Taxonomy of the Dermaptera

Biological ranks of earwigs

This taxonomy of the Dermaptera follows Engel & Haas (2007) to the rank of genus.[1]

## Taxonomy[edit]

* Order Dermaptera de Geer, 1773

* Suborder †Archidermaptera Bey-Bienko, 1936

* Superfamily †Protodiplatyoidea Martynov, 1925a

* Family †Protodiplatyidae Martynov, 1925a

* Genus †Abrderma Xing, Shih, & Ren, 2016
* Genus †Aneuroderma Xiong, Engel, & Ren, 2021
* Genus †Anisodiplatys Vishniakova, 1980
* Genus †Applanatiforceps Yin, Shih, Engel, & Ren, 2021
* Genus †Archidermapteron Vishniakova, 1980
* Genus †Asiodiplatys Vishniakova, 1980
* Genus †Babderma Xing, Shih, & Ren, 2016
* Genus †Brevicula Whalley, 1985
* Genus †Longicerciata Zhang, 1994
* Genus †Microdiplatys Vishniakova, 1980
* Genus †Perissoderma Xing, Shih, & Ren, 2016
* Genus †Protodiplatys Martynov, 1925
* Genus †Sinoprotodiplatys Nel, Aria, Garrouste, & Waller, 2012

* Family †Dermapteridae Vishniakova, 1980

* Genus †Dacryoderma Engel, 2021
* Genus †Dimapteron Kelly, Ross, & Jarzembowski, 2016
* Genus †Dermapteron Martynov, 1925
* Genus †Jurassimedeola Zhang, 2002
* Genus †Palaeodermapteron Zhao, Shih, & Ren, 2011
* Genus †Phanerogramma Cockerell, 1915
* Genus †Sinoplaeodermata Zhang, 2002
* Genus †Valdopteron Kelly, Ross, & Jarzembowski, 2016

* Family †Turanoviidae Engel, 2003

* Genus †Turanovia Vishniakova, 1980

* Family †Incertae sedis in †Archidermaptera Bey-Bienko, 1936

* Genus †Trivenapteron Kelly, Ross, & Jarzembowski, 2016

* Suborder †Eodermaptera Engel, 2003

* Superfamily †Bellodermatoidea Zhao, Shih, & Ren, 2010

* Family †Bellodermatidae Zhao, Shih, & Ren, 2010

* Genus †Belloderma Zhao, Shih, & Ren, 2010

* Superfamily †Semenovioloidea Vishniakova, 1980

* Family †Semenoviolidae Vishniakova, 1980

* Subfamily †Semenoviolinae Vishniakova, 1980

* Genus †Semenoviola Vishniakova, 1980
* Genus †Semenovioloides Vishniakova, 1980

* Subfamily †Aglyptodermatinae Xiong, Engel, & Ren, 2021

* Genus †Aglyptoderma Xiong, Engel, & Ren, 2021
* Genus †Ekpagloderma Yin, Shih, Engel, & Ren, 2023

* Infraorder †Turanodermaptera Engel, 2019

* Superfamily †Turanodermatoidea Engel, 2003

* Family †Turanodermatidae Engel, 2003

* Genus †Turanoderma Vishniakova, 1980

* Suborder Neodermaptera Engel, 2003

* Infraorder Protodermaptera Zacher, 1910

* Superfamily Diplatyoidea Verhoeff, 1902a

* Family Diplatyidae Verhoeff, 1902a

* Genus †Acanthodiplatys Ren, Zhang, Shih, & Ren, 2018
* Genus Circodiplatys Steinmann, 1986
* Genus Diplatys Audinet-Serville, 1831
* Genus †Hirtidiplatys Ren, Zhang, Shih, & Ren, 2018
* Genus Lobodiplatys Steinmann, 1974
* Genus Mesodiplatys Steinmann, 1986
* Genus Schizodiplatys Steinmann, 1974
* Genus †Tytthodiplatys Engel, 2011

* Family Haplodiplatyidae Engel, 2017

* Genus Haplodiplatys Hincks, 1955

* Superfamily Pygidicranoidea Verhoeff, 1902a

* Family Pygidicranidae Verhoeff, 1902a

* Subfamily †Astreptolabidinae Engel, 2011

* Genus †Astreptolabis Engel, 2011

* Subfamily †Burmapygiinae Engel & Grimaldi, 2004

* Genus †Burmapygia Engel & Grimaldi, 2004

* Subfamily Anataeliinae Burr, 1909b

* Genus Anataelia Bolivar, 1899

* Subfamily Blandicinae Burr, 1915

* Genus Alloblandex Hincks, 1957
* Genus Austroblandex Brindle, 1986
* Genus Blandex Burr, 1912
* Genus Parablandex Brindle, 1966

