List of fossil bird genera

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A replica of the Berlin specimen of Archaeopteryx, most famous of prehistoric "birds". Modern research considers it unlikely to be a bird ancestor, though it was certainly a close relative of these.

Birds evolved from certain feathered theropod dinosaurs, and there is no real dividing line between birds and non-avian dinosaurs except that some of the former survived the Cretaceous–Paleogene extinction event while the latter did not. For the purposes of this article, a 'bird' is considered to be any member of the clade Aves in the broadest sense.[1] Some dinosaur groups which may or may not be true birds are listed below under Proto-birds.

This page contains a listing of prehistoric bird taxa only known from completely fossilized specimens. These extinctions took place before the Late Quaternary and thus took place in the absence of significant human interference. While the earliest hominids had been eating birds and especially their eggs, human population and technology was simply insufficient to seriously affect healthy bird populations until the Upper Paleolithic Revolution. Rather, reasons for the extinctions listed here are stochastic abiotic events such as bolide impacts, climate changes due to orbital shifts, mass volcanic eruptions etc. Alternatively, species may have gone extinct due to evolutionary displacement by successor or competitor taxa – it is notable that an extremely large number of seabirds have gone extinct during the mid-Tertiary; this seems at least partly due to competition by the contemporary radiation of marine mammals.

The relationships of these taxa are often hard to determine, as many are known only from very fragmentary remains and due to the complete fossilization precluding analysis of information from DNA, RNA or protein sequencing. The taxa listed in this article should be classified with the Wikipedia conservation status category "Fossil".

Before the late 19th century, when minerals were still considered one of the kingdoms of binomial nomenclature, fossils were often treated according to a parallel taxonomy. Rather than assigning them to animal or plant genera, they were treated as mineral genera and given binomial names typically using Osteornis ("bone-bird") or Ornitholithus ("bird fossil") as "genus". The latter name, however, is still in use for an oogenus of fossil bird eggs. Also, other animals (in particular pterosaurs) were placed in these "genera". In sources pre-dating the Linnean system, the above terms are also seen in the more extensive descriptions used to name taxa back then.


Taxonomic list of fossil prehistoric birds

Higher-level taxa are presented in likely or suspected phylogenetic order. Genus-level taxa and lower are sorted chronologically, in ascending order (i.e., older taxa first).

The higher-level groups of non-Neornithes are arranged based on the phylogeny proposed by Luis Chiappe,[2] updated and expanded to incorporate recent research.[3] The categories are inclusive in ascending order.

Taxonomic assignments, especially in the pygostylian to early neornithine genera, are still very provisional and subject to quite frequent change.[4][5]

Basal Avialae (extinct)

The most primitive "birds", usually still possessing a long bony tail with generally unfused vertebrae. Not all of these may be on the line of bird ancestors; whether they are not closer to other theropods groups than to the Avialae remains to be thoroughly tested (see Xiaotingia).

Basal Pygostylia (extinct)

The earliest birds with a modern pygostyle: a reduction and fusion of the tail vertebrae; possibly a paraphyletic group. Two types of pygostyle are known, a rod-shaped one found in Confuciusornithidae, Enantiornithes and some non-avian theropods such as Nomingia, and a plowshare-shaped one, only known in the lineage leading to modern birds. It is not certain that the pygostyles found in birds are indeed synapomorphies.[6]

Enantiornithes (extinct)

Reconstruction of Iberomesornis romerali, a tiny primitive enantiornithine.
It was no bigger than a modern-day finch.

The taxonomic list of enantiornithine groups presented here follows a summary published by Thomas R. Holz Jr. in 2011.[7]

Enantiornithes

Note that Holtz (2011) also included Zhyraornis in his classification of euenantiornithines, though this genus is more often classified as an ornithuran.[13] Holtz also placed Liaoningornis as an ornithuromorph, though more recent studies have placed it as a close relative of Eoalulavis.[14]

Basal Euornithes (extinct)

Also called "basal Ornithuromorpha".[15] Essentially modern birds, except many still possess a few primitive features such as teeth or wing claws. These have the plowshare-shaped pygostyle and proper tail fan as seen in most living birds. The taxonomy of this group is confusing; the name "Ornithurae" was first proposed by Ernst Haeckel in 1866 and has been revised in meaning several times since.

The following is a list of primitive euornithian genera and those that cannot be confidently referred to any subgroups, following Holtz (2011).[7]

Note that Holtz also included the genera Eurolimnornis, Holbotia, Palaeocursornis and Piksi as euornitheans, though they have since been re-identified as pterosaurs.[16]

Basal Ornithurae (extinct)

