon..27..235K. doi:10.1007/s10531-017-1431-8. S2CID 22699049.
8. ^ Phillips, W. W. A. (1984). Manual of the mammals of Sri Lanka. Vol. Part III (Second revised ed.). Colombo: Wildlife and Nature Protection Society of Sri Lanka.
9. ^ Kittle, A. M.; Watson, A. C.; Chanaka Kumara, P. H.; Nimalka Sanjeewani, H. K. (2014). "Status and distribution of the leopard in the central hills of Sri Lanka". Cat News. 56: 28−31.
10. ^ a b c Kittle, A.M.; Watson, A.C.; Fernando, T.S.P. (2017). "The ecology and behaviour of a protected area Sri Lankan leopard (Panthera pardus kotiya) population". Tropical Ecology. 58: 71–86.
11. ^ Kittle, A. M.; Watson, A. C.; Kumara, P. H. C.; Sandanayake, S. D. K.; Sanjeewani, H. K. N.; Fernando, S. (2014). "Notes on the diet, prey and habitat selection of the Sri Lankan leopard in the central highlands of Sri Lanka". Journal of Threatened Taxa. 6 (9): 6214–6221\. doi:10.11609/jott.o3731.6214-21.
12. ^ "Sri Lankan Leopard | Mammals in Sri Lanka | Dilmah Conservation". www.dilmahconservation.org. Retrieved 2020-12-25.
13. ^ Hadley, B. (2008). The Sloth Bear (PDF). IUCN/SSC Bear Specialist Group. Archived from the original (PDF) on 2008-12-21.
14. ^ Brown, G. (1993). The Great Bear Almanac. Lyons & Burford. ISBN 1558212108.
15. ^ "SL Black Leopard not extinct; spotted again". Daily Mirror. Retrieved 2020-05-31.
16. ^ "Rescued black leopard dies; wildlife officials yet to determine cause of death". Economy Next. 29 May 2020. Retrieved 2020-05-31.
17. ^ "Rare Black Leopard recently rescued from a snare dies". News First. 29 May 2020. Retrieved 2020-05-31.
18. ^ Kittle, A.M.; Watson, A.C.; Samaranayake, P.K.L. (2021). "Edge effects and distribution of prey forage resources influence how an apex predator utilizes Sri Lanka's largest protected area". Journal of Zoology. 314: 31–42\. doi:10.1111/jzo.12870. S2CID 234011936.
19. ^ Sarah Lazarus and Jon Jensen (15 January 2020). "Sri Lanka's leopards are under threat, but this woman is determined to save them". CNN. Retrieved 2021-01-17.
20. ^ Kittle, A. M.; Watson, A. C.; Fernando, S. (2012). "Notes on the status, distribution and abundance of the Sri Lankan leopard in the central hills of Sri Lanka". Cat News (56).
21. ^ International Species Information System (2011). "ISIS Species Holdings: Panthera pardus kotiya, December 2011".

## External links

[edit]

Wikimedia Commons has media related to Panthera pardus kotiya.

Wikispecies has information related to Panthera pardus kotiya.

* "Panthera pardus kotiya". IUCN/SSC Cat Specialist Group.
* "The Leopard Project". The Wilderness and Wildlife Conservation Trust, Sri Lanka.
* "Images and movies of the Sri Lankan leopard". ARKive. Archived from the original on 2008-07-25.
* Anthonis, L. (1997). "Lanka gets her own endemic leopard". Sunday Times.

* v
* t
* e

Extant Carnivora species

* Kingdom: Animalia
* Phylum: Chordata
* Class: Mammalia
* Infraclass: Eutheria
* Superorder: Laurasiatheria

Feliformia ("cat-like" carnivorans)

Feliformia

* African palm civet (Nandinia binotata)

Feloidea

| Prionodon (Asiatic linsangs)|

* Banded linsang (P. linsang)
* Spotted linsang (P. pardicolor)

|

Felidae (cats)

| Pantherinae|

| Neofelis|

* Sunda clouded leopard (N. diardi)
* Clouded leopard (N. nebulosa)

|

Panthera|

* Snow leopard (P. uncia)
* Tiger (P. tigris)
* Jaguar (P. onca)
* Lion (P. leo)
* Leopard (P. pardus)

Felinae sensu stricto

| Bay cat
lineage|

| Pardofelis|

* Marbled cat (P. marmorata)

|

Catopuma|

* Bay cat (C. badia)
* Asian golden cat (C. temminckii)