* Subfamily Brindlesiinae Srivastava, 1985b

* Genus Brindlesia Srivastava, 1985b

* Subfamily Challiinae Steinmann, 1973

* Genus Challia Burr, 1904

* Subfamily Cylindrogastrinae Maccagno, 1929

* Genus Cylindrogaster Stål, 1855

* Subfamily Diplatymorphinae Boeseman, 1954

* Genus Diplatymorpha Boeseman, 1954

* Subfamily Echinosomatinae Burr, 1910a

* Genus Parapsalis Borelli, 1921
* Genus Echinosoma Audinet-Serville, 1831

* Subfamily Esphalmeninae Burr, 1909a

* Genus Esphalmenus Burr, 1909

* Subfamily Karschiellinae Verhoeff, 1902a

* Genus Bormansia Verhoeff, 1902
* Genus Karschiella Verhoeff, 1902

* Subfamily †Prodacnodinae Peng, Engel, Liu, & Nel, 2024

* Genus †Prodacnodes Peng, Engel, Liu, & Nel, 2024

* Subfamily Pygidicraninae Verhoeff, 1902a

* Genus Acrania Burr, 1915
* Genus Cranopygia Burr, 1908
* Genus Dacnodes Burr, 1907
* Genus Paracranopygia Steinmann, 1986
* Genus Pygidicrana Audinet-Serville, 1831
* Genus Tagalina Dohrn, 1863

* Subfamily Pyragrinae Verhoeff, 1902a

* Genus Echinopsalis Bormans, 1893
* Genus †Gracilipygia Ren, Zhang, Shih, & Ren, 2017
* Genus Pyragra Audinet-Serville, 1831
* Genus Pyragropsis Borelli, 1908

* Subfamily †Stonychopygiinae Engel & Huang, 2017

* Genus †Stonychopygia Engel & Huang, 2017

* Subfamily Incertae sedis in Pygidicranidae Verhoeff, 1902a

* Genus †Archaeosoma Zhang, 1994
* Genus †Eminepygia Chen & Zhang, 2021
* Genus †Gallinympha Perrichot & Engel, 2011
* Genus †Geosoma Zhang, 1997
* Genus †Robustipygia Ren, Zhang, Shih, & Ren, 2018

* Infraorder Epidermaptera Engel, 2003

* Parvorder Paradermaptera Verhoeff, 1902a

* Superfamily Hemimeroidea Sharp, 1895

* Family Hemimeridae Sharp, 1895

* Genus Araeomerus Maa, 1974
* Genus Hemimerus Walker, 1871

* Superfamily Apachyoidea Verhoeff, 1902a

* Family Apachyidae Verhoeff, 1902a

* Genus Apachys Audinet-Serville, 1831
* Genus Dendroiketes Burr, 1909

* Parvorder Metadermaptera Engel, 2003

* Superfamily Anisolabidoidea Verhoeff, 1902a

* Family Anisolabididae Verhoeff, 1902a

* Genus Aborolabis Srivastava, 1969
* Genus Anisolabis Fieber, 1853
* Genus Canarilabis Steinmann, 1985
* Genus Carcinophora Scudder, 1876
* Genus Epilandex Hebard, 1927
* Genus Euborellia Burr, 1909
* Genus Flexiolabis Steinmann, 1989
* Genus Gonolabis Burr, 1910
* Genus Indolabis Steinmann, 1989
* Genus Mongolabis Zacher, 1911
* Genus Neolabis Brindle, 1981
* Genus Ornatolabis Steinmann, 1989
* Genus Paraflexiolabis Steinmann, 1989
* Genus Placolabis Bey-Bienko, 1959
* Genus Socotralabis Kočárek, 2014
* Genus Thekalabis Kapoor, 1967
* Genus Zacheria Steinmann, 1975

* Subfamily Anophthalmolabidinae Steinmann, 1975

* Genus Anophthalmolabis Brindle, 1968

* Subfamily Antisolabidinae Brindle, 1978a

* Genus Antisolabis Burr, 1911

* Subfamily Brachylabidinae Burr, 1908a

* Genus Brachylabis Dohrn, 1864
* Genus Ctenisolabis Verhoeff, 1902
* Genus Metisolabis Burr, 1910