  • Limenavis patagonica Clarke & Chiappe 2001 (Late Cretaceous)– paleognath?
  • Palintropidae
    • Palintropus retusus (Marsh 1892) Brodkorb 1970 [Cimolopteryx retusa Marsh 1892; Apatornis retusus (Marsh 1892) Brodkorb 1962] (Late Cretaceous) (?Galliformes: Quercymegapodiidae)
  • Ambiortiformes Kuročkin 1982
  • Apsaraviformes Kuročkin 1982
  • Gansuiformes Hou & Liu 1984
    • Gansuidae Hou & Liu 1984
      • Gansus Hou & Liu 1984 (Early Cretaceous)
  • Ichthyornithiformes Marsh 1873 [Pteropappi Stejneger 1885]
    • Apatornithidae Fürbringer 1888
      • Apatornis celer (Marsh 1873) Marsh 1873 [Ichthyornis celer Marsh 1873] (Late Cretaceous)
      • Guildavis tener (Marsh 1880) Clarke 2004 [Ichthyornis tener Marsh 1880] (Cretaceous of Wallace County, US)
      • Iaceornis marshi Clarke 2004 (Late Cretaceous)
    • Ichthyornithidae Marsh 1873
      • Ichthyornis Marsh 1872 [Angelinornis Kashin 1972; Colonosaurus Marsh 1872; Graculavus Marsh 1873 non Marsh 1872; Plegadornis Wetmore 1962 non Brehm 1855] (Late Cretaceous)
  • Hesperornithiformes Fürbringer 1888 (Large, toothed, loon-like diving birds)
    Hesperornis
    • Chupkaornis keraorum Tanaka et al. 2017
    • Potamornis skutchi Elzanowski, Paul & Stidham 2000 (Late Cretaceous)
    • Pasquiaornis Tokaryk, Cumbaa & Storer 1997 (Late Cretaceous)
    • Enaliornithidae Fürbringer 1888
      • Enaliornis Seeley 1876 [Palaeocolyntus (sic) Seeley 1864 nomen nudum; Palaeocolymbus Seeley 1869; Pelargonis (sic) Seeley 1864 nomen nudum; Pelagornis Seeley 1866 nomen nudum non Lartet 1857] (Early Cretaceous)
    • Baptornithidae AOU 1910
      • Baptornis Marsh 1877 [Parascaniornis Lambrecht 1933]
    • Brodavidae Martin, Kuročkin & Tokaryk 2012
      • Brodavis Martin, Kuročkin & Tokaryk 2012 (Late Cretaceous)
    • Judinornithidae Nesov 1983
    • Hesperornithidae Marsh 1880

Neornithes

The subclass that contains all modern birds.

Unresolved and basal forms
These modern birds are known from remains that cannot be placed in relation to any one modern group and are neither autapomorphic enough to assign them to own orders. Especially the Late Cretaceous/early Paleogene taxa are probably basal to several modern orders, while later Paleogene taxa often represent extinct lineages outside the modern families.

