A replica of the Berlin specimen of Archaeopteryx, most famous of prehistoric "birds". Modern research considers it unlikely to be a bird ancestor, though it was certainly a close relative of these.
Birds evolved from certain feathered theropod dinosaurs, and there is no real dividing line between birds and non-avian dinosaurs except that some of the former survived the Cretaceous–Paleogene extinction event while the latter did not. For the purposes of this article, a 'bird' is considered to be any member of the clade Aves in the broadest sense.[1] Some dinosaur groups which may or may not be true birds are listed below under Proto-birds.
This page contains a listing of prehistoric bird taxa only known from completely fossilized specimens. These extinctions took place before the Late Quaternary and thus took place in the absence of significant human interference. While the earliest hominids had been eating birds and especially their eggs, human population and technology was simply insufficient to seriously affect healthy bird populations until the Upper Paleolithic Revolution. Rather, reasons for the extinctions listed here are stochastic abiotic events such as bolide impacts, climate changes due to orbital shifts, mass volcanic eruptions etc. Alternatively, species may have gone extinct due to evolutionary displacement by successor or competitor taxa – it is notable that an extremely large number of seabirds have gone extinct during the mid-Tertiary; this seems at least partly due to competition by the contemporary radiation of marine mammals.
The relationships of these taxa are often hard to determine, as many are known only from very fragmentary remains and due to the complete fossilization precluding analysis of information from DNA, RNA or protein sequencing. The taxa listed in this article should be classified with the Wikipedia conservation status category "Fossil".
Before the late 19th century, when minerals were still considered one of the kingdoms of binomial nomenclature, fossils were often treated according to a parallel taxonomy. Rather than assigning them to animal or plant genera, they were treated as mineral genera and given binomial names typically using Osteornis ("bone-bird") or Ornitholithus ("bird fossil") as "genus". The latter name, however, is still in use for an oogenus of fossil bird eggs. Also, other animals (in particular pterosaurs) were placed in these "genera". In sources pre-dating the Linnean system, the above terms are also seen in the more extensive descriptions used to name taxa back then.
Higher-level taxa are presented in likely or suspected phylogenetic order. Genus-level taxa and lower are sorted chronologically, in ascending order (i.e., older taxa first).
The higher-level groups of non-Neornithes are arranged based on the phylogeny proposed by Luis Chiappe,[2] updated and expanded to incorporate recent research.[3] The categories are inclusive in ascending order.
Taxonomic assignments, especially in the pygostylian to early neornithine genera, are still very provisional and subject to quite frequent change.[4][5]
Basal Avialae (extinct)
The most primitive "birds", usually still possessing a long bony tail with generally unfused vertebrae. Not all of these may be on the line of bird ancestors; whether they are not closer to other theropods groups than to the Avialae remains to be thoroughly tested (see Xiaotingia).
The earliest birds with a modern pygostyle: a reduction and fusion of the tail vertebrae; possibly a paraphyletic group. Two types of pygostyle are known, a rod-shaped one found in Confuciusornithidae, Enantiornithes and some non-avian theropods such as Nomingia, and a plowshare-shaped one, only known in the lineage leading to modern birds. It is not certain that the pygostyles found in birds are indeed synapomorphies.[6]
Note that Holtz (2011) also included Zhyraornis in his classification of euenantiornithines, though this genus is more often classified as an ornithuran.[13] Holtz also placed Liaoningornis as an ornithuromorph, though more recent studies have placed it as a close relative of Eoalulavis.[14]
Also called "basal Ornithuromorpha".[15] Essentially modern birds, except many still possess a few primitive features such as teeth or wing claws. These have the plowshare-shaped pygostyle and proper tail fan as seen in most living birds. The taxonomy of this group is confusing; the name "Ornithurae" was first proposed by Ernst Haeckel in 1866 and has been revised in meaning several times since.
The following is a list of primitive euornithian genera and those that cannot be confidently referred to any subgroups, following Holtz (2011).[7]
†YanornisZhou & Zhang 2001 [ArchaeoraptorSloan 1999 (nomen nudum); ArchaeovolansCzerkas & Xu 2002; AberratiodontusGong, Hou & Wang 2004] (Early Cretaceous)
Note that Holtz also included the genera Eurolimnornis, Holbotia, Palaeocursornis and Piksi as euornitheans, though they have since been re-identified as pterosaurs.[16]
Unresolved and basal forms
These modern birds are known from remains that cannot be placed in relation to any one modern group and are neither autapomorphic enough to assign them to own orders. Especially the Late Cretaceous/early Paleogene taxa are probably basal to several modern orders, while later Paleogene taxa often represent extinct lineages outside the modern families.
†"Palaeotringa" vetusMarsh 1870 (Lance Late Cretaceous of Wyoming – Hornerstown Late Cretaceous/?Early Palaeocene of New Jersey, US) - gruiform? anseriform (presbyornithid)?
†PlesiocathartesGaillard 1908 (Late Eocene–? Early Miocene of SW Europe) - cathartid, leptosomid?
†P. europaeusGaillard 1908
†P. gaillardi
†P. wyomingensisWeidig 2006
†P. majorWeidig 2006
†P. geiselensisMayr 2002
†P. kelleriMayr 2002
†Botauroides parvusShufeldt 1915 (Eocene of Wyoming, US) – coliiform?
†Aminornis excavatusAmeghino 1899 (Deseado Early Oligocene of Rio Deseado, Argentina) – gruiform (aramid)?
†Ciconiopsis antarcticaAmeghino 1899 (Deseado Early Oligocene of Patagonia, Argentina) – ciconiiform (ciconiid)?
†Climacarthrus incompletusAmeghino 1899 (Deseado Early Oligocene of Argentina) – falconiform (accipitrid)? A nomen dubium
†Cruschedula revolaAmeghino 1899 (Deseado Early Oligocene of Golfo San Jorge, Argentina) Aves incertae sedis; A nomen dubium
†Dolichopterus viatorMilne-Edwards 1867 [DolicopterusAymard 1856 nomen nudum; CamaskelusAymard 1856 nomen nudum; CamaskelusMilne-Edwards 1867; Dolicopterus viatorAymard 1856 nomen nudum; Camaskelus palustrisAymard 1856 nomen nudum; Camaskelus palustrisMilne-Edwards 1867] (Early Oligocene of Ronzon, France) – charadriiform (charadriid)? Not Dolicopterus as sometimes claimed
†Loncornis erectusAmeghino 1899 (Deseado Early Oligocene of Rio Deseado, Argentina)– gruiform (aramid)?
†Loxornis clivusAmeghino 1894– anatid? (Deseado Early Oligocene of Argentina)
†Palaeopapia eous>(Harrison & Walker 1976) Harrison & Walker 1979 [HowardiaHarrison & Walker 1976 non Berlese & Leonardi 1896; Howardia eousHarrison & Walker 1976] (Hampstead Early Oligocene of Isle of Wight, England) – anseriform?
†Paracygnopterus scottiHarrison & Walker 1979 (Early Oligocene of Belgium and England) – anseriform (anatid)?
