Trogiomorpha

Group of booklice

Trogiomorpha

Temporal range: Barremian–Recent

PreꞒ

Dorypteryx domestica
Scientific classification
Kingdom:  | Animalia
Phylum:  | Arthropoda
Class:  | Insecta
Order:  | Psocodea
Suborder:  | Trogiomorpha
Roesler, 1940
Infraorders[1]

See text

Trogiomorpha is one of the three major suborders of barklice, booklice, and parasitic lice in the order Psocodea (formerly Psocoptera),[2][3] alongside Troctomorpha and Psocomorpha. There are about 8 families and more than 430 described species in Trogiomorpha.[4][5][6] Trogiomorpha is widely agreed to be earliest diverging of the three suborders, and retains the most primitive characteristics.[7]

Trogium pulsatorium

## Contents

* 1 Internal phylogeny
* 2 Families
* 3 References
* 4 Further reading

## Internal phylogeny[edit]

Here is a cladogram showing the phylogeny of Psocodea, showing the position of Trogiomorpha:[1]

Psocodea |  |  |  | Troctomorpha |  |  |  |  |  |  |

Phthiraptera

|

Liposcelididae

|  |  |

Pachytroctidae

|

Sphaeropsocidae

Amphientometae

Psocomorpha

Trogiomorpha |  |  |  |  |

Atropetae

|

Psyllipsocetae

Prionoglaridetae (paraphyletic)

## Families[edit]

Psocomorpha contains 3 infraorders and 5 extant (living) families, as well as three identified extinct families:

* Atropetae
* †Archaeatropidae Baz & Ortuño, 2000
* †Empheriidae Baz & Ortuño, 2000
* Lepidopsocidae Enderlein, 1903 (scaly-winged barklice)
* Psoquillidae Lienhard & Smithers, 2002 (bird nest barklice)
* Trogiidae Roesler, 1944 (granary booklice)
* Psyllipsocetae
* Psyllipsocidae Lienhard & Smithers, 2002 (cave barklice)
* Prionoglaridetae (paraphyletic)
* Prionoglarididae Azar, Huang & Nel, 2017 (large-winged psocids)
* Unplaced:
*     * †Cormopsocidae Yoshizawa & Lienhard, 2020

## References[edit]

1. ^ a b de Moya, Robert S; Yoshizawa, Kazunori; Walden, Kimberly K O; Sweet, Andrew D; Dietrich, Christopher H; Kevin P, Johnson (2021-06-16). Buckley, Thomas (ed.). "Phylogenomics of Parasitic and Nonparasitic Lice (Insecta: Psocodea): Combining Sequence Data and Exploring Compositional Bias Solutions in Next Generation Data Sets". Systematic Biology. 70 (4): 719–738. doi:10.1093/sysbio/syaa075. ISSN 1063-5157.
2. ^ Johnson, Kevin P.; Smith, Vincent S. (2019). "Psocodea Species File Online". Retrieved 2019-06-14.
3. ^ Bess, Emilie; Smith, Vince; Lienhard, Charles; Johnson, Kevin P. (2006). "Psocodea". Tree of Life. Retrieved 2019-06-14.
4. ^ "Trogiomorpha Report". Integrated Taxonomic Information System. Retrieved 2019-05-04.
5. ^ "Trogiomorpha suborder Information". BugGuide.net. Retrieved 2019-05-04.
6. ^ Johnson, Kevin P.; Smith, Vincent S. (2019). "suborder Trogiomorpha". Psocodea species file online, Version 5.0. Retrieved 2019-05-04.
7. ^ Yoshizawa, Kazunori; Lienhard, Charles (June 2020). "†Cormopsocidae: A new family of the suborder Trogiomorpha (Insecta: Psocodea) from Burmese amber". Entomological Science. 23 (2): 208–215. doi:10.1111/ens.12414. ISSN 1343-8786.