  • Australornis lovei Mayr & Scofield 2014 (late early Paleocene)
  • Gallornis straeleni Lambrecht 1931 (Late Cretaceous)
  • Ceramornis major Brodkorb 1963 (Late Cretaceous) – charadriiform?
  • †"Presbyornithidae" gen. et sp. indet. (Barun Goyot Late Cretaceous of Udan Sayr, Mongolia) - anseriform (presbyornithid)?
  • Torotix clemensi Brodkorb 1963 (Late Cretaceous) – pelecaniform, charadriiform, procellariiform or phoenicopteriform
  • Neornithes incerta sedis Kurochkin 1995 (Nemegt Late Cretaceous of S Mongolia) - phalacrocoracid?
  • Neornithes incerta sedis AMNH FR 25272 (Lance Creek Late Cretaceous of Converse County, US) - phalacrocoracid?
  • Neornithes incerta sedis PVPH 237 (Portezuelo Late Cretaceous of Sierra de Portezuelo, Argentina) - galliform?
  • Neornithes incerta sedis UCMP 117598 (Hell Creek Late Cretaceous of Bug Creek West, US)
  • Neornithes incerta sedis UCMP 117599 (Hell Creek Late Cretaceous of Bug Creek West, US) – anseriform?
  • "Lonchodytes" pterygius Brodkorb 1963 (Late Cretaceous/?Early Palaeocene) – charadriiform?
  • Novacaesareala hungerfordi Parris & Hope 2002 (Late Cretaceous/Early Palaeocene) – related to Torotix?
  • "Palaeotringa" vetus Marsh 1870 (Lance Late Cretaceous of Wyoming – Hornerstown Late Cretaceous/?Early Palaeocene of New Jersey, US) - gruiform? anseriform (presbyornithid)?
  • Volgavis marina Nessov & Jarkow 1989 (Early Palaeocene of Volgograd, Russia) – charadriiform? phalacrocoraciform?
  • Tshulia litorea Nessov 1988 (Late Paleocene of Zhylga, Kazakhstan)
  • Eupterornis remensis Lemoine 1878 (Paleocene of Cernay, France) – charadriiform (larid?)? gaviiform?
  • Gradiornis walbeckensis Mayr 2007 (Paleocene of Walbeck, Germany) – cariamid?
  • "Messelornis" russelli Mourer-Chauviré 1995 sensu Mayr 2007 (Paleocene of Cernay, France) – messelornithid?
  • Walbeckornis creber Mayr 2007 (Paleocene of Walbeck, Germany) – charadriiform? messelornithid?
  • Neornithes incerta sedis (Late Paleocene/Early Eocene of Ouled Abdoun Basin, Morocco) – charadriiform, ciconiiform, gruiform?
  • Argillipes Harrison & Walker 1977 (London Clay Early Eocene of England) – galliform?
    • A. aurorum Harrison & Walker 1977
    • A. magnus Harrison & Walker 1979
    • A. paralectoris Harrison & Walker 1977
  • Coturnipes cooperi Harrison & Walker 1977 (Early Eocene of England, and Virginia, US?)– galliform, falconiform?
  • Fluviatilavis antunesi Harrison 1983 (Early Eocene of Silveirinha, Portugal)– charadriiform?
  • Mopsitta tanta Waterhouse et al. 2008 (Early Eocene) – threskiornithid (may belong in Rhynchaeites), psittacid?
  • Neanis schucherti Shufeldt 1913 (Early Eocene)– coraciiform (primobucconid), piciform?
  • Paleophasianus meleagroides Wetmore 1940 (Willwood Early Eocene of Bighorn County, US)– galliform (tetraonine or cracid) or gruiform (aramid)?
  • Precursor parvus (Early Eocene)– several species? psittaciform (pseudasturid or psittacid) + charadriiform (glareolid)?
  • "Precursor" litorum Harrison & Walker 1977
  • "Precursor" magnus Harrison & Walker 1977
  • Procuculus minutus Harrison & Walker 1977 (Early Eocene of Bognor Regis, England)– cuculiform (parvicuculid), coraciiform (primobucconid), close to Primapus?
  • Pulchrapollia Dyke & Cooper 2000 (Early Eocene) – psittaciform (pseudasturid or psittacid)?
    • P. gracilis Dyke & Cooper 2000
    • P. olsoni (Feduccia & Martin 1976) [Primobucco olsoni Feduccia & Martin 1976]
  • Neornithes incerta sedis USNM 496384 (Nanjemoy Early Eocene of Virginia, US)– parvicuculid? aegithalornithid?
  • Palaeopsittacus georgei Harrison 1982b (Early – middle Eocene of NW Europe) – caprimulgiform (podargid?) or quercypsittid?
  • Amitabha urbsinterdictensis Gulas-Wroblewski & Wroblewski 2003 (Bridger middle Eocene of Forbidden City, US) - galliform (phasianid) or gruiform (rallid?)?
  • Eociconia sangequanensis Hou 1989 (middle Eocene of China)– ciconiiform (ciconiid)?
  • Protocypselomorphus manfredkelleri Mayr 2005 (middle Eocene of Messel, Germany)– caprimulgiform, apodiform or ancestral to both
  • Pumiliornis tessellatus Mayr 1999 (middle Eocene of Messel, Germany)
  • Ludiortyx hoffmanni (Gervais 1852) Brunet 1970 (Late Eocene) – rallid, quercymegapodid? [Includes Tringa hoffmanni Gervais 1852; Palaeortyx blanchardi Milne-Edwards 1869; Palaeortyx hoffmanni (Gervais 1852); Ludiortyx blanchardi (Milne-Edwards 1869); Eortyx hoffmanni (Gervais 1852)]
  • Minggangia changgouensis Hou 1982 (Late Eocene of China) – rallid, threskiornithid?
  • Petropluvialis simplex Harrison & Walker 1976 (Late Eocene of England)– may be same as Palaeopapia; anseriform?
  • "Phasianus" alfhildae Shufeldt 1915 (Washakie B Late Eocene of Haystack Butte, US)– gruiform, ciconiiform, phoenicopteriform?
  • Telecrex grangeri Wetmore 1934 (Irdin Manha Late Eocene of Chimney Butte, Mongolia)– meleagrid or gruiform (rallid?)
  • Neornithes incerta sedis AMNH FR 2941 (Irdin Manha Late Eocene of Chimney Butte, China) – falconiform (accipitrid)? gruiform (Eogrus)?
  • Zheroia kurochkini Nesov 1988 (Late Eocene of Kazakhstan) – gruiform? pelagornithid?