Caracal
lineage|

* Serval (Leptailurus serval)

Caracal|

* African golden cat (C. aurata)
* Caracal (C. caracal)

Leopardus|

* Pampas cat (L. colocola)
* Geoffroy's cat (L. geoffroyi)
* Kodkod (L. guigna)
* Southern tiger cat (L. guttulus)
* Andean mountain cat (L. jacobita)
* Ocelot (L. pardalis)
* Oncilla (L. tigrinus)
* Margay (L. wiedii)

Lynx|

* Canada lynx (L. canadensis)
* Eurasian lynx (L. lynx)
* Iberian lynx (L. pardinus)
* Bobcat (L. rufus)

Puma
lineage|

| Acinonyx|

* Cheetah (A. jubatus)

|

* Jaguarundi (Herpailurus yagouaroundi)

Puma|

* Cougar (P. concolor)

Leopard cat
lineage|

* Pallas's cat (Otocolobus manul)

Prionailurus|

* Leopard cat (P. bengalensis)
* Sunda leopard cat (P. javanensis)
* Flat-headed cat (P. planiceps)
* Rusty-spotted cat (P. rubiginosus)
* Fishing cat (P. viverrinus)

Felis|

* Chinese mountain cat (F. bieti)
* Domestic cat (F. catus)
* Jungle cat (F. chaus)
* African wildcat (F. lybica)
* Sand cat (F. margarita)
* Black-footed cat (F. nigripes)
* European wildcat (F. silvestris)

Viverroidea|

*     * see below↓

Viverroidea

Viverridae

Palm civets

| Hemigalinae|

* Owston's palm civet (Chrotogale owstoni)
* Otter civet (Cynogale bennettii)
* Hose's palm civet (Diplogale hosei)
* Banded palm civet (Hemigalus derbyanus)

|

Paradoxurinae|

* Binturong (Arctictis binturong)
* Small-toothed palm civet (Arctogalidia trivirgata)
* Sulawesi palm civet (Macrogalidia musschenbroekii)
* Masked palm civet (Paguma larvata)

Paradoxurus|

* Asian palm civet (P. hermaphroditus)
* Brown palm civet (P. jerdoni)
* Golden palm civet (P. zeylonensis)

Viverrinae sensu lato

| Viverrinae
sensu stricto|

* Small Indian civet (Viverricula indica)
* African civet (Civettictis civetta)

Viverra|

* Malabar large-spotted civet (V. civettina)
* Large-spotted civet (V. megaspila)
* Malayan civet (V. tangalunga)
* Large Indian civet (V. zibetha)

Genettinae

| Poiana
(African linsangs)|

* Central African oyan (P. richardsonii)
* West African oyan (P. leightoni)

|

Genetta (genets)

| subgenus Genetta
(paraphyletic)|

* Abyssinian genet (G. abyssinica)
* Common genet (G. genetta)
* South African small-spotted genet (G. felina)

|

subgenus Eugenetta
(paraphyletic)|

* Angolan genet (G. angolensis)
* Cape genet (G. tigrina)

subgenus Herpailuropoda
(paraphyletic)|

* Bourlon's genet (G. bourloni)
* Johnston's genet (G. johnstoni)
* Hausa genet (G. thierryi)
* Giant forest genet (G. victoriae)

subgenus Pardogale
(paraphyletic)|

* Pardine genet (G. pardina)
* King genet (G. poensis)

subgenus Prionailuropoda|

* Letaba genet (G. letabae)
* Rusty-spotted genet (G. maculata)
* Schouteden's genet (G. schoutedeni)

subgenus Leptailuropoda
(paraphyletic)|

* Crested servaline genet (G. cristata)
* Servaline genet (G. servalina)

subgenus Osbornictis|

* Aquatic genet (G. piscivora)

| Herpestoidea|

*     * see below↓

|

Herpestoidea

| Hyaenidae
(hyenas)|

| Proteles|

* Aardwolf (P. cristata)

|

Hyaeninae
(bone-crushing hyenas)|

* Striped hyena (Hyaena hyaena)
* Brown hyena (Parahyaena brunnea)

Crocuta|

* Spotted hyena (C. crocuta)

Herpestidae sensu lato

| Eupleridae
(Malagasy
carnivorans)|

| Euplerinae
(Malagasy civets)|

* Fossa (Cryptoprocta ferox)
* Malagasy civet (Fossa fossana)