†"Pararallus" hassenkampiMartini 1967 (Sieblos Dysodil Early Oligocene of Sieblos, Germany)
†Riacama caligineaAmeghino 1899 (Deseado Early Oligocene of Argentina) – gruiform?
†Smiliornis penetransAmeghino 1899 (Deseado Early Oligocene of Argentina)– gruiform?
†Teracus littoralisMilne-Edwards 1871 [leracus Aymard 1856 nomen nudum; leracus littoralisAymard 1856 nomen nudum] (Early Oligocene of France)
†Teleornis impressusAmeghino 1899 (Deseado Early Oligocene of Argentina)– anatid?
†Pseudolarus guaraniticusAmeghino 1899 (Deseado Early Oligocene – Miocene of Argentina) – gruiform?
Neornithes incerta sedis BMNH PAL 4989 (Hampstead Early Oligocene of Isle of Wight, England) – formerly "Ptenornis" and included in Headonornis; anseriform?
†"Anas" creccoidesvan Beneden 1871 (Early-mid Oligocene of Belgium) – anseriform?
†Headonornis hantoniensis(Lydekker 1891) Harrison & Walker 1976 [Agnopterus hantoniensisLydekker 1891; Presbyornis hantoniensis(Lydekker 1891) Bocheński 1997; Ptenornis] – only BMNH PAL 30325 belongs to this bird, may belong to Presbyornis.
†PresbyornisWetmore 1926 [NautilornisWetmore 1926; ColtoniaHardy 1959] (Paleocene– Early Oligocene)
The diatrymas, a group of huge flightless Paleogene birds of unclear affinities. Traditionally placed within the Gruiformes, they are usually considered a distinct order nowadays and appear closer to the Anseriformes.
†Grallavis edwardsi(Lydekker 1891c) Cheneval 1984 [Propelargus edwardsiLydekker 1891c; Palaeoephippiorhynchus edwardsi(Lydekker 1891)] (Early Miocene) – may be same as Prociconia
The group that includes modern pelicans and cormorants. As presented here paraphyletic; the tropicbird lineage is not part of this group and relationships with Procellariiformes and Sphenisciformes require more research. Also, as the pelicans are at least as close to the Ciconiiformes as to cormorants, the latter group is being recognized as Phalacrocoraciiformes by some recent authors and the core Pelecaniformes are occasionally merged into the Ciconiiformes.
†BorvocarboMourer-Chauviré, Berthet & Hugueney 2004Lua error: Internal error: The interpreter has terminated with signal "24". (Late Oligocene of C Europe) – phalacrocoracid? basal phalacrococacoid?
†B. guillotiMourer-Chauviré, Berthet & Hugueney 2004Lua error: Internal error: The interpreter has terminated with signal "24".
†B. tardatusGöhlich & Mourer-Chauviré 2010Lua error: Internal error: The interpreter has terminated with signal "24".
†LimicorallusKuročkin 1968Lua error: Internal error: The interpreter has terminated with signal "24". (middle Oligocene)
†L. saiensisKuročkin 1968Lua error: Internal error: The interpreter has terminated with signal "24".
†L. carbunculusMayr 2009Lua error: Internal error: The interpreter has terminated with signal "24".
†NambashagWorthy 2011Lua error: Internal error: The interpreter has terminated with signal "24".
†N. billerooensisWorthy 2011Lua error: Internal error: The interpreter has terminated with signal "24".
†N. microglaucusWorthy 2011Lua error: Internal error: The interpreter has terminated with signal "24".
†NectornisCheneval 1984Lua error: Internal error: The interpreter has terminated with signal "24". (Late Oligocene?/Early Miocene – Middle Miocene)
†N. africanusMayr 2014Lua error: Internal error: The interpreter has terminated with signal "24".
†N. miocaenus(Milne-Edwards 1868)Lua error: Internal error: The interpreter has terminated with signal "24". [?Phalacrocorax miocaenus(Milne-Edwards 1868)Lua error: Internal error: The interpreter has terminated with signal "24".]
†N. anatolicus(Mourer-Chauviré 1978) Mlíkovský 1998Lua error: Internal error: The interpreter has terminated with signal "24". [Phalacrocorax anatolicusMourer-Chauviré 1978Lua error: Internal error: The interpreter has terminated with signal "24".]
†Valenticarbo praetermissusHarrison 1979Lua error: Internal error: The interpreter has terminated with signal "24". (Late Pliocene/Early Pleistocene) – a nomen dubium
†Oligocorax Lambrecht 1933
†O. littoralisMilne-Edwards 1868Lua error: Internal error: The interpreter has terminated with signal "24". [Phalacrocorax littoralis(Milne-Edwards 1868)Lua error: Internal error: The interpreter has terminated with signal "24".]
†O. stoeffelensis(Mayr 2007) Mayr 2014Lua error: Internal error: The interpreter has terminated with signal "24". [Oligocorax sp.Mayr 2001Lua error: Internal error: The interpreter has terminated with signal "24".; Borvocarbo stoeffelensisMayr 2007Lua error: Internal error: The interpreter has terminated with signal "24".]
Placement unresolved
†Phalacrocoracidae gen. et sp. indet. (Late Eocene ?–? mid-Oligocene)
Extant genera present in the fossil record
Phalacrocorax (Oligocene –? Recent) – may be several genera. Includes Australocorax, Miocorax
†P. gregoriiDe Vis 1905Lua error: Internal error: The interpreter has terminated with signal "24".
†P. gregorii(De Vis 1905)Lua error: Internal error: The interpreter has terminated with signal "24". [Australocorax vetustusDe Vis 1905Lua error: Internal error: The interpreter has terminated with signal "24".]
†P. chapalensisAlvarez 1977Lua error: Internal error: The interpreter has terminated with signal "24".
†P. destefani(Regalia 1902)Lua error: Internal error: The interpreter has terminated with signal "24". [Paracorax destefaniRegalia 1902Lua error: Internal error: The interpreter has terminated with signal "24".]
†P. femoralis(Miller 1929)Lua error: Internal error: The interpreter has terminated with signal "24". [Miocorax femoralisMiller 1929Lua error: Internal error: The interpreter has terminated with signal "24".]
†P. filyawiEmslie 1995bLua error: Internal error: The interpreter has terminated with signal "24".
†P. goletensisHoward 1965Lua error: Internal error: The interpreter has terminated with signal "24".
†P. ibericusVillalta 1963Lua error: Internal error: The interpreter has terminated with signal "24".
†P. idahensis(Marsh 1870)Lua error: Internal error: The interpreter has terminated with signal "24". [Graculus idahensisMarsh 1870Lua error: Internal error: The interpreter has terminated with signal "24".]
†P. intermedius(Milne-Edwards 1867)Lua error: Internal error: The interpreter has terminated with signal "24". [Phalacrocorax praecarbo, Ardea brunhuberi; Phalacrocorax brunhuberi(Milne-Edwards)Lua error: Internal error: The interpreter has terminated with signal "24".; Botaurites avitus]
†P. kennelli(Howard 1949)Lua error: Internal error: The interpreter has terminated with signal "24".
†P. kuehneanusSchlüter 1991Lua error: Internal error: The interpreter has terminated with signal "24".
