Urocoptidae

Family of gastropods

Urocoptidae

Apertural view of a shell of Holospira elizabethae
Scientific classification
Domain:  | Eukaryota
Kingdom:  | Animalia
Phylum:  | Mollusca
Class:  | Gastropoda
Subclass:  | Heterobranchia
Order:  | Stylommatophora
Superfamily:  | Urocoptoidea
Family:  | Urocoptidae
Pilsbry, 1898[1] (1868)

Urocoptidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Urocoptoidea.[2]

## Taxonomy[edit]

### 2005 taxonomy[edit]

The family Urocoptidae was classified in the superfamily Orthalicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[3]

The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

* Urocoptinae Pilsbry, 1898 (1868) - synonym: Cylindrellidae Tryon, 1868
* Apomatinae Paul, 1982
* Brachypodellinae H. B. Baker, 1956
* Eucalodiinae P. Fischer & Crosse, 1873
* Holospirinae Pilsbry, 1946
* Microceraminae Pilsbry, 1904 - synonyms: Johaniceraminae Jaume & de la Torre, 1972; Macroceraminae Jaume & de la Torre, 1972
* Tetrentodontinae Bartsch, 1943

### 2008 taxonomy[edit]

Uit de Weerd (2008)[2] moved the Urocoptidae to the newly established superfamily Urocoptoidea based on molecular phylogeny research.[2]

The following cladogram based on Bayesian analysis of sequences of 28S ribosomal RNA shows the phylogenic relations within the Urocoptoidea.[2] There are up to 2008 genetically analyzed species of Urocoptidae and one species of Cerionidae marked in khaki color in the cladogram.[2]

Urocoptoidea |  |  |  |  |  |  |  |  |  |  |  |  |

Macroceramus microdon

|

Microceramus pontificus

|  |  |

Torrecoptis holguinensis

|

Autocoptis menkeana

|  |  |

Simplicervix inornata

|

Brachypodella dominicensis gabbi

|  |  |  |  |  |  |

Dissotropis sp. nov. [1]

|

Anisospira lebmanni

Archegocoptis crenata

Eucalodium sp.

Cerionidae

Cerion striatellum

|  |  |

Coelostemma sp. nov. [2]

|

Hendersoniella lux lux

Holospira fortisculpta

Epirobia polygyra

### 2012 taxonomy[edit]

Fred G. Thompson established the family Epirobiidae in 2012 for five genera of snails (Epirobia, Propilsbrya, Pectinistemma, Gyrocion, Prionoloplax), that were previously placed within Urocoptidae.[4] Data from molecular phylogeny research as well as morphology supports monophyly of Holospiridae Pilsbry, 1946 and the Eucalodiidae Crosse & Fischer, 1868.[4]

## Genera[edit]

Genera in the family Urocoptidae include:

Urocoptinae

* Autocoptis Pilsbry, 1902[2]
* Brachypodella L. Pfeiffer, 1840 (as Cylindrella)
* Urocoptis Beck, 1837 - type genus of the family Urocoptidae[3]

Apominae

* Apoma Beck, 1837 - type genus of the subfamily Apomatinae[3]
* Simplicervix Pilsbry, 1903[2]

Brachypodellinae

* Brachypodella Beck, 1837 - type genus of the subfamily Brachypodellinae[3]

Eucalodiinae

* Anisospira Strebel, 1880[2]
* Archegocoptis Pilsbry, 1903[2]
* †Condonella McLellan, 1927[5]
* Dissotropis Bartsch, 1906[2]
* Eucalodium Crosse & P. Fischer, 1868 - type genus of the subfamily Eucalodiinae[3]

Holospirinae

* Coelostemma Dall, 1895[2]
* Hendersoniella Dall, 1905[2]
* Holospira Martens, 1860 - type genus of the subfamily Holospirinae[3]

Microceraminae

* Microceramus Pilsbry & Vanatta, 1898 - type genus of the subfamily Microceraminae[3]
* Macroceramus Guilding, 1828[2]

Tetrentodontinae

* Tetrentodon Pilsbry, 1903 - type genus of the subfamily Tetrentodontinae[3]
* Torrecoptis Bartsch, 1943[2]

Other genera include:[6]