* Subfamily †Cretolabiinae Engel & Haas, 2007

* Genus †Cretolabia Popham, 1990

* Subfamily Gonolabininae Popham and Brindle, 1966c

* Genus Gonolabina Verhoeff, 1902

* Subfamily Idolopsalidinae Steinmann, 1975

* Genus Idolopsalis Borelli, 1910

* Subfamily Isolabidinae Verhoeff, 1902b

* Genus Africolabis Brindle, 1978
* Genus Geracodes Hebard, 1917
* Genus Isolabis Verhoeff, 1902
* Genus Pterolabis Steinmann, 1989

* Subfamily Parisolabidinae Verhoeff, 1904

* Genus Parisolabis Verhoeff, 1904
* Genus Parisopsalis Burr, 1914

* Subfamily Platylabiinae Burr, 1911

* Genus Platylabia Dohrn, 1867

* Subfamily Titanolabidinae Srivastava, 1982

* Genus Titanolabis Burr, 1910

* Subfamily Incertae sedis in Anisolabididae Verhoeff, 1902a

* Genus †Geroncolabis Engel & Chatzimanolis, 2010
* Genus †Kotejalabis Engel & Chatzimanolis, 2005
* Genus †Palaeocarcinophora Engel & Chatzimanolis, 2010
* Genus †Spiladopygia Engel & Chatzimanolis, 2010
* Genus †Toxolabis Engel & Grimaldi, 2014

* Parvorder Eteodermaptera Engel, 2003

* Nanorder Plesiodermaptera Engel, 2003

* Superfamily Labiduroidea Verhoeff, 1902a

* Family Labiduridae Verhoeff, 1902a

* Subfamily Allostethinae Verhoeff, 1904

* Genus Allostethella Zacher, 1910
* Genus Allostethus Verhoeff, 1904
* Genus Gonolabidura Zacher, 1910
* Genus Protolabidura Steinmann, 1985

* Subfamily Labidurinae Verhoeff, 1902a

* Genus Forcipula Bolivar, 1897
* Genus Labidura Leach, 1815
* Genus Tomopygia Burr, 1904

* Subfamily Nalinae Steinmann, 1975

* Genus Nala Zacher, 1910

* Subfamily †Zigrasolabidinae Engel, 2022

* Genus †Zigrasolabis Engel & Grimaldi, 2014
* Genus †Tricholabidura Peng & Engel, 2022

* Subfamily Incertae sedis in Labiduridae Verhoeff, 1902a

* Genus †Acantholabis Mao, Engel, & Ren, 2020
* Genus †Caririlabia Martins-Neto, 1990
* Genus †Myrrholabia Engel & Grimaldi, 2004

* Nanorder Eudermaptera Verhoeff, 1902a

* Superfamily Forficuloidea Latreille, 1810

* Family Arixeniidae Jordan, 1909

* Genus Arixenia Jordan, 1909
* Genus Xeniaria Maa, 1974

* Family Spongiphoridae Verhoeff, 1902a

* Subfamily Caecolabiinae Steinmann, 1990

* Genus Caecolabia Brindle, 1975

* Subfamily Cosmogeracinae Brindle, 1982

* Genus Cosmogerax Hebard, 1933

* Subfamily Geracinae Brindle, 1971

* Genus Apterogerax Engel, 2017
* Genus Barygerax Hebard, 1917
* Genus Eugerax Hebard, 1917
* Genus Gerax Hebard, 1917
* Genus †Nesogerax Engel, 2017
* Genus Pseudovostox Borelli, 1926
* Genus Yepezia Brindle, 1982

* Subfamily Isolaboidinae Brindle, 1978b

* Genus Isolaboides Hincks, 1958
* Genus Paralaboides Steinmann, 1989

* Subfamily Isopyginae Hincks, 1951

* Genus Isopyge Borelli, 1931

* Subfamily Labiinae Burr, 1909b

* Genus Chaetolabia Brindle, 1972
* Genus Circolabia Steinmann, 1987
* Genus Isolabella Verhoeff, 1902
* Genus Labia Leach, 1815
* Genus Paralabella Steinmann, 1990
* Genus Paraspania Steinmann, 1985
* Genus Sphingolabis Bormans, 1883
* Genus Spirolabia Steinmann, 1987