## Further reading[edit]

* Lyal, Ch H. C. (1985). "Phylogeny and classification of the Psocodea, with particular reference to the lice (Psocodea: Phthiraptera)". Systematic Entomology. 10 (2): 145–165. doi:10.1111/j.1365-3113.1985.tb00525.x. ISSN 0307-6970.

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Extant Psocodea families

* Kingdom: Animalia
* Phylum: Arthropoda
* Class: Insecta
* Subclass: Pterygota
* Infraclass: Neoptera
* Superorder: Paraneoptera

Suborder Trogiomorpha

| Atropetae|

* Lepidopsocidae
* Psoquillidae
* Trogiidae

|

Prionoglaridetae|

* Prionoglarididae (paraphyletic)

Psyllipsocetae|

* Psyllipsocidae

Suborder Troctomorpha

| Amphientometae|

* Amphientomidae
* Compsocidae
* Manicapsocidae
* Musapsocidae
* Protroctopsocidae
* Troctopsocidae

|

Nanopsocetae|

* Liposcelididae
* Pachytroctidae
* Sphaeropsocidae

| Phthiraptera
(lice)|

| Amblycera|

* Ancistronidae
* Boopidae
* Colpocephalidae
* Gliricolidae
* Gyropidae
* Laemobothriidae
* Menoponidae (chicken body lice)
* Pseudomenoponidae
* Ricinidae
* Somaphantidae
* Trimenoponidae
* Trinotonidae

|

Anoplura
(sucking lice)|

* Echinophthiriidae (seal lice)
* Enderleinellidae
* Haematopinidae (ungulate lice)
* Hamophthiriidae
* Hoplopleuridae (armoured lice)
* Hybothiridae
* Linognathidae (pale lice)
* Microthoraciidae
* Neolinognathidae
* Pecaroecidae
* Pedicinidae
* Pediculidae (body lice, head lice)
* Polyplacidae (spiny rat lice)
* Pthiridae (crab lice or pubic lice)
* Ratemiidae

Ischnocera
(paraphyletic)|

* Bovicolidae
* Dasyonygidae
* Goniodidae
* Heptapsogasteridae
* Lipeuridae
* Philopteridae (paraphyletic)
* Trichodectidae
* Trichophilopteridae

Rhyncophthirina|

* Haematomyzidae

Ischnocera, Rhyncophthirina and Amblycera comprise paraphyletic suborder Mallophaga (chewing lice).

Suborder Psocomorpha

| Archipsocetae|

* Archipsocidae

|

Caeciliusetae|

* Amphipsocidae
* Asiopsocidae
* Caeciliusidae
* Dasydemellidae
* Paracaeciliidae
* Stenopsocidae

Epipsocetae|

* Cladiopsocidae
* Dolabellopsocidae
* Epipsocidae
* Ptiloneuridae
* Spurostigmatidae

Homilopsocidea|

* Ectopsocidae
* Elipsocidae
* Lachesillidae
* Lesneiidae
* Mesopsocidae
* Peripsocidae
* Sabulopsocidae

Philotarsetae|

* Philotarsidae
* Pseudocaeciliidae
* Trichopsocidae

Psocetae|

* Hemipsocidae
* Myopsocidae
* Psilopsocidae
* Psocidae

Order Psocodea is comprised from paraphyletic order Psocoptera (book lice or bark lice) and polyphyletic order Phthiraptera (lice).

Taxon identifiers|

* Wikidata: Q4051792
* Wikispecies: Trogiomorpha
* AFD: Trogiomorpha
* BugGuide: 536518
* CoL: 8MP9C
* Fauna Europaea: 12129
* Fauna Europaea (new): a08cf520-9a72-4e20-bd7b-cdc64327fea7
* Fossilworks: 216237
* iNaturalist: 212153
* ITIS: 103321
* NCBI: 159969
* NZOR: bd6063c7-8068-405d-81b9-fa5856263c5b
* Psocodea Species File: 1191651

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*[†]: Extinct
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