  • "Falco" falconellus Shufeldt 1915 (or falconella; Eocene of Wyoming, US) – falconiform (falconid)?
  • Agnopterus Milne-Edwards 1868 (Late Eocene– Late Oligocene of Europe)– phoenicopteriform or anseriform
    • A. laurillardi Milne-Edwards 1868
    • A. sicki Alvarenga 1990
    • A. turgaiensis Tugarinov 1940 [Cygnopterus lambrechti Kuročkin 1968]
  • Plesiocathartes Gaillard 1908 (Late Eocene–? Early Miocene of SW Europe) - cathartid, leptosomid?
    • P. europaeus Gaillard 1908
    • P. gaillardi
    • P. wyomingensis Weidig 2006
    • P. major Weidig 2006
    • P. geiselensis Mayr 2002
    • P. kelleri Mayr 2002
  • Botauroides parvus Shufeldt 1915 (Eocene of Wyoming, US) – coliiform?
  • Aminornis excavatus Ameghino 1899 (Deseado Early Oligocene of Rio Deseado, Argentina) – gruiform (aramid)?
  • Ciconiopsis antarctica Ameghino 1899 (Deseado Early Oligocene of Patagonia, Argentina) – ciconiiform (ciconiid)?
  • Climacarthrus incompletus Ameghino 1899 (Deseado Early Oligocene of Argentina) – falconiform (accipitrid)? A nomen dubium
  • Cruschedula revola Ameghino 1899 (Deseado Early Oligocene of Golfo San Jorge, Argentina) Aves incertae sedis; A nomen dubium
  • Dolichopterus viator Milne-Edwards 1867 [Dolicopterus Aymard 1856 nomen nudum; Camaskelus Aymard 1856 nomen nudum; Camaskelus Milne-Edwards 1867; Dolicopterus viator Aymard 1856 nomen nudum; Camaskelus palustris Aymard 1856 nomen nudum; Camaskelus palustris Milne-Edwards 1867] (Early Oligocene of Ronzon, France) – charadriiform (charadriid)? Not Dolicopterus as sometimes claimed
  • Loncornis erectus Ameghino 1899 (Deseado Early Oligocene of Rio Deseado, Argentina)– gruiform (aramid)?
  • Loxornis clivus Ameghino 1894– anatid? (Deseado Early Oligocene of Argentina)
  • Manu antiquus Marples 1946 (Early Oligocene) – pelagornithid? procellariiform (diomedeid)?
  • Palaeocrex rex Wetmore 1927 (Early Oligocene of Trigonias Quarry, US) – gruiform (rallid)?
  • Palaeopapia eous >(Harrison & Walker 1976) Harrison & Walker 1979 [Howardia Harrison & Walker 1976 non Berlese & Leonardi 1896; Howardia eous Harrison & Walker 1976] (Hampstead Early Oligocene of Isle of Wight, England) – anseriform?
  • Paracygnopterus scotti Harrison & Walker 1979 (Early Oligocene of Belgium and England) – anseriform (anatid)?
  • "Pararallus" hassenkampi Martini 1967 (Sieblos Dysodil Early Oligocene of Sieblos, Germany)
  • Riacama caliginea Ameghino 1899 (Deseado Early Oligocene of Argentina) – gruiform?
  • Smiliornis penetrans Ameghino 1899 (Deseado Early Oligocene of Argentina)– gruiform?
  • Teracus littoralis Milne-Edwards 1871 [leracus Aymard 1856 nomen nudum; leracus littoralis Aymard 1856 nomen nudum] (Early Oligocene of France)
  • Teleornis impressus Ameghino 1899 (Deseado Early Oligocene of Argentina)– anatid?
  • Pseudolarus guaraniticus Ameghino 1899 (Deseado Early Oligocene – Miocene of Argentina) – gruiform?
  • Neornithes incerta sedis BMNH PAL 4989 (Hampstead Early Oligocene of Isle of Wight, England) – formerly "Ptenornis" and included in Headonornis; anseriform?
  • "Anas" creccoides van Beneden 1871 (Early-mid Oligocene of Belgium) – anseriform?
  • "Charadrius" sheppardianus Cope 1881 (Florissant middle Oligocene of Florissant, US) – charadriiform (charadriid?)
  • Megagallinula harundinea Kuročkin 1968 (Indricotherium middle Oligocene of Chelkar-Teniz, Kazakhstan)
  • "Palaeorallus" alienus Kuročkin 1968 (middle Oligocene of Tatal-Gol, Mongolia)– galliform?
  • "Vanellus" selysii van Beneden 1871 (middle Oligocene of Rupelmonde, Belgium) – charadriiform (charadriid)?
  • Anserpica kiliani Mourer-Chauviré, Berthet & Hugueney 2004 (Late Oligocene of France) – gruiform (gruid?) or anseriform (anseranatid?)?
  • Gnotornis Wetmore 1942 (Brule Late Oligocene of Shannon County, US) – gruiform (aramid)?
    • G. aramiellus Wetmore 1942
    • G. roardeola
    • G. walkeri
  • Guguschia nailiae Aslanova & Burčak-Abramovič 1968 (Late Oligocene of Pirəkəşkül, Azerbaijan) – anseriform (anserine)? pelagornithid (same as Caspiodontornis)?
  • Tiliornis senex Ameghino 1899 (Late? Oligocene of Argentina)– phoenicopteriform? A nomen dubium
  • Neornithes incerta sedis QM F40203 (Late Oligocene of Riversleigh, Australia)– gruiform (rallid)?
  • Gaviella pusilla Shufeldt 1915 (Oligocene? of Wyoming, US)– gaviiform? plotopterid?
  • "Anas" skalicensis Bayer 1882 (Early Miocene of "Skalitz", Czech Republic) - anseriform?
  • Chenornis graculoides Portis 1884 (Early Miocene) - Anseriformes (Anatidae) or Pelecaniformes (Phalacrocoracidae)?
  • "Propelargus" olseni Brodkorb 1963 (Hawthorne Early Miocene of Tallahassee, US) – ciconiiform?
  • Neornithes incerta sedis MNHN SA 1259-1263 (Early/Middle Miocene of Sansan, France) – passeriform?
  • Anisolornis excavatus Ameghino 1891 (Santa Cruz Middle Miocene of Karaihen, Argentina) – gruiform, galliform, tinamiform?
  • "Ardea" perplexa Milne-Edwards 1869 (Middle Miocene of Sansan, France) – ardeid? strigiform?
  • "Cygnus herrenthalsi" van Beneden 1871 (Middle Miocene of Belgium)
  • "Anas" risgoviensis Ammon 1918 (Late Miocene of Bavaria, Germany) – anseriform?