* Subfamily Nesogastrinae Verhoeff, 1902a

* Genus Nesogaster Verhoeff, 1902

* Subfamily Nesolabiinae Engel, 2017

* Genus Nesolabia Hincks, 1957

* Subfamily Pericominae Burr, 1911a

* Genus Parapericomus Ramamurthi, 1967
* Genus Pericomus Burr, 1911

* Subfamily Ramamurthiinae Steinmann, 1975

* Genus Ramamurthia Steinmann, 1975

* Subfamily Rudracinae Srivastava, 1995

* Genus Rudrax Srivastava, 1995

* Subfamily Sparattinae Verhoeff, 1902a

* Tribe Auchenomini Burr, 1909b

* Genus Auchenomus Karsch, 1886

* Tribe Chaetospaniini Steinmann, 1990

* Genus Chaetospania Karsch, 1886

* Tribe Sparattini Verhoeff, 1902a

* Genus Mecomera Audinet-Serville, 1838
* Genus Sparatta Audinet-Serville, 1838

* Subfamily Spongiphorinae Verhoeff, 1902a

* Genus Filolabia Steinmann, 1989
* Genus Formicilabia Rehn & Hebard, 1917
* Genus Homotages Burr, 1909
* Genus †Ikelus Estrada-Álvarez & Núñez-Bázan, 2023
* Genus Irdex Burr, 1911
* Genus Marava Burr, 1911
* Genus Pseudomarava Steinmann, 1989
* Genus Purex Burr, 1911
* Genus Spongiphora Audinet-Serville, 1831
* Genus Spongovostox Burr, 1911
* Genus Vostox Burr, 1911

* Subfamily Strongylopsalinae Burr, 1911a

* Genus Strongylolabis Steinmann, 1986
* Genus Strongylopsalis Burr, 1900

* Subfamily Vandicinae Burr, 1911a

* Genus Vandex Burr, 1911

* Subfamily Incertae sedis in Spongiphoridae Verhoeff, 1902

* Genus Paratages Srivastava, 1987

* Family Chelisochidae Verhoeff, 1902a

* Subfamily Chelisochinae Verhoeff, 1902a

* Genus Adiathella Brindle, 1970
* Genus Adiathetus Burr, 1907
* Genus Chelisochella Verhoeff, 1902
* Genus Chelisoches Scudder, 1876
* Genus †Chelisoficula Nel, Waller, Albouy, Menier, & de Ploëg, 2003
* Genus Euenkrates Rehn, 1927
* Genus Exypnus Burr, 1907
* Genus Gressitolabis Brindle, 1970
* Genus Hamaxas Burr, 1907
* Genus Lamprophorella Mjöberg, 1924
* Genus Proreus Burr, 1907
* Genus Schizochelisoches Steinmann, 1987
* Genus Schizoproreus Steinmann, 1987
* Genus Solenosoma Burr, 1907

* Subfamily Genitalatinae Steinmann, 1987

* Genus Genitalata Kapoor, 1974

* Subfamily Kinesinae Srivastava, 2003

* Genus Kinesis Burr, 1907

* Subfamily Incertae sedis in Chelisochidae Verhoeff, 1902a

* Genus †Amblypygia Engel, 2016
* Genus †Sthenotropis Engel, 2016

* Family Forficulidae Latreille, 1810

* Subfamily Allodahliinae Verhoeff, 1902a

* Genus Allodahlia Verhoeff, 1902
* Genus Brindleiana Steinmann, 1975
* Genus Eulithinus Hincks, 1935

* Subfamily Ancistrogastrinae Verhoeff, 1902a

* Genus Ancistrogaster Stål, 1855
* Genus Litocosmia Hebard, 1917
* Genus Osteulcus Burr, 1907
* Genus Paracosmia Borelli, 1909
* Genus Praos Burr, 1907
* Genus Sarcinatrix Rehn, 1903

* Subfamily Anechurinae Verhoeff, 1902a

* Genus Anechura Scudder, 1876
* Genus Chelidura Latreille, 1825
* Genus Chelidurella Verhoeff, 1902
* Genus Eumegalura Bey-Bienko, 1934
* Genus Mesasiobia Semenov, 1908
* Genus Mesochelidura Verhoeff, 1902
* Genus Neopterygidea Srivastava, 1984
* Genus Oreasiobia Semenov, 1936
* Genus Perirrhytus Burr, 1911
* Genus Pseudochelidura Verhoeff, 1902
* Genus Pterygidea Verhoeff, 1902