†P. longipes(Tugarinov 1940)Lua error: Internal error: The interpreter has terminated with signal "24". [Pliocarbo longipesTugarinov 1940Lua error: Internal error: The interpreter has terminated with signal "24".]
†P. leptopusBrodkorb 1961Lua error: Internal error: The interpreter has terminated with signal "24".
†P. macerBrodkorbLua error: Internal error: The interpreter has terminated with signal "24".
†P. macropus(Cope 1878)Lua error: Internal error: The interpreter has terminated with signal "24". [Graculus macropusCope 1878Lua error: Internal error: The interpreter has terminated with signal "24".]
†P. marinavis(Shufeldt 1915)Lua error: Internal error: The interpreter has terminated with signal "24". [Oligocorax marinavisShufeldt 1915Lua error: Internal error: The interpreter has terminated with signal "24".]
†P. mongoliensisKuročkin 1971Lua error: Internal error: The interpreter has terminated with signal "24".
†P. owreiBrodkorb & Mourer-Chauviré 1984Lua error: Internal error: The interpreter has terminated with signal "24".
†P. reliquusKuročkin 1976Lua error: Internal error: The interpreter has terminated with signal "24".
†P. rogersi(Howard 1932)Lua error: Internal error: The interpreter has terminated with signal "24".
†P. serdicensisBurčak-Abramovič & Nikolov 1984Lua error: Internal error: The interpreter has terminated with signal "24".
†P. tanzaniaeHarrison & Walker 1979Lua error: Internal error: The interpreter has terminated with signal "24".
†P. wetmoreiBrodkorb 1955Lua error: Internal error: The interpreter has terminated with signal "24".
†CopepteryxOlson & Hasegawa 1996Lua error: Internal error: The interpreter has terminated with signal "24".
†C. hexerisOlson & Hasegawa 1996Lua error: Internal error: The interpreter has terminated with signal "24".
†C. titanOlson & Hasegawa 1996Lua error: Internal error: The interpreter has terminated with signal "24".
†Hokkaidornis abashiriensisSakurai, Kimura & Katoh 2008Lua error: Internal error: The interpreter has terminated with signal "24".
†Phocavis maritimusGoedert 1988Lua error: Internal error: The interpreter has terminated with signal "24".
†Plotopterum joaquinensisHoward 1969Lua error: Internal error: The interpreter has terminated with signal "24".
†TonsalaOlson 1980Lua error: Internal error: The interpreter has terminated with signal "24".
†T. hildegardaeOlson 1980Lua error: Internal error: The interpreter has terminated with signal "24".
†T. buchananiDyke, Xia & Habib 2011 Lua error: Internal error: The interpreter has terminated with signal "24".
†Protoplotidae
†Protoplotus beaufortiLambrecht 1931Lua error: Internal error: The interpreter has terminated with signal "24". (Paleocene? – middle Eocene of Sumatra)
Anhingidae – darters
†Meganhinga chilensisAlvarenga 1995Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene)
†MacranhingaNoriega 1992Lua error: Internal error: The interpreter has terminated with signal "24". (giant snake-birds) (Middle/Late Miocene –? Early Pliocene)
†M. paranensisNoriega 1992Lua error: Internal error: The interpreter has terminated with signal "24". [Meganhinga paranensis(Noriega 1992)Lua error: Internal error: The interpreter has terminated with signal "24".]
†M. ranziiAlvarenga & Guilherme 2003Lua error: Internal error: The interpreter has terminated with signal "24".
†"Paranavis" (Middle/Late Miocene) – a nomen nudum
†Giganhinga kiyuensisInderknecht & Noriega 2002Lua error: Internal error: The interpreter has terminated with signal "24". (Late Pliocene/Early Pleistocene)
†A. beckeriEmslie 1998Lua error: Internal error: The interpreter has terminated with signal "24".
†A. hadarensisBrodkorb & Mourer-Chauviré 1982Lua error: Internal error: The interpreter has terminated with signal "24".
†A. laticeps(De Vis 1906)Lua error: Internal error: The interpreter has terminated with signal "24". [Plotus laticepsDe Vis 1906Lua error: Internal error: The interpreter has terminated with signal "24".]
†A. malaguralaMackness 1995Lua error: Internal error: The interpreter has terminated with signal "24".
†A. pannonicaLambrecht 1916Lua error: Internal error: The interpreter has terminated with signal "24".
†A. subvolans(Brodkorb 1956) Becker 1986Lua error: Internal error: The interpreter has terminated with signal "24". [Phalacrocorax subvolansBrodkorb 1956Lua error: Internal error: The interpreter has terminated with signal "24".]
†A. walterbolesiWorthy 2012Lua error: Internal error: The interpreter has terminated with signal "24".
†A. fraileyiCampbell 1996Lua error: Internal error: The interpreter has terminated with signal "24". [?Macranhinga fraileyi(Campbell 1996)Lua error: Internal error: The interpreter has terminated with signal "24".]
†A. grandisMartin & Mengel 1975Lua error: Internal error: The interpreter has terminated with signal "24".
†A. minutaAlvarenga & Guilherme 2003Lua error: Internal error: The interpreter has terminated with signal "24".
†Protopelicanus cuvieriReichenbach 1852Lua error: Internal error: The interpreter has terminated with signal "24".
†Miopelecanus Cheneval 1984
†M. gracilis(Milne-Edwards 1863) Cheneval 1984Lua error: Internal error: The interpreter has terminated with signal "24". [Pelecanus gracilisMilne-Edwards 1863Lua error: Internal error: The interpreter has terminated with signal "24".]
†M. intermedius(Fraas 1870) Cheneval 1984Lua error: Internal error: The interpreter has terminated with signal "24". [Pelecanus intermediusFraas 1870Lua error: Internal error: The interpreter has terminated with signal "24".]
†Pseudodontornis Lambrecht 1930 [Neodontornis Harrison & Walker 1976; Macrodontopteryx Harrison & Walker 1976; Palaeochenoides Schufeldt 1916] (Late Paleocene – London Clay Early Eocene of England)
†P. longidentata Harrison & Walker 1976 [Odontopteryx oweni (Harrison & Walker 1976); Macrodontopteryx oweni Harrison & Walker 1976]
†P. tenuirostris Harrison 1985
†P. tschulensis (Averianov et al. 1991) [Odontopteryx tschulensis]
?Pelagornithidae gen. et sp. indet. (middle Eocene)
†XenerodiopidaeRasmussen 1987Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene)
†Xenerodiopus mycterRasmussen, Olson & Simons 1987Lua error: Internal error: The interpreter has terminated with signal "24".
Ardeidae – herons
†Ardeagrandis arboreaKurockin & Ganya 1972Lua error: Internal error: The interpreter has terminated with signal "24".
†Calcardea junneiGingerich 1987Lua error: Internal error: The interpreter has terminated with signal "24". (Paleocene)
†Matuku otagoenseScofield, Worthy & Tennyson 2010Lua error: Internal error: The interpreter has terminated with signal "24".
†Nyctisoma robustaElżanowski & Zelenkov 2015Lua error: Internal error: The interpreter has terminated with signal "24".