  • "Ardea" aureliensis (Late Miocene of France) – ardeid?
  • Eoneornis australis Ameghino 1895 nomen dubium (Miocene of Argentina) – anatid? A nomen dubium
  • Eutelornis patagonica Ameghino 1895 (Miocene of Argentina) – anatid?
  • Protibis cnemialis Ameghino 1891 (Miocene of Argentina) – ciconiiform (threskiornithid)?
  • "Limnatornis" paludicola Milne-Edwards 1871 (Miocene of France) – coliid? phoeniculid?
  • "Picus" gaudryi (Miocene of France) – piciform?
  • "Ardea" lignitum Giebel 1860 (Late Pliocene of Germany) – ardeid? strigid (genus Bubo)?
  • Bathoceleus hyphalus Brodkorb 1959 (Pliocene of New Providence, Bahamas) – picid?
  • "Homalopus" Milne-Edwards 1870 non Chevrolat 1837– piciform? Preoccupied by a subgenus of Cryptocephalus leaf beetles described in 1835.
  • "Liptornis hesternus" Ameghino 1894 [Liptornis cuvierii] – pelecaniform (pelecanid)? A nomen dubium
  • Proceriavis martini Harrison & Walker 1979 – pelagornithid?
  • Protopelicanus cuvieri Reichenbach 1852 – pelecaniform (pelecanid)? pelagornithid?
  • Eurofluvioviridavis robustipes Mayr 2005a (middle Eocene of Messel, Germany)
  • Archaeotrogonidae Wetmore 1926 – basal Cypselomorphae?
    • †Archaeotrogonidae gen. et sp. indet. (Early Eocene)
    • Hassiavis laticauda Mayr 1998
    • Archaeotrogon Milne-Edwards 1892 (Late Eocene/Early Oligocene)
  • Cimolopterygidae Brodkorb 1963 – charadriiform?
    • Cimolopteryx Marsh 1892 (Late Cretaceous)
    • Lamarqueavis Agnolin 2010 (Late Cretaceous)
      • L. australis Agnolin 2010
      • L. minima (Brodkorb 1963) Agnolin 2010 [Cimolopteryx minima Brodkorb 1963]
      • L. petra (Hope 2002) Agnolin 2010 [Cimolopteryx petra Hope 2002]
  • Cladornithidae Wetmore 1930 [Cladornithes Wetmore 1960; Cladornidae Ameghino 1895] (pelecaniform?
  • Eremopezidae Rothschild 1911 – pelecaniform? ratite?
    • Eremopezus eocaenus Andrews 1904 (Late Eocene) - includes Stromeria fajumensis Lambrecht 1929
  • Gracilitarsidae Mayr 2001 – close to Sylphornithidae?
  • Halcyornithidae Harrison & Walker 1972 – psittaciform (= Pseudasturidae?), coraciiform?
    • Halcyornis toliapicus (Koenig 1825) Owen 1846 (London Clay Early Eocene of England)
    • Cyrilavis Martin 2010 (Green River Early Eocene of C US) - psittaciform (halcyornithid or psittacid)?
      • C. olsoni (Feduccia & Martin 1976) Martin 2010 [Primobucco olsoni Feduccia & Martin 1976]
      • C. colburnorum Ksepka, Clarke & Grande 2011
  • Juncitarsidae Peters 1987
    • Kashinia magnum (Harrison & Walker 1976) Harrison & Walker 1979 [Tenuicrus Harrison & Walker 1976 non Womersley 1940; Tenuicrus magnum Harrison & Walker 1976] – phoenicopteriform?
    • Juncitarsus Olson & Feduccia 1980 – phoenicopteriform?
  • Laornithidae Cracraft 1973 – charadriiform? gruiform? pelagornithid?
  • Lonchodytidae Brodkorb 1963
    • Lonchodytes estesi Brodkorb 1963 (Late Cretaceous/?Early Palaeocene) – gaviiform/pelecaniform? procellariiform?
  • Palaeospizidae – passeriform? coraciiform? coliiform?
  • Parvicuculidae Harrison 1982 – cypselomorph, cuculiform, coraciiform (primobucconid)?
    • Parvicuculus minor Harrison & Walker 1977 (Early Eocene of NW Europe)
  • Remiornithidae Martin 1992 (Paleocene of France) – palaeognath?
  • Sylphornithidae Mourer-Chauviré 1988 – cuculiform? coraciiform? close to Gracilitarsidae?
    • Sylphornis bretouensis Mourer-Chauvire 1988 (middle Eocene of France)
    • Oligosylphe mourerchauvireae Mayr & Smith 2002 (Borgloon Early Oligocene of Hoogbutsel, Belgium)
  • Tytthostonychidae Olson & Parris 1987 – procellariiform, pelecaniform?
  • Zygodactylidae Brodkorb 1971 [Primoscenidae Harrison 1977]– near passerine
    • Eozygodactylus americanus Weidig 2010
    • Zygodactylus Ballmann 1969 (Early Oligocene – Middle? Miocene of C Europe)
      • Z. ignotus Ballmann 1969
      • Z. luberonensis Mayr 2008
      • Z. grivensis Ballmann 1969
    • Primoscens minutus Harrison & Walker 1977
    • Primozygodactylus Mayr 1998
      • P. ballmanni Mayr 1998
      • P. danielsi Mayr 1998
      • P. major Mayr 1998
      • P. eunjooae Mayr & Zelenkov 2009
  • "Graculavidae" Fürbringer 1888 [Dakotornithidae Erickson 1975; Scaniornithidae; Telmatornithidae Cracraft 1972] – a paraphyletic form taxon, the "transitional shorebirds"
    • Limosavis Shufeldt 1915 [Graculavus Marsh 1872 non Marsh 1973] (Late Cretaceous –? Early Palaeocene) – charadriiform?
      • L. velox (Marsh 1872) Lambrecht 1933 [Graculavus velox Marsh 1872]
      • L. augustus (Hope 1999) [Graculavus augustus Hope 1999]
    • Palaeotringa Marsh 1870 (Late Cretaceous/Early Palaeocene) – charadriiform?
    • Telmatornis priscus Marsh 1870 [Telmatornis affinis Marsh 1870; Graculavus pumilis Marsh 1872; Palaeotringa vetus Marsh 1870] (Late Cretaceous?) – charadriiform? gruiform? podicipediform?
    • Zhylgaia aestiflua Nesov 1988 (Early Paleocene) – presbyornithid?
    • Scaniornis lundgreni Dames 1890 (Early/Middle Paleocene) – phoenicopteriform?
    • Dakotornis cooperi Erickson 1975 (Paleocene of North Dakota, US)
    • Placement unresolved
      • "Graculavidae" gen. et sp. indet. (Gloucester County, US)