* Allocoptis Thompson & Franz, 1976
* Amphicosmia Pilsbry & Vanatta, 1898
* Anoma
* Arangia
* Bactrocoptis
* Badiofax
* Bostrichocentrum
* Bostrychocentrum
* Callonia
* Centralia
* Cochlodinella
* Coelocentrum
* Geoscala
* Gongylostoma
* Gyraxis
* Haplocion
* Heterocoptis
* Idiostemma
* Liocallonia
* Metastoma
* Mychostoma
* Nesocoptis
* Nodulia
* Oligostylus
* Organocoptis
* Pfeiffericoptis
* Pineria
* Pleurostemma
* Poeycoptis
* Pseudopineria
* Pycnoptychia Pilsbry & Vanatta, 1898
* Sagracoptis
* Spiroceramus
* Spirostemma
* Strophina
* Tomelasmus
* Trilamellaxis
* Uncinicoptis

## References[edit]

1. ^ Pilsbry H. A. (1898). The Nautilus 11(9): 107.
2. ^ a b c d e f g h i j k l m n Uit De Weerd, D. R. (2008). "Delimitation and phylogenetics of the diverse land-snail family Urocoptidae (Gastropoda: Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion". Journal of Molluscan Studies. 74 (4): 317–29. doi:10.1093/mollus/eyn023.
3. ^ a b c d e f g h Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
4. ^ a b Thompson F. G. (2012). "The land snail genus Epirobia and allied genera in México and Central America, with the description of a new family, the Epirobiidae (Gastropoda, Pulmonata, Urocoptoidea)". Bulletin of the Florida Museum of Natural History 51: 167-215. PDF.
5. ^ Roth, B. (2000). "Upper Cretaceous (Campanian) land snails (Gastropoda: Stylommatophora) from Washington and California". Journal of Molluscan Studies. 66 (3): 373–381. doi:10.1093/mollus/66.3.373.
6. ^ "Urocoptidae - MICROCERAMIDAE". Archived from the original on 2012-07-16. Retrieved 2024-01-22.

## Further reading[edit]

* Bartsch, P. (1943). "Notes on Mexican Urocoptid mollusks". Journal of the Washington Academy of Sciences. 33 (2): 54–59.; 7 figs.
* Carlos de la Torre. 1912. New Cuban Urocoptis of the U. cinerea group. The Nautilus, volume 26, number 4, pages 55-58, plate 6.
* Kabat A. R., Herschler R. & González G. A. (2012). "Resolution of taxonomic problems associated with the complex publication history of the seminal Torre and Bartsch monograph on Cuban Urocoptidae (Gastropoda, Pulmonata)". Zootaxa 3362: 43–53. PDF.
* Pilsbry H. A. & Henderson J. B. jr. (February 1913). Two new Cuban Urocoptidae. The Nautilus, volume 26, number 10, pages 109-110.
* (in Spanish) Miguel L. Jaume & Alfredo de la Torre. 1976. Los Urocoptidae de Cuba (Mollusca-Pulmonata). Série 4: Ciencias biologicas; no. 53. La Habana, Cuba: Centro de Información Cientifica y Tecnica, 122 pp.
* Mesa Garcia, R; Jaume, ML (1982). "Zoogeography, ecology, and protection of the Urocoptidae family in Cuba (Gastropoda-Pulmonata). II". Revista Cubana de Medicina Tropical. 34 (2): 172–92. PMID 6763741.
* Richardson C. L. 1991. Urocoptidae: catalogue of species. Tryonia 22: 1–245.
* Tryon G. W. (1903) Manual of Conchology. Second series: Pulmonata. Volume 15. Urocoptidae. Continued by H. A. Pilsbry.

## External links[edit]

Wikimedia Commons has media related to Urocoptidae.

* photos
* photos
* blog Bram's Snail Site with photos

Taxon identifiers|

* Wikidata: Q3074668
* Wikispecies: Urocoptidae
* BOLD: 277415
* CoL: 7NLHB
* EoL: 2662
* Fossilworks: 365180
* GBIF: 2618
* iNaturalist: 171962
* IRMNG: 114082
* ITIS: 77033
* NCBI: 513510
* WoRMS: 932609

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Authority control databases: National |

* Israel

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