†Palaeophoyx columbianaMcCoy 1963Lua error: Internal error: The interpreter has terminated with signal "24".
†Pikaihao bartleiWorthy et al. 2013Lua error: Internal error: The interpreter has terminated with signal "24".
†Proardea amissa(Milne-Edwards 1892) Lambrecht 1933Lua error: Internal error: The interpreter has terminated with signal "24". [Ardea amissaMilne-Edwards 1892Lua error: Internal error: The interpreter has terminated with signal "24".; Egretta amissa(Milne-Edwards 1892) Mlíkovský & Švec 1989Lua error: Internal error: The interpreter has terminated with signal "24".] (Late Eocene –? Late Oligocene)
†Proardeola walkeriHarrison 1979)Lua error: Internal error: The interpreter has terminated with signal "24". [Ardea FormosaMilne-Edwards 1871 (nomen nudum)Lua error: Internal error: The interpreter has terminated with signal "24".; Ardeola walker(Harrison 1979) Mlíkovský & Švec 1989Lua error: Internal error: The interpreter has terminated with signal "24".] – possibly same as Proardea
†Zeltornis ginsburgiBalouet 1981Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene)
†A. aurelianensisMilne-Edwards 1871Lua error: Internal error: The interpreter has terminated with signal "24". [Proardea aurelianensis(Milne-Edwards 1871)Lua error: Internal error: The interpreter has terminated with signal "24".]
†A. polkensisBrodkorb 1955Lua error: Internal error: The interpreter has terminated with signal "24".
†A. howardaeBrodkorb 1980Lua error: Internal error: The interpreter has terminated with signal "24".
†A. paloccidentalisShufeldt 1892Lua error: Internal error: The interpreter has terminated with signal "24".
†A. sytchevskayaeZelenkov 2011Lua error: Internal error: The interpreter has terminated with signal "24".
Threskiornithidae gen. et sp. indet. NMMP-KU 1301 (Pondaung middle Eocene of Paukkaung, Myanmar) – Threskiornithinae?
†Gerandibis paganus(Milne-Edwards 1868) De Pietri 2013Lua error: Internal error: The interpreter has terminated with signal "24". [Milnea Lydekker1891a non Reichenbach 1866Lua error: Internal error: The interpreter has terminated with signal "24".; Ibis paganaMilne-Edwards 1868Lua error: Internal error: The interpreter has terminated with signal "24".; Eudocimus paganus(Milne-Edvards 1868) Sharpe 1899Lua error: Internal error: The interpreter has terminated with signal "24".; Plegadis paganus(Milne-Edvards 1868) Olson 1981Lua error: Internal error: The interpreter has terminated with signal "24".; Actiornis pagana(Milne-Edwards 1868) Harrison 1976Lua error: Internal error: The interpreter has terminated with signal "24".; Milnea gracilisLydekker 1891aLua error: Internal error: The interpreter has terminated with signal "24".; Plegadis gracilis(Lydekker 1891) Cheneval 1984 non Miller & Bowman 1956Lua error: Internal error: The interpreter has terminated with signal "24".]
†Rhynchaeites messelensisWittich 1898Lua error: Internal error: The interpreter has terminated with signal "24". [PlumumidaHoch 1980Lua error: Internal error: The interpreter has terminated with signal "24".; Plumumida lutetialisHoch 1980Lua error: Internal error: The interpreter has terminated with signal "24".] (middle Eocene of Messel, Germany) – may include Mopsitta
†Vadaravis brownaeSmith, Grande & Clarke 2013Lua error: Internal error: The interpreter has terminated with signal "24".
†G. perplexus(Milne-Edwards 1869) Cheneval 2000Lua error: Internal error: The interpreter has terminated with signal "24". [Ardea perplexaMilne-Edwards 1869Lua error: Internal error: The interpreter has terminated with signal "24".; Proardea perplexus(Milne-Edwards 1869) Gaillard 1939Lua error: Internal error: The interpreter has terminated with signal "24".]
†G. apelexOlson 1985Lua error: Internal error: The interpreter has terminated with signal "24".
†G. olsoniMourer-Chauviré & Geraads 2010Lua error: Internal error: The interpreter has terminated with signal "24".
Platalea
†P. chione(Emslie 1995)Lua error: Internal error: The interpreter has terminated with signal "24". [Ajaia chioneEmslie 1995Lua error: Internal error: The interpreter has terminated with signal "24".]
†P. tiangangensisHou 1987Lua error: Internal error: The interpreter has terminated with signal "24". (Xiacaowan Middle Miocene of Sihong, China)
†P. pharangitesOlson 1981Lua error: Internal error: The interpreter has terminated with signal "24". [Plegadis gracilisMiller & Bowman 1956 non (Lydekker 1891)Lua error: Internal error: The interpreter has terminated with signal "24".] (Late Pliocene of WC US)
†E. leiseyiEmslie 1995aLua error: Internal error: The interpreter has terminated with signal "24". (Early Pleistocene of Florida)
†E. peruvianusCampbell 1979Lua error: Internal error: The interpreter has terminated with signal "24". (Late Pleistocene of Peru; may be living species)
Goliathia andrewsiLambrecht 1930Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene/Early Oligocene of Egypt)
Paludiavis richaeHarrison & Walker 1982Lua error: Internal error: The interpreter has terminated with signal "24". (Late Miocene of Tunisia and Pakistan)
†Eopuffinus kazachstanensisNesov 1986Lua error: Internal error: The interpreter has terminated with signal "24". (Late Paleocene of Zhylga, Kazakhstan)
†Makahala miraeMayr 2015Lua error: Internal error: The interpreter has terminated with signal "24".
†ProaestrelataImber 1985Lua error: Internal error: The interpreter has terminated with signal "24".
†MarinavidaeHarrison & Walker 1977Lua error: Internal error: The interpreter has terminated with signal "24".
†Marinavis longirostrisHarrison & Walker 1977 Lua error: Internal error: The interpreter has terminated with signal "24".
†TytthostonychidaeOlson & Parris 1987Lua error: Internal error: The interpreter has terminated with signal "24".
†Tytthostonyx glauconiticusOlson & Parris 1987Lua error: Internal error: The interpreter has terminated with signal "24".
†DiomedeoididaeFischer 1985Lua error: Internal error: The interpreter has terminated with signal "24".
†Rupelornis van Beneden 1871 [GaviotaFischer 1983 non Miller & Sibley 1941Lua error: Internal error: The interpreter has terminated with signal "24".; ArdeitaCheneval 1996Lua error: Internal error: The interpreter has terminated with signal "24".; DiomedeoidesFischer 1985Lua error: Internal error: The interpreter has terminated with signal "24".; FrigidafronsCheneval 1995Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Oligocene– Early Miocene of C Europe and Iran)
†R. harmati(Kessler 2009) Mayr & Smith 2012Lua error: Internal error: The interpreter has terminated with signal "24". [Diomedeoides harmatiKessler 2009Lua error: Internal error: The interpreter has terminated with signal "24".]