Ostrich and related ratites.

  • Struthionidae Vigors 1825 [†Struthiolithidae Vjalov 1971; †Palaeotididae Houde & Haubold 1987] (ostriches)
    • Palaeotis weigelti Lambrecht 1928 [Palaeogrus geiseltalensis Lambrecht 1935; Ornitocnemus geiseltalensis (Lambrecht 1935) Brodkorb 1967] (middle Eocene)
    • Remiornis heberti Lemoine 1881
    • Orientornis linxiaensis (Hou et al. 2005) Wang 2008 [Struthio linxiaensis Hou et al. 2005]
    • Prehistoric species of extant genera
      • Struthio Linnaeus 1758 [†Palaeostruthio Burchak-Abramovich 1953; †Struthiolithus Brandt 1873 ootaxa; Struthio (Pachystruthio) Kretzoi 1954a; Megaloscelornis Lydekker 1879] (Early Miocene – Recent)

Cassowaries, emus and related ratites.

  • Casuariidae Kaup 1847 [Dromaiidae Huxley 1868; Dramaiinae Gray 1870; Dramiceiidae Richmond 1908; Dramaeidae Newton 1896] (emus and cassowaries)
    • Diogenornis fragilis Alvarenga 1983 (Early Eocene) – possible casuariiform
    • Emuarius Boles 1992 (emuwaries) (Late Oligocene – Late Miocene)
    • Prehistoric species of extant genera
      • Dromaius Vieillot 1816 [Tachea Fleming 1822; Emou Griffith & Pidgeon 1829; Peronista Mathews 1912; Metapteryx De Vis 1892] (emus) (Middle Miocene – Recent)
      • Casuarius Brisson 1760 [Cela Moehring 1752 [Oken1816] non Reichenbach 1853 non Illiger 1826; Cela Moehring 1752 nom. rej.; Rhea Lacépède 1800 non Latham 1790; Chelarga Billberg 1828; Oxyporus Brookes 1828 non (Bourdot & Galzin 1925) Donk 1933; Thrasys Billberg 1828; Cassowara Perry 1811; Hippalectryo Gloger 1842]

Rheas and related ratites.