Eupleres (falanoucs)|

* Eastern falanouc (E. goudotii)
* Western falanouc (E. major)

Galidiinae
(vontsira)|

* Ring-tailed vontsira (Galidia elegans)

Galidictis|

* Broad-striped Malagasy mongoose (G. fasciata)
* Grandidier's mongoose (G. grandidieri)

* Narrow-striped mongoose (Mungotictis decemlineata)

Salanoia|

* Brown-tailed mongoose (S. concolor)
* Durrell's vontsira (S. durrelli)

Herpestidae sensu stricto (mongooses)

Mungotinae

| Suricata|

* Meerkat (S. suricatta)

|

* Liberian mongoose (Liberiictus kuhni)

Mungos|

* Gambian mongoose (M. gambianus)
* Banded mongoose (M. mungo)

* Pousargues's mongoose (Dologale dybowskii)

Helogale|

* Ethiopian dwarf mongoose (H. hirtula)
* Common dwarf mongoose (H. parvula)

Crossarchus
(kusimanses)|

* Alexander's kusimanse (C. alexandri)
* Angolan kusimanse (C. ansorgei)
* Common kusimanse (C. obscurus)
* Flat-headed kusimanse (C. platycephalus)

Herpestinae

* Marsh mongoose (Atilax paludinosus)
* Long-nosed mongoose (Xenogale naso)

Urva
(Asian mongooses)|

* Small Indian mongoose (U. auropunctata)
* Short-tailed mongoose (U. brachyura)
* Indian grey mongoose (U. edwardsii)
* Indian brown mongoose (U. fusca)
* Javan mongoose (U. javanica)
* Collared mongoose (U. semitorquata)
* Ruddy mongoose (U. smithii)
* Crab-eating mongoose (U. urva)
* Stripe-necked mongoose (U. vitticolla)

* White-tailed mongoose (Ichneumia albicauda)
* Yellow mongoose (Cynictis penicillata)
* Selous's mongoose (Paracynictis selousi)
* Meller's mongoose (Rhynchogale melleri)

Bdeogale|

* Bushy-tailed mongoose (B. crassicauda)
* Jackson's mongoose (B. jacksoni)
* Black-footed mongoose (B. nigripes)

Herpestes
(slender mongooses)|

* Angolan slender mongoose (H. flavescens)
* Egyptian mongoose (H. ichneumon)
* Somalian slender mongoose (H. ochracea)
* Cape gray mongoose (H. pulverulenta)
* Common slender mongoose (H. sanguinea)

Caniformia ("dog-like" carnivorans)

Canidae (dogs)

| Urocyon|

* Gray fox (U. cinereoargenteus)
* Island fox (U. littoralis)

|

Vulpini

* Bat-eared fox (Otocyon megalotis)

Nyctereutes
(raccoon dogs)|

* Common raccoon dog (N. procyonoides)
* Japanese raccoon dog (N. viverrinus)

Vulpes
(true foxes)|

* Bengal fox (V. bengalensis)
* Blanford's fox (V. cana)
* Cape fox (V. chama)
* Corsac fox (V. corsac)
* Tibetan fox (V. ferrilata)
* Arctic fox (V. lagopus)
* Kit fox (V. macrotis)
* Pale fox (V. pallida)
* Rüppell's fox (V. rueppelli)
* Swift fox (V. velox)
* Red fox (V. vulpes)
* Fennec fox (V. zerda)

Canini (true dogs)

Cerdocyonina
(zorro)

* Maned wolf (Chrysocyon brachyurus)

Speothos|

* Bush dog (S. venaticus)

* Short-eared dog (Atelocynus microtis)
* Crab-eating fox (Cerdocyon thous)

Lycalopex
(South American foxes)|

* Culpeo (L. culpaeus)
* Darwin's fox (L. fulvipes)
* South American gray fox (L. griseus)
* Pampas fox (L. gymnocercus)
* Sechuran fox (L. sechurae)
* Hoary fox (L. vetulus)

Canina
(wolf-like canids)

| Lupulella|

* Side-striped jackal (L. adustus)
* Black-backed jackal (L. mesomelas)

|

Lycaon|

* African wild dog (L. pictus)

* Dhole (Cuon alpinus)