†R. definitusvan Beneden 1871Lua error: Internal error: The interpreter has terminated with signal "24". [Vanellus selysiivan Beneden 1871Lua error: Internal error: The interpreter has terminated with signal "24".; Anas creccoidesvan Beneden 1871Lua error: Internal error: The interpreter has terminated with signal "24".; Gaviota lipsiensisFischer 1983Lua error: Internal error: The interpreter has terminated with signal "24".; Diomedeoides minimusFischer 1985Lua error: Internal error: The interpreter has terminated with signal "24".; Frigidafrons brodkorbiCheneval 1995Lua error: Internal error: The interpreter has terminated with signal "24".; Ardeita gracilisCheneval 1996 (nomen nudum)Lua error: Internal error: The interpreter has terminated with signal "24".; Diomedeoides lipsiensis(Fischer 1983) Mayr et al. 2002Lua error: Internal error: The interpreter has terminated with signal "24".; Diomedeoides brodkorbi(Cheneval 1995) Fischer 2003Lua error: Internal error: The interpreter has terminated with signal "24".]
†R. babaheydariensis(Peters & Hamedani 2000) Mayr & Smith 2012Lua error: Internal error: The interpreter has terminated with signal "24". [Frigidafons babaheydariensisPeters & Hamedani 2000Lua error: Internal error: The interpreter has terminated with signal "24".; Diomedeoides babaheydariensis(Peters & Hamedani 2000) Mayr, Peters & Rietschel 2002Lua error: Internal error: The interpreter has terminated with signal "24".]
†Murunkus subitusPanteleyev & Nesov 1993Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene)
†PlotornisMilne-Edwards 1874Lua error: Internal error: The interpreter has terminated with signal "24". [ChenornisPortis 1884Lua error: Internal error: The interpreter has terminated with signal "24".] (Early – Middle Miocene)
†P. arvernensis(Milne-Edwards 1871) Cheneval 1984aLua error: Internal error: The interpreter has terminated with signal "24". [Puffinus arvernensisMilne-Edwards 1871 nomen nudumLua error: Internal error: The interpreter has terminated with signal "24".; Puffinus arvernensisShufeldt 1896Lua error: Internal error: The interpreter has terminated with signal "24".]
†P. deltortriiMilne-Edwards 1874Lua error: Internal error: The interpreter has terminated with signal "24".
†P. graculoides(Portis 1884) Mayr & Pavia 2014Lua error: Internal error: The interpreter has terminated with signal "24". [Chenornis graculoidesPortis 1884Lua error: Internal error: The interpreter has terminated with signal "24".]
†Tydea septentrionalisMayr & Smith 2012Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene of the North Sea basin)
†Diomedeidae gen. et sp. indet. (Late Oligocene of South Carolina)
†Primodroma bourneiHarrison & Walker 1977Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene)
Prehistoric species of extant genera
†"Oceanodroma" hubbsiMiller 1951Lua error: Internal error: The interpreter has terminated with signal "24". (Hubbs' Storm-petrel)(Capistrano Middle/Late Miocene of Orange County, US)
†Argyrodyptes microtarsusAmeghino 1905Lua error: Internal error: The interpreter has terminated with signal "24". (San Julián Late Eocene/Early Oligocene of Chubut, Argentina)
†Pterodromoides minoricensisSequí et al. 2001Lua error: Internal error: The interpreter has terminated with signal "24".
†P. aquitanicus(Milne-Edwards) Brodkorb 1963Lua error: Internal error: The interpreter has terminated with signal "24".Procellaria aquitanicaMilne-Edwards 1874Lua error: Internal error: The interpreter has terminated with signal "24".
†P. antiquus(Milne-Edwards) Brodkorb 1963cLua error: Internal error: The interpreter has terminated with signal "24".Procellaria antiquaMilne-Edwards 1874Lua error: Internal error: The interpreter has terminated with signal "24".
†P. barnesiHoward 1978Lua error: Internal error: The interpreter has terminated with signal "24".
†P. calhouniHoward 1968Lua error: Internal error: The interpreter has terminated with signal "24".
†P. conradiMarsh 1870Lua error: Internal error: The interpreter has terminated with signal "24".
†P. diatomicusMiller 1925Lua error: Internal error: The interpreter has terminated with signal "24".
†P. felthamiHoward 1949Lua error: Internal error: The interpreter has terminated with signal "24".
†P. gilmoreiChandler 1990Lua error: Internal error: The interpreter has terminated with signal "24".
†P. inceptorWetmore 1930Lua error: Internal error: The interpreter has terminated with signal "24".
†P. kanakoffiHoward 1949Lua error: Internal error: The interpreter has terminated with signal "24".
†P. micraulaxBrodkorb 1963Lua error: Internal error: The interpreter has terminated with signal "24".
†P. mitchelliMiller 1961Lua error: Internal error: The interpreter has terminated with signal "24".
†P. nestoriAlcover 1989Lua error: Internal error: The interpreter has terminated with signal "24".
†P. priscusMiller 1961Lua error: Internal error: The interpreter has terminated with signal "24".
†P. raemdonckii(Van Beneden 1871) Brodkorb 1962Lua error: Internal error: The interpreter has terminated with signal "24". [Larus Raemdonckii Beneden 1871]
†P. tedfordiHoward 1971Lua error: Internal error: The interpreter has terminated with signal "24".
OpisthocomidaeSwainson 1837Lua error: Internal error: The interpreter has terminated with signal "24". [Foratidae Olson 1992Lua error: Internal error: The interpreter has terminated with signal "24".; Onychopterygidae Cracraft 1971Lua error: Internal error: The interpreter has terminated with signal "24".]
†Foro panariumOlson 1992Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene) - cuculiform?
†Hoatzi panariumThomas 1996Lua error: Internal error: The interpreter has terminated with signal "24".
†Onychopteryx simpsoniCracraft 1971Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene of Argentina) – falconid? A nomen dubium
†Protoazin parisiensisMayr & De Pietri 2014Lua error: Internal error: The interpreter has terminated with signal "24".
†Namibiavis senutaeMourer-Chauviré 2003Lua error: Internal error: The interpreter has terminated with signal "24". (Late Oligocene of Namibia)
†Hoazinavis lacustrisAlvarenga, Mayr & Mourer-Chauviré 2011Lua error: Internal error: The interpreter has terminated with signal "24". (Miocene of Brazil)
†Hoazinoides magdalenaeMiller 1953Lua error: Internal error: The interpreter has terminated with signal "24". (Late Miocene of Colombia)
Musophagidae gen. et sp. indet. (Late Oligocene – Middle Miocene of WC Europe)
Musophagidae gen. et sp. indet. (Egypt)
†VeflintornisKashin 1976Lua error: Internal error: The interpreter has terminated with signal "24". [ApopempsisBrodkorb 1971 non Schenkling 1903Lua error: Internal error: The interpreter has terminated with signal "24".] (Middle Miocene)
†V. meini(Ballmann 1969)Lua error: Internal error: The interpreter has terminated with signal "24". [Musophaga meiniBallmann 1969Lua error: Internal error: The interpreter has terminated with signal "24".]