  • †Opisthodactylidae Ameghino 1895
  • Rheidae (Bonaparte 1853) [Rheinae Bonaparte 1853] (rheas)
    • Heterorhea dabbenei Rovereto 1914 (Pliocene)
    • Hinasuri nehuensis Tambussi 1995
    • Prehistoric species of extant genera
      • Rhea Brisson 1760 [Rhea Moehring 1758 nomen dubium; Pterocnemia Gray 1870; Toujou Lacépède 1801; Tujus Rafinesque 1815]

Moas.

  • Megalapterygidae - (Upland moas)
    • Megalapteryx didinus (Owen 1883) Haast 1886 [Palaeocasuarius Rothschild 1907 (Forbes 1892); Dinornis didinus Owen 1883; Megalapteryx benhami Archey 1941; Megalapteryx hamiltoni Rothschild 1907; Megalapteryx hectori Haast 1886; Megalapteryx tenuipes Lydekker 1891; Anomalopteryx didina (Owen 1883) Lydekker 1891; Palaeocasuarius velox Rothschild 1907; Palaeocasuarius elegans Rothschild 1907; Palaeocasuarius haasti Rothschild 1907]
  • Dinornithidae Owen 1843 [Palapteryginae Bonaparte 1854; Palapterygidae Haast 1874; Dinornithnideae Stejneger 1884] (great moas)
    • Dinornis Owen 1843 [Palapteryx Owen 1846; Megalornis Owen 1843 non Gray 1841; Movia Reichenbach 1853; Moa Reichenbach 1853; Owenia Gray 1855; Tylopteryx Hutton 1891]
  • Emeidae (Bonaparte 1854) [Emeinae Bonaparte 1854; Anomalopterygidae Oliver 1930; Anomalapteryginae Archey 1941] (lesser moas)
    • Anomalopteryx didiformis (Owen 1844) Reichenbach 1853 (Little bush moas)
    • Emeus crassus (Owen 1846) Reichenbach 1853 (Eastern moas)
    • Euryapteryx Haast 1874 (Broad-billed or turkey moas)
    • Pachyornis Lydekker 1891 (Stout moas)

Two unnamed Saint Bathans Fauna species.[17]

†Lithornithiformes Houde 1988

  • Tinamidae Gray 1840 [Crypturidae Bonaparte 1831; Tinamotidae Bonaparte 1854; Eudromiidae Bonaparte 1854; Rhynchotidae von Boetticher 1934] (tinamous)
    • Querandiornis romani Rusconi 1958 (Ensenada Early/Middle Pleistocene of Argentina)
    • Roveretornis Brodkorb 1961
    • Tinamidae gen. et sp. indet. MACN-SC Fleagle Collection (Early – Middle Miocene of S Argentina) - at least 2 species
    • Prehistoric species of extant genera
      • Eudromia [non Eudromias Boie 1832; Calodromas Sclater & Salvin 1873 non Calodromus Guein 1832; Calopezus Ridgway 1884; Tinamisornis Rovereto 1914; Roveretornis Brodkorb 1961]
      • Nothura Wagler 1827 [†Cayetanornis Brodkorb 1963]
      • Crypturellus Brabourne & Chubb 1914 [Microcrypturus Chubb 1917; Crypturornis Oberholser 1922; Orthocrypturus Miranda-Ribeiro 1937]

Vegaviiformes

Dromornis
Genyornis

The group that includes modern ducks and geese.