Canis|

* Golden jackal (C. aureus)
* Domestic dog (C. familiaris)
* Coyote (C. latrans)
* African wolf (C. lupaster)
* Wolf (C. lupus)
* Eastern wolf (C. lycaon)
* Red wolf (C. rufus)
* Ethiopian wolf (C. simensis)

Arctoidea

Ursidae (bears)

| Ailuropoda|

* Giant panda (A. melanoleuca)

|

Tremarctos|

* Spectacled bear (T. ornatus)

Ursinae|

* Sloth bear (Melursus ursinus)
* Sun bear (Helarctos malayanus)

Ursus|

* American black bear (U. americanus)
* Asian black bear (U. thibetanus)
* Brown bear (U. arctos)
* Polar bear (U. maritimus)

| Mustelida|

| Pinnipedia (seals)|

*     * see below↓

|

Musteloidea|

*     * see below↓

Pinnipedia (seals)

| Odobenidae|

* Walrus (Odobenus rosmarus)

|

Otariidae (eared seals)

| Callorhinus
(northern fur seals)|

* Northern fur seal (C. ursinus)

|

Otariinae
(sea lions)|

* Steller sea lion (Eumetopias jubatus)

Zalophus|

* California sea lion (Z. californianus)
* Galápagos sea lion (Z. wollebaeki)

* South American sea lion (Otaria flavescens)

Neophoca|

* Australian sea lion (N. cinerea)

* New Zealand sea lion (Phocarctos hookeri)

Arctocephalus
(southern fur seals)|

* South American fur seal (A. australis)
* Australasian fur seal (A. forsteri)
* Galápagos fur seal (A. galapagoensis)
* Antarctic fur seal (A. gazella)
* Juan Fernández fur seal (A. philippii)
* Brown fur seal (A. pusillus)
* Guadalupe fur seal (A. townsendi)
* Subantarctic fur seal (A. tropicalis)

Phocidae (earless seals or true seals)

Phocinae
("northern seals")

* Bearded seal (Erignathus barbatus)
* Hooded seal (Cystophora cristata)

Phocini

* Harp seal (Pagophilus groenlandicus)
* Ribbon seal (Histriophoca fasciata)
* Grey seal (Halichoerus grypus)

Phoca|

* Spotted seal (P. largha)
* Harbor seal (P. vitulina)

Pusa|

* Caspian seal (P. caspica)
* Ringed seal (P. hispida)
* Baikal seal (P. sibirica)

Monachinae
("southern seals")

| Monachini
(monk seals)|

* Mediterranean monk seal (Monachus monachus)

Neomonachus|

* Hawaiian monk seal (N. schauinslandi)

Mirounga
(elephant seals)|

* Northern elephant seal (M. angustirostris)
* Southern elephant seal (M. leonina)

Lobodontini
(Antarctic seals)|

* Leopard seal (Hydrurga leptonyx)
* Weddell seal (Leptonychotes weddellii)
* Crabeater seal (Lobodon carcinophagus)
* Ross seal (Ommatophoca rossi)

Musteloidea

| Ailuridae|

* Red panda (Ailurus fulgens)

|

Mephitidae (skunks)

| Conepatus
(hog-nosed skunks)|

* Molina's hog-nosed skunk (C. chinga)
* Humboldt's hog-nosed skunk (C. humboldtii)
* American hog-nosed skunk (C. leuconotus)
* Striped hog-nosed skunk (C. semistriatus)

|

Mephitis|

* Hooded skunk (M. macroura)
* Striped skunk (M. mephitis)

Mydaus
(stink badgers)|

* Sunda stink badger (M. javanensis)
* Palawan stink badger (M. marchei)

Spilogale
(spotted skunks)|

* Southern spotted skunk (S. angustifrons)
* Western spotted skunk (S. gracilis)
* Eastern spotted skunk (S. putorius)
* Pygmy spotted skunk (S. pygmaea)

Procyonidae

* Kinkajou (Potos flavus)

Bassariscus|

* Ringtail (B. astutus)
* Cacomistle (B. sumichrasti)

Procyon
(raccoons)|

* Crab-eating raccoon (P. cancrivorus)
* Raccoon (P. lotor)
* Cozumel raccoon (P. pygmaeus)

Bassaricyon
(olingos)|

* Eastern lowland olingo (B. alleni)
* Northern olingo (B. gabbii)
* Western lowland olingo (B. medius)
* Olinguito (B. neblina)