†V. africanus(Harrison 1980)Lua error: Internal error: The interpreter has terminated with signal "24". [Musophaga africanusHarrison 1980Lua error: Internal error: The interpreter has terminated with signal "24".; Apopempsis africanus(Harrison 1980)Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Miocene)
†TeratornithidaeMiller 1909Lua error: Internal error: The interpreter has terminated with signal "24".– teratorns
†Argentavis magnificensCampbell & Tonni 1980Lua error: Internal error: The interpreter has terminated with signal "24". (Late Miocene)
†Aiolornis incredibilis(Howard 1952) Campbell, Scott & Springer 1999Lua error: Internal error: The interpreter has terminated with signal "24". [Teratornis incredibilisHoward 1952Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Pliocene – Late Pleistocene)
†Oscaravis olsoni(Arredondo & Arredondo 2002) Suarez & Olson 2009Lua error: Internal error: The interpreter has terminated with signal "24". [Teratornis olsoniArredondo & Arredondo 2002Lua error: Internal error: The interpreter has terminated with signal "24".] (Cuban teratorn)(Pleistocene)
†Cathartornis gracilisMiller 1910Lua error: Internal error: The interpreter has terminated with signal "24".
†Taubatornis campbelliOlson & Alvarenga 2002Lua error: Internal error: The interpreter has terminated with signal "24".
†TeratornisMiller 1909Lua error: Internal error: The interpreter has terminated with signal "24". (Early Pleistocene – Late Pleistocene)
†T. woodburnensisCampbell & Stenger 2002Lua error: Internal error: The interpreter has terminated with signal "24".
†T. merriamiMiller 1909Lua error: Internal error: The interpreter has terminated with signal "24". (Merriam's teratorn)
†Phasmagyps patritusWetmore 1927Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene)
†Hadrogyps aigialeusEmslie 1988Lua error: Internal error: The interpreter has terminated with signal "24". (American dwarf vulture) (Middle Miocene)
†PliogypsBecker 1986Lua error: Internal error: The interpreter has terminated with signal "24". (Miocene vulture) (Late Miocene – Late Pliocene)
†P. charonBecker 1986Lua error: Internal error: The interpreter has terminated with signal "24".
†P. fisheriTordoff 1959Lua error: Internal error: The interpreter has terminated with signal "24".
†Perugyps diaziStucchi & Emslie 2005Lua error: Internal error: The interpreter has terminated with signal "24". (Peruvian vulture) (Late Miocene/Early Pliocene)
†Dryornis (Argentinean vulture) Moreno & Mercerat, 1891Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene – Late Pliocene) – similar to the extant genus Vultur
†D. pampeanusMoreno & Mercerat, 1891Lua error: Internal error: The interpreter has terminated with signal "24".
†D. hatcheriDegrange 2022Lua error: Internal error: The interpreter has terminated with signal "24".
†Aizenogyps toomeyaeEmslie 1998Lua error: Internal error: The interpreter has terminated with signal "24". (South American vulture) (Late Pliocene)
†Breagyps clarkiMiller 1938Lua error: Internal error: The interpreter has terminated with signal "24". (long-legged vulture) (Late Pleistocene)
†Geronogyps reliquusCampbell 1979Lua error: Internal error: The interpreter has terminated with signal "24". (Late Pleistocene)
†Wingegyps cartelleiAlvarenga & Olson 2004Lua error: Internal error: The interpreter has terminated with signal "24". (Amazonian vulture) (Late Pleistocene)
†Parasarcoramphus milneedwardsiMourer-Chauviré 2002Lua error: Internal error: The interpreter has terminated with signal "24".
Placement unresolved
Cathartidae gen. et sp. indet. (Late Oligocene of Mongolia)
Cathartidae gen. et sp. indet. (Late Miocene/Early Pliocene of Lee Creek Mine, US)
Cathartidae gen. et sp. indet. (Middle Pliocene of Argentina)
†V. fossilisMoreno & Mercerat 1891Lua error: Internal error: The interpreter has terminated with signal "24". [Cathartes fossilis; Sarcoramphus fossilis]
†HorusornithidaeMourer-Chauviré 1991Lua error: Internal error: The interpreter has terminated with signal "24".
†Horusornis vianeyliaudaeMourer-Chauviré 1991Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene)
†Amanuensis pickfordiMourer-Chauviré 2003Lua error: Internal error: The interpreter has terminated with signal "24".
†PelargopappusStejneger 1885Lua error: Internal error: The interpreter has terminated with signal "24".(Late Eocene/Early Oligocene – Late Oligocene/Early Miocene of France) – formerly Amphiserpentarius/Amynoptilon/Pelargopsis
†P. schlosseri
†P. magnus
Accipitridae – hawks, eagles and Old World vultures
Milvoides (Late Eocene)
Aquilavus (Late Eocene/Early Oligocene – Early Miocene)
†MasillaraptoridaeMayr 2009Lua error: Internal error: The interpreter has terminated with signal "24".
†Masillaraptor parvunguisMayr 2006Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene of Messel, Germany)
†Danielsraptor phorusrhacoidesMayr & Kitchener 2022Lua error: Internal error: The interpreter has terminated with signal "24". (early Eocene of Essex, England)
†Lagopterus minutusMoreno & Mercerat 1891Lua error: Internal error: The interpreter has terminated with signal "24".
†Parvulivenator watteliHarrison 1982Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene)
†Stintonornis mitchelliHarrison 1984Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene)
†Badiostes patagonicusAmeghino 1895Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene)
†Pediohierax ramenta(Wetmore 1936) Becker 1987Lua error: Internal error: The interpreter has terminated with signal "24". [Falco ramentaWetmore 1936Lua error: Internal error: The interpreter has terminated with signal "24".] - (Middle Miocene)
†Thegornis musculosusAmeghino 1895Lua error: Internal error: The interpreter has terminated with signal "24". [Thegornis debilisAmeghino 1895Lua error: Internal error: The interpreter has terminated with signal "24".; Buteo musculosus(Ameghino 1895) Agnolin 2006Lua error: Internal error: The interpreter has terminated with signal "24".]
†Petrosushkinia pliocaena(Tugarinov 1935) Zelenkov 2015Lua error: Internal error: The interpreter has terminated with signal "24". [SushkiniaTugarinov 1935 non Martynov 1930Lua error: Internal error: The interpreter has terminated with signal "24".; Sushkinia pliocaenaTugarinov 1935Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Pliocene)
†FluvioviridavidaeMayr 2005Lua error: Internal error: The interpreter has terminated with signal "24".
†Fluvioviridavis platyrhamphusMayr & Daniels 2001Lua error: Internal error: The interpreter has terminated with signal "24". (Green River Early Eocene of N America)
†Wyomingcypselus pohliWeidig 2003 nomen nudumLua error: Internal error: The interpreter has terminated with signal "24".
†Procypseloides mourerchauvireae(Mlíkovský) Harrison & Walker 1977Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene/Early Oligocene – Early Miocene)
†Laputavis robusta(Dyke 2001a) Dyke 2001bLua error: Internal error: The interpreter has terminated with signal "24". [LaputaDyke 2001a nonLua error: Internal error: The interpreter has terminated with signal "24".; Laputa robustaDyke 2001aLua error: Internal error: The interpreter has terminated with signal "24".]