* Subfamily Diaperasticinae Burr, 1907

* Genus Diaperasticus Burr, 1907

* Subfamily Forficulinae Latreille, 1810

* Genus Afroforficula Steinmann, 1990
* Genus Apterygidea Westwood, 1840
* Genus Champaipites Burr, 1907
* Genus Doru Burr, 1907
* Genus Elaunon Burr, 1907
* Genus Forficula Linnaeus, 1758
* Genus Guanchia Burr, 1911
* Genus Mesolabia Shiraki, 1905
* Genus Parlax Burr, 1911
* Genus Proforficula Steinmann, 1990
* Genus Skalistes Burr, 1907
* Genus †Succiniforcipula Engel, 2016
* Genus Tauropygia Brindle, 1970

* Subfamily Neolobophorinae Burr, 1907

* Genus Dubianus Estrada-Álvarez & Núñez-Bázan, 2022
* Genus Eudohrnia Burr, 1907
* Genus Metresura Rehn, 1922
* Genus Neolobophora Scudder, 1875
* Genus Setocordax Brindle, 1970
* Genus Tristanella Borelli, 1909

* Subfamily Opisthocosmiinae Verhoeff, 1902a

* Genus Acanthocordax Günther, 1929
* Genus Chaetocosmia Nishikawa, 1973
* Genus Cipex Burr, 1910
* Genus Cordax Burr, 1910
* Genus Eparchus Burr, 1907
* Genus Eutimomena Bey-Bienko, 1970
* Genus Hypurgus Burr, 1907
* Genus Neoopisthocosmia Steinmann, 1990
* Genus Opisthocosmia Dohrn, 1865
* Genus Parasondax Srivastava, 1978
* Genus Parasyntonus Steinmann, 1990
* Genus Paratimomenus Steinmann, 1974
* Genus Pareparchus Burr, 1911
* Genus Prosadiya Hebard, 1923
* Genus Sondax Burr, 1910
* Genus Spinosocordax Steinmann, 1988
* Genus Syntonus Burr, 1910
* Genus Timomenus Burr, 1907

* Subfamily Skendylinae Burr, 1907

* Genus Afrocosmia Hincks, 1960
* Genus Brachycosmiella Steinmann, 1990
* Genus Cosmiella Verhoeff, 1902
* Genus Cosmiola Bey-Bienko, 1959
* Genus Forcepsia Moreira, 1930
* Genus Kleter Burr, 1907
* Genus Liparura Burr, 1907
* Genus Lipodes Burr, 1907
* Genus Mixocosmia Borelli, 1909
* Genus Neocosmiella Hebard, 1919
* Genus Obelura Burr, 1907
* Genus Paracosmiella Steinmann, 1990

* Subfamily Incertae sedis in Forficulidae Latreille, 1810

* Genus †Rupiforficula Engel & Chatzimanolis, 2010

* Family Incertae sedis in Forficuloidea Latreille, 1810

* Genus †Litholabis Engel & Chatzimanolis, 2010
* Genus †Petrolabis Engel & Chatzimanolis, 2010

* Superfamily Incertae sedis in Epidermaptera Engel, 2003

* Genus †Autrigonoforceps Engel & Peris, 2015
* Genus †Laasbium Scudder, 1900
* Genus †Labiduromma Scudder, 1890
* Genus †Rhadinolabis Engel, Ortega-Blanco, & Azar, 2011

## Nomina dubia[edit]

Family †Ocelliidae Ewing, 1942 is considered a nomen dubium.

## References[edit]

1. ^ Engel, M. S.; Haas, F. (2007). "Family-Group Names for Earwigs (Dermaptera)". American Museum Novitates (3567): 1–20. doi:10.1206/0003-0082(2007)539[1:FNFED]2.0.CO;2. hdl:2246/5858.

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Extant Dermaptera families

* Kingdom: Animalia
* Phylum: Arthropoda
* Class: Insecta
* Subclass: Pterygota
* Infraclass: Neoptera

* Anisolabididae
* Apachyidae
* Arixeniidae
* Chelisochidae
* Diplatyidae
* Forficulidae
* Hemimeridae
* Karschiellidae
* Labiduridae
* Pygidicranidae
* Spongiphoridae

See also: Taxonomy of the Dermaptera

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Taxonomy of ...

Life|

| Monophyletic|

* Bacteria

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* Plantae
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Animals|

* Animalia
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Polyphyletic|

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*[†]: Extinct
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