Nasuina
(coatis)|

| Nasua|

* White-nosed coati (N. narica)
* South American coati (N. nasua)

|

Nasuella
(mountain coatis)|

* Eastern mountain coati (N. meridensis)
* Western mountain coati (N. olivacea)

| Mustelidae|

*     * see below↓

|

Mustelidae

Mustelidae

* American badger (Taxidea taxus)

Mellivora|

* Honey badger (M. capensis)

Melinae

| Arctonyx
(hog badgers)|

* Northern hog badger (A. albogularis)
* Greater hog badger (A. collaris)
* Sumatran hog badger (A. hoevenii)

|

Meles
(Eurasian badgers)|

* Japanese badger (M. anakuma)
* Caucasian badger (M. canescens)
* Asian badger (M. leucurus)
* European badger (M. meles)

Melogale
(ferret-badgers)|

* Vietnam ferret-badger (M. cucphuongensis)
* Bornean ferret badger (M. everetti)
* Chinese ferret-badger (M. moschata)
* Javan ferret-badger (M. orientalis)
* Burmese ferret-badger (M. personata)
* Formosan ferret-badger (M. subaurantiaca)

Guloninae

* Tayra (Eira barbara)

Pekania|

* Fisher (P. pennanti)

Gulo|

* Wolverine (G. gulo)

Martes
(martens)|

* American marten (M. americana)
* Pacific marten (M. caurina)
* Yellow-throated marten (M. flavigula)
* Beech marten (M. foina)
* Nilgiri marten (M. gwatkinsii)
* European pine marten (M. martes)
* Japanese marten (M. melampus)
* Sable (M. zibellina)

Ictonychinae

| Lyncodontini|

* Patagonian weasel (Lyncodon patagonicus)

Galictis
(grisons)|

* Lesser grison (G. cuja)
* Greater grison (G. vittata)

Ictonychini
(African polecats)|

| Vormela|

* Marbled polecat (V. peregusna)

|

* African striped weasel (Poecilogale albinucha)

Ictonyx|

* Saharan striped polecat (I. libycus)
* Striped polecat (I. striatus)

Lutrinae (otters)

* Giant otter (Pteronura brasiliensis)

Lontra|

* North American river otter (L. canadensis)
* Marine otter (L. felina)
* Neotropical otter (L. longicaudis)
* Southern river otter (L. provocax)

Enhydra|

* Sea otter (E. lutris)

* Spotted-necked otter (Hydrictis maculicollis)

Lutra|

* Eurasian otter (L. lutra)
* Hairy-nosed otter (L. sumatrana)

Lutrogale|

* Smooth-coated otter (L. perspicillata)

Aonyx|

* African clawless otter (A. capensis)
* Asian small-clawed otter (A. cinereus)
* Congo clawless otter (A. congicus)

Mustelinae

| Neogale
(New World weasels)|

* Amazon weasel (N. africana)
* Colombian weasel (N. felipei)
* Long-tailed weasel (N. frenata)
* American mink (N. vison)

|

Mustela
(weasels)

| subgenus Mustela
(paraphyletic)|

* Sichuan weasel (M. aistoodonnivalis)
* Mountain weasel (M. altaica)
* Stoat/Beringian ermine (M. erminea)
* Haida ermine (M. haidarum)
* Yellow-bellied weasel (M. kathiah)
* Least weasel (M. nivalis)
* American ermine (M. richardsonii)

|

subgenus Lutreola
(paraphyletic)|

* Japanese weasel (M. itatsi)
* European mink (M. lutreola)
* Indonesian mountain weasel (M. lutreolina)
* Malayan weasel (M. nudipes)
* Siberian weasel (M. sibirica)
* Back-striped weasel (M. strigidorsa)

subgenus Putorius|

* Steppe polecat (M. eversmannii)
* Ferret (M. furo)
* Black-footed ferret (M. nigripes)
* European polecat (M. putorius)

* See also: Mink
* Polecat

Taxon identifiers

Panthera pardus kotiya|

* Wikidata: Q475046
* Wikispecies: Panthera pardus kotiya
* ARKive: panthera-pardus-kotiya
* CoL: 7KH89
* EoL: 1253181
* GBIF: 5219437
* iNaturalist: 147732
* ITIS: 726468
* IUCN: 15959
* MSW: 14000255
* NCBI: 2816903
* Observation.org: 186420

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