†Palescyvus escampensisKarhu 1988Lua error: Internal error: The interpreter has terminated with signal "24".
†ScaniacypselusHarrison 1984Lua error: Internal error: The interpreter has terminated with signal "24". (Early – middle Eocene)
†S. wardiHarrison 1984Lua error: Internal error: The interpreter has terminated with signal "24".
†S. szarskii(Peters 1985) Meyr & Peters 1999Lua error: Internal error: The interpreter has terminated with signal "24". [Aegialornis szarskiiPeters 1985Lua error: Internal error: The interpreter has terminated with signal "24".; Primapus szarskii(Peters 1985) Mourer-Chauviré 1988Lua error: Internal error: The interpreter has terminated with signal "24".]
†AegialornithidaeLydekker 1891Lua error: Internal error: The interpreter has terminated with signal "24".
†Primapus lackiHarrison & Walker 1975Lua error: Internal error: The interpreter has terminated with signal "24". [?Procuculus minutusHarrison & Walker 1977Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Eocene) ?apodid
†AegialornisLydekker 1891Lua error: Internal error: The interpreter has terminated with signal "24". [TachyornisMilne-Edwards 1892Lua error: Internal error: The interpreter has terminated with signal "24".; BelornisMilne-Edwards 1893Lua error: Internal error: The interpreter has terminated with signal "24".; MesogiornisMlíkovský 2002Lua error: Internal error: The interpreter has terminated with signal "24".](Early? – Late Eocene)
†A. germanicusMlíkovský 2002Lua error: Internal error: The interpreter has terminated with signal "24".
†A. gallicusLydekker 1891Lua error: Internal error: The interpreter has terminated with signal "24". [Tachyornis hirundoMilne-Edwards 1891Lua error: Internal error: The interpreter has terminated with signal "24".]
†A. wetmoreiCollins 1976Lua error: Internal error: The interpreter has terminated with signal "24". [Mesogiornis wetmorei(Collins 1976) Mlíkovský 2002Lua error: Internal error: The interpreter has terminated with signal "24".]
†A. broweriCollins 1976Lua error: Internal error: The interpreter has terminated with signal "24". [Mesogiornis broweri(Collins 1976) Mlíkovský 2002Lua error: Internal error: The interpreter has terminated with signal "24".]
†EocypselidaeHarrison 1984Lua error: Internal error: The interpreter has terminated with signal "24". – apodiform (hemiprocnid?)? caprimulgiform? basal Cypselomorphae?
†EocypselusHarrison 1984Lua error: Internal error: The interpreter has terminated with signal "24". (Late Paleocene ?- Early Eocene of NC Europe)
†E. vincentiHarrison 1984 Lua error: Internal error: The interpreter has terminated with signal "24".
†E. roweiKsepka et al. 2013Lua error: Internal error: The interpreter has terminated with signal "24".
†CypselavidaeMourer-Chauviré 2006Lua error: Internal error: The interpreter has terminated with signal "24".
†Parargornis messelensisMayr 2003Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene) jungornithid, trochilid, basal as Argornis?
†Argornis caucasicusKarhu 1999Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene) ?Trochilidae
†Cypselavus gallicusGaillard 1908Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene Early Oligocene) aegialornithid or hemiprocnid
†JungornithidaeKarhu 1988Lua error: Internal error: The interpreter has terminated with signal "24".
†JungornisKarhu 1988Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene of N Caucasus, Russia)
†J. tesselatusKarhu 1988Lua error: Internal error: The interpreter has terminated with signal "24".
†J. geraldmayriMourer-Chauviré & Sigé 2006Lua error: Internal error: The interpreter has terminated with signal "24".
Trochilidae hummingbirds
†EurotrochilusMayr 2004Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene)
†E. inexpectatusMayr 2004Lua error: Internal error: The interpreter has terminated with signal "24".
†E. noniewicziBochenski & Bochenski 2008Lua error: Internal error: The interpreter has terminated with signal "24".
Placement unresolved
Trochilidae sp. et gen. indet. (Bahamas, West Indies)
†Botauroides parvusShufeldt 1915Lua error: Internal error: The interpreter has terminated with signal "24". (Eocene of Wyoming, US)
†Eobucco brodkorbiFeduccia & Martin 1976Lua error: Internal error: The interpreter has terminated with signal "24". - sandcoleid?
†Eocolius walkeriDyke & Waterhouse 2001Lua error: Internal error: The interpreter has terminated with signal "24". (London Clay Early Eocene of Walton-on-the-Naze, England) - sandcoleid or coliid
†LimnatornisMilne-Edwards 1871Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene of Saint-Gérand-le-Puy, France) - coliid? (Urocolius?)
†Coliiformes gen. et sp. indet. (Late Miocene of Kohfidisch, Austria)[20]
†UintornisMarsh 1872Lua error: Internal error: The interpreter has terminated with signal "24". - sandcoleid?
Family †ChascacocoliidaeZelenkov & Dyke 2008Lua error: Internal error: The interpreter has terminated with signal "24".
Genus †ChascacocoliusHoude & Olson 1992Lua error: Internal error: The interpreter has terminated with signal "24". (Late Paleocene ?- Early Eocene) - basal? sandcoleid?
Family †SelmeidaeZelenkov & Dyke 2008Lua error: Internal error: The interpreter has terminated with signal "24".
†Selmes absurdipesPeters 1999Lua error: Internal error: The interpreter has terminated with signal "24". (Middle Eocene ?-Late Oligocene of C Europe) - coliid? (synonym of Primocolius?)
Family †SandcoleidaeHoude & Olson 1992 sensu Mayr & Mourer-Chauviré 2004Lua error: Internal error: The interpreter has terminated with signal "24".
†Sandcoleus copiosusHoude & Olson 1992Lua error: Internal error: The interpreter has terminated with signal "24". (Paleocene)
†Anneavis anneaeHoude & Olson 1992Lua error: Internal error: The interpreter has terminated with signal "24".
†Eoglaucidium pallasFischer 1987Lua error: Internal error: The interpreter has terminated with signal "24".
Family ColiidaeSwainson 1837 sensu Mayr & Mourer-Chauviré 2004Lua error: Internal error: The interpreter has terminated with signal "24".
†PrimocoliusMourer-Chauviré 1988Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene/Oligocene of Quercy, France)
†OligocoliusMayr 2000Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene of Frauenweiler, Germany)
†Masillacolius brevidactylusMayr & Peters 1998Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene of Messel, Germany)
Extant genera present in the fossil record
Colius [NecrornisMilne-Edwards 1871Lua error: Internal error: The interpreter has terminated with signal "24".]
†Zealandornithidae
†Zealandornis relictusWorthy et al., 2022Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene of Otago, New Zealand)
†Quasisyndactylus longibrachisMayr 1998Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene) – alcediniform, basal?
†Paracoracias occidentalisClarke et al. 2009Lua error: Internal error: The interpreter has terminated with signal "24".
†Cryptornis antiquusGervais 1848–52 [nomen nudum]Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene) – bucerotid? geranopterid?
†Protornis glarniensisvon Meyer 1856Lua error: Internal error: The interpreter has terminated with signal "24". (Oligocene) – basal to motmotids and meropids?
† Coraciiformes gen. et spp. indet. PQ 1216, QU 15640 (Late Eocene) – 2 species
†GeranopteridaeMayr & Mourer-Chauviré 2000Lua error: Internal error: The interpreter has terminated with signal "24".
†GeranopterusMilne-Edvards 1892Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene – Early Miocene)
†G. alatus(Mayr & Mourer-Chauvire 2000)Lua error: Internal error: The interpreter has terminated with signal "24".
†G. bohemicus(Mlikovsky 1999) Mourer-Chauvire 1999Lua error: Internal error: The interpreter has terminated with signal "24". [Nupharanassa bohemicaMlikovsky 1999Lua error: Internal error: The interpreter has terminated with signal "24".]
†G. milneendwardsiMayr & Mourer-Chauvire 2000Lua error: Internal error: The interpreter has terminated with signal "24".
†EocoraciidaeMayr & Mourer-Chauviré 2000Lua error: Internal error: The interpreter has terminated with signal "24".
†Eocoracias brachypteraMayr & Mourer-Chauviré 2000Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene)
†PrimobucconidaeFeduccia & Martin 1976 sensu Mayr, Mourer-Chauviré & Weidig 2004Lua error: Internal error: The interpreter has terminated with signal "24".
†PrimobuccoBrodkorb 1970Lua error: Internal error: The interpreter has terminated with signal "24". [NeanisBrodkorb 1965 non Gistl 1848Lua error: Internal error: The interpreter has terminated with signal "24".] (Early – middle Eocene)
†P. kestneri Feduccia 1973Lua error: Internal error: The interpreter has terminated with signal "24". [Neanis kestneri(Feduccia 1973) Feduaccia 1976Lua error: Internal error: The interpreter has terminated with signal "24".]
†P. frugilegusMayr, Mourer-Chauviré & Weidig 2004Lua error: Internal error: The interpreter has terminated with signal "24".
†P. mcgrewiBrodkorb 1970Lua error: Internal error: The interpreter has terminated with signal "24".
†P. perneriMayr, Mourer-Chauviré & Weidig 2004Lua error: Internal error: The interpreter has terminated with signal "24".
†Euroceros bulgaricusBoev & Kovachev 2007Lua error: Internal error: The interpreter has terminated with signal "24". (Late Miocene of Hadzhidimovo, Bulgaria)
†Foshanornis songiZhao et al. 2015Lua error: Internal error: The interpreter has terminated with signal "24".
†Septentrogon madseni Kristoffersen 2002 (Fur Late Paleocene/Early Eocene of Ejerslev, Denmark)
†Paratrogon gallicus(Milne-Edwards 1871) Lambrecht 1933Lua error: Internal error: The interpreter has terminated with signal "24". [Trogon gallicusMilne-Edwards 1871Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Miocene of France)
†PrimotrogonMayr 1999Lua error: Internal error: The interpreter has terminated with signal "24". [MasillatrogonMayr 2009Lua error: Internal error: The interpreter has terminated with signal "24".] (middle Eocene of Messel, Germany? - Early Oligocene of France)
†P. wintersteiniMayr 1999Lua error: Internal error: The interpreter has terminated with signal "24".
†P. pumilioMayr 2005Lua error: Internal error: The interpreter has terminated with signal "24". [Masillatrogon pumilio(Mayr 2005) Mayr 2009Lua error: Internal error: The interpreter has terminated with signal "24".]
These fossil taxa cannot be assigned to any major group with reasonable certainty. The "proto-birds" above are of some indeterminate basal position in the entire avialan (and paravian) radiation, but known from such diagnostic material that their relationships at the family level are known. In contrast, the taxa here have a hypodigm that is usually just sufficient for giving them a valid scientific name, but not for phylogenetic purposes beyond classing them as pygostylians or more modern birds. Some, however, are known from such fragmentary remains that the possibility that they are non-avian "reptiles" such as dinosaurs cannot be ruled out at present.
†"Ichthyornis" minusculus Nesov 1990 (Bissekty Late Cretaceous of Kyzyl Kum, Uzbekistan) – enantiornithine?
↑Brodkob, Pierce (1963). "Catalogue of fossil birds 1- Archaeopterygiformes through Ardeiformes". Biological Sciences, Bulletin of the Florida State Museum7 (4): 180–293.
↑O'Connor, J. K.; Zhang, Y.; Chiappe, L. M.; Meng, Q.; Quanguo, L.; Di, L. (2013). "A new enantiornithine from the Yixian Formation with the first recognized avian enamel specialization". Journal of Vertebrate Paleontology33 (1): 1–12. doi:10.1080/02724634.2012.719176. Bibcode: 2013JVPal..33....1O.
↑Tennyson, A.J.D.; Worthy, T.H.; Jones, C.M.; Scofield, R.P.; Hand, S.J. (2010). "Moa's Ark: Miocene fossils reveal the great antiquity of moa (Aves: Dinornithiformes) in Zealandia". Records of the Australian Museum62: 105–114. doi:10.3853/j.0067-1975.62.2010.1546.
↑ 18.018.1Nikita V. Zelenkov; Thomas A. Stidham; Nicolay V. Martynovich; Natalia V. Volkova; Qiang Li; Zhuding Qiu (2018). "The middle Miocene duck Chenoanas (Aves, Anatidae): new species, phylogeny and geographical range". Papers in Palaeontology4 (3): 309–326. doi:10.1002/spp2.1107.
↑Suárez, William; Olson, Storrs L. (2021). "A new fossil raptor (Accipitridae: Buteogallus) from Quaternary cave deposits in Cuba and Hispaniola, West Indies". Bulletin of the British Ornithologists' Club141 (3). doi:10.25226/bboc.v141i3.2021.a3.
↑Similar to Urocolius and Limnatornis (if distinct): Mlíkovský (2002)
Lua error: Internal error: The interpreter has terminated with signal "24".
References
Chiappe, Luis M. (2001): The rise of birds. In: Briggs, Derek E.G. & Crowther, P.R. (eds.): Palaeobiology II: A Synthesis: 102-106. Cambridge University Press, Cambridge, UK.
Chiappe, Luis M. (2002): Basal bird phylogeny: problems and solutions. In: Chiappe, L.M. and Witmer, L.M. (eds.): Mesozoic Birds: Above the Heads of Dinosaurs: 448-472. University of California Press, Berkeley, US.
Gauthier, Jacques A. & de Queiroz, Kevin (2001): Feathered dinosaurs, flying dinosaurs, crown dinosaurs, and the name "Aves". In: Gauthier, Jacques & Gall, L.F. (eds.): New Perspectives on the Origin and Early Evolution of Birds: Proceedings of the International Symposium in Honor of John H. Ostrom: 7-41. Peabody Museum of Natural History, Yale University, New Haven, Connecticut.
Mortimer, Michael (2004): The Theropod Database: Phylogeny of taxa. Retrieved 2013-MAR-02.
Olson, Storrs L. (1985): The fossil record of birds. In: Farner, D.S.; King, J.R. & Parkes, Kenneth C. (eds.): Avian Biology8: 79-238. Academic Press, New York. Not in copyright; PDF fulltext