  • Basal and unresolved forms
    • Anatalavis Olson & Parris 1987 [Nettapterornis Mlikovsky 2002] (Late Cretaceous/Early Paleocene– Early Eocene)– anseranatid or basal.
      • A. rex (Shufeldt 1915) Olson & Parris 1987 [Telmatornis rex Shufeldt 1915]
      • A. oxfordi Olson 1999 [Nettapterornis oxfordi (Olson 1999) Mlikovskmiy 2002]
    • Romainvillia stehlini Lebedinský 1927 (Late Eocene/Early Oligocene)– anseranatide or anatide
    • Saintandrea chenoides Mayr & De Pietri 2013
    • Proherodius oweni Lydekker 1891 (Early Eocene) – presbyornithid?
    • Paranyroca magna Miller & Compton 1939 (Early Miocene) – anatide or own family?
    • Garganornis ballmanni Meijer 2014
  • Anhimidae Stejneger 1885 [Palamedeinae Bonaparte 1831; Palamedeidae Gray 1840] (screamers
  • Dromornithidae Fürbringer 1888 – Australian mihirungs or "demon ducks"
  • Anseranatidae Sclater 1880 sensu Stejneger 1885 (magpie-geese
    • Eoanseranas handae Worthy & Scanlon 2009 (Hand's Dawn Magpie Goose)
    • Anseranatidae gen. et sp. nov (Late Oligocene)
  • Presbyornithidae Wetmore 1926 [Telmabatidae Howard 1955]
    • Teviornis gobiensis Kuročkin, Dyke & Karhu 2002 (Late Cretaceous)
    • Headonornis hantoniensis (Lydekker 1891) Harrison & Walker 1976 [Agnopterus hantoniensis Lydekker 1891; Presbyornis hantoniensis (Lydekker 1891) Bocheński 1997; Ptenornis] – only BMNH PAL 30325 belongs to this bird, may belong to Presbyornis.
    • Presbyornis Wetmore 1926 [Nautilornis Wetmore 1926; Coltonia Hardy 1959] (Paleocene– Early Oligocene)
    • Telmabates Howard 1955
      • T. antiquus Howard 1955
      • T. howardae Cracraft 1970
    • Wilaru Boles et al. 2013
  • Anatidae Leach 1820 [Cnemiornithidae Stejneger 1855; Cygnidae Kaup 1850 fide Gray; Anseridae Kaup 1850 fide Gray; Cereopsidae Kaup 1850 fide Gray; Fuligulidae Carus 1868; Merganidae Swainson 1837; Mergidae Gray 1840 fide 1871; Plectropteridae Carus; Erismaturidae Carus 1868; Tadornidae Carus 1868] (ducks, geese and swans)
    • Allgoviachen tortonica Mayr, Lechner & Böhme, 2022 (Late Miocene)
    • Annakacygna Matsuoka & Hasegawa 2022 (Late Miocene)
    • Bonibernicla ponderosa Kuročkin 1985 [Branta woolfendeni Bickart 1990]
    • Eonessa anaticula Wetmore 1938 (Middle Eocene)
    • Cygnavus (Early Oligocene – Early Miocene)
      • C. senckenbergi Lambrecht 1931
      • C. formosus Korochkin 1968
    • Cygnopterus Lambrecht 1931 (middle Oligocene – Early Miocene)– sometimes included in Cygnavus
      • C. neogradiensis Kessler & Hir 2009
      • C. affinis (Beneden 1883) Lambrecht 1931 [Sula affinis Beneden 1883; Palaeopapia hamsteadiensis Harrison & Walker 1979b]
    • Mionetta Livezey & Martin 1988 (Late Oligocene – Early Miocene)
      • M. robusta (Milne-Edwards 1868) Mlíkovský 2002 [Anas robusta Milne-Edwards, 1868; Anserobranta robusta (Milne-Edwards 1868) Cheneval 1987]
      • M. arvernensis (Lydekker 1891a) [Aythya arvernensis (Lydekker 1891a) Brodkorb 1964; Fuligula arvernensis Lydekker, 1891a]
      • M. blanchardi (Milne-Edwards 1863) Livezey & Martin 1988 [Anas blanchardi Milne-Edwards 1863; Palaeortyx phasianoides Milne-Edwards 1869; Anas macroptera Milne-Edwards 1871 Nomen nudum; Taoperdix phasianoides (Milne-Edwards 1869) Brodkorb 1964; Dendrochen blanchardi (Milne-Edwards 1863) Cheneval 1983b]
      • M. consobrina (Milne-Edwards 1867) Livezey & Martin 1988 [Anas consobrina Milne-Edwards 1867; Dendrochen consobrina (Milne-Edwards 1867) Cheneval 1983b]
      • M. natator (Milne-Edwards 1867) Livezey & Martin 1988 [Anas natator Milne-Edwards 1867; Querquedula natator (Milne-Edwards 1867) Brodkorb 1962; Dendrochen natator (Milne-Edwards 1867) Cheneval 1983b]
    • Notochen bannockburnensis Worthy et al., 2022 (Early Miocene)
    • Australotadorna alecwilsoni Worthy 2009 (Late Oligocene – Early Miocene)
    • Pinpanetta Worthy 2009 (Late Oligocene – Early Miocene)
    • Dunstanetta johnstoneorum Worthy et al. 2007 (Johnstone's ducks) (Early/Middle Miocene)
    • Manuherikia Worthy et al. 2007 (Early/Middle Miocene)
      • M. lacustrina Worthy et al. 2007 (Manuherikia duck)
      • M. minuta Worthy et al. 2007 (Minute Manuherikia duck)
      • M. douglasi Worthy et al. 2008 (Douglas' duck)
    • Matanas enrighti Worthy et al. 2007 (Enright's ducks) (Early/Middle Miocene)
    • Miotadorna Worthy et al. 2007 (Early/Middle Miocene)
      • M. sanctibathansi Worthy et al. 2007 (St. Bathans shelducks) (Early/Middle Miocene)
      • M. catrionae Tennyson et al. 2022 (Catriona's shelduck) (late Early/early Middle Miocene) (synonym of M. sanctibathansi)
    • Megalodytes morejohni Howard 1992 (Middle Miocene)
    • Sinanas diatomas Yeh 1980 (Middle Miocene)
    • Dendrochen Miller 1944 (Late Miocene)
      • D. integra (Miller 1944) Cheneval 1987 [Querquedula integra Miller 1944; Anas integra (Miller 1944); Spatula integra (Miller 1944)]
      • D. oligocaena (Tugarinov 1940) [Anas oligocaena Tugarinov 1940]
      • D. robusta Miller 1944
    • Presbychen abavus Wetmore 1930 (Late Miocene)
    • Afrocygnus chauvireae Louchart et al. 2005 (Late Miocene – Early Pliocene)
    • Balcanas pliocaenica (Early Pliocene) – may belong in Tadorna
    • Wasonaka yepomerae Howard 1966 (Middle Pliocene)
    • Paracygnus plattensis Short 1969 (Late Pliocene)
    • Anabernicula Ross 1935 (Late Pliocene? – Late Pleistocene)
    • Eremochen russelli Brodkorb 1961 (Pliocene)
    • Tirarinetta kanunka Worthy 2008 (Pliocene)
    • Brantadorna Howard 1964 (Middle Pleistocene)
      • B. downsi Howard 1964
      • B. robusta (Short 1970) Livezey 1997 [Anabernicula robusta Short 1970]
    • Nannonetta invisitata Campbell 1979 (Late Pleistocene)
    • Aldabranas cabri Harrison & Walker 1978 (Late Pleistocene)
    • Ankonetta larriestrai Cenizo & Agnolín 2010
    • Cayaoa bruneti Tonni 1979
    • Chenoanas Zelenkov 2012
    • Helonetta brodkorbi Emslie 1992
    • Mioquerquedula minutissima Zelenkov & Kuročkin 2012 [Anas velox Milne-Edwards 1867; Anas meyerii Milne-Edwards, 1868; Nettion velox (Milne-Edwards 1868) Brodkorb 1964; Aythya meyerii (Milne-Edwards 1868) Brodkorb 1964] (Middle – Late? Miocene)
    • Proanser major Umanskaya 1979
    • Shiriyanetta hasegawai Watanabe & Matsuoka 2015
    • Pleistoanser bravardi Agnolín 2006
    • Lavadytis pyrenae Stidham & Hilton 2015
    • Asiavis phosphatica Nesov 1986
    • Nogusunna conflictoides Zelenkov 2011
    • Sharganetta mongolica Zelenkov 2011
    • Protomelanitta Zelenkov 2011
      • P. gracilis Zelenkov 2011
      • P. shihuibas (Hou 1985) Zelenkov 2012 [Aythya shihuibas Hou 1985]
    • Bambolinetta lignitifila (Portis 1884) Mayr & Pavia 2014 [Anas lignitifila Portis 1884] (Early Pliocene of Monte Bamboli, Italy)
    • Heteroanser vicinus (Kuročkin 1976) Zelenkov 2012 [Heterochen vicinus Kuročkin 1976; Anser vicinus (Kuročkin 1976) Mlíkovský & Švec 1986]
    • Placement unresolved
      • "Anas" albae Jánossy 1979 (Late Miocene) – formerly in Mergus
      • "Anas" amotape Campbell 1979
      • "Anas" eppelsheimensis Lambrecht 1933 (Early Pliocene)
      • "Anas" isarensis Lambrecht 1933 (Late Miocene)
      • "Anas" luederitzensis (Early Miocene)
      • "Anas" sanctaehelenae Campbell 1979
      • "Anser" scaldii [Anser scaldii Beneden 1872 nomen nudum; "Anas" scaldii] (Late Miocene)
      • "Aythya" chauvirae (Middle Miocene) – 2 species
      • "Chenopis" nanus De Vis 1905 (Pleistocene) – at least 2 taxa, may be living species
      • "Cygnopterus" alphonsi Cheneval 1984
      • "Oxyura" doksana Mlíkovský 2002 (Early Miocene)
      • †"cf. Megalodytes" (Middle Miocene)
      • †Anatidae gen. et sp. indet. MNZ S42797 (Early/Middle Miocene)
      • †Anatidae gen. et spp. indet. (Middle Miocene)
      • †Anatidae gen. et spp. indet. (Late Miocene)
    • Prehistoric species of extant genera
      • Biziura Stephens 1824
        • B. exhumata DeVis 1889
        • B. delautouri Forbes 1892 [Biziura lobata delautouri (Forbes 1892)] (New Zealand musk duck)
      • Dendrocygna
        • D. eversa Wetmore 1924
        • D. valdipinnis DeVis 1888
        • D. soporata (Kuročkin 1976) Mlíkovský & Švec 1986 [Anas soporata Kuročkin 1976]
      • Somateria (middle Oligocene – Recent)
      • Bucephala
      • Clangula
      • Aix
      • Cygnus
        • C. mariae Bickart 1990
        • C. csakvarensis (Lambrecht 1933) Mlíkovský 1992 [Cygnus csákvárensis Lambrecht 1931a nomen nudum; Cygnanser csakvarensis (Lambrecht 1933) Kretzoi 1957; Olor csakvarensis (Lambrecht 1933) Mlíkovský 1992b]
        • C. liskunae (Kuročkin 1976) [Anser liskunae Kuročkin 1976]
        • C. atavus (Fraas 1870) Mlíkovský 1992 [Anas atava Fraas 1870; Anas cygniformis Fraas 1870; Palaelodus steinheimensis Fraas 1870; Anser atavus (Fraas 1870) Lambrecht 1933; Anser cygniformis (Fraas 1870) Lambrecht 1933]
        • C. paloregonus Cope 1878 [Anser condoni Schufeldt 1892; Cygnus matthewi Schufeldt 1913]
        • C. hibbardi Brodkorb 1958
        • C. lacustris (De Vis 1905) [Archaeocygnus lacustris De Vis 1905]
        • C. verae Boev 2000
      • Chloephaga
      • Histrionicus
      • Mergus
      • Anas
        • A. elapsa DeVis 1888 [Nettion elapsum (DeVis 1888)] (Chinchilla Late Pleistocene of Condamine River, Australia) ("Nettion")
        • A. gracilipes DeVis 1905 [Nettion gracilipes (DeVis 1905)] (Late Pleistocene of Australia) ("Nettion")
        • A. moldovica (Late Pliocene of Tchichmiknaia, Moldova?Georgia?)
        • A. strenuum (Late Pleistocene of Patteramordu, Australia) ("Nettion")
        • A. kisatibiensis [Anser kisatibiensis] (Early Pliocene of Kisatibi, Georgia)
        • A. lambrechti (Spillman 1942) [Archeoquerquedula lambrechti Spillman 1942; Archaeoquerquedula (sic) lambrechti Stephens; Querquedula lambrechti (Spillman 1942)](Late? Pleistocene of Santa Elena peninsula, Ecuador)
        • A. itchtucknee McCoy 1963
        • A. kurochkini Zelenkov & Panteleyev 2015
        • A. pullulans (Brodkorb 1961)
        • A. schneideri Emslie 1985
        • A. bunkeri (Wetmore 1944) [Nettion bunkeri Wetmore 1944]
        • A. greeni (Brodkorb 1964) [Nettion greeni Brodkorb 1964]
        • A. ogallalae (Brodkorb 1962) [Nettion ogallalae Brodkorb 1962]
        • A. cheuen Agnolín 2006
      • Anser
        • A. arenosus Bickart 1990
        • A. arizonae Bickart 1990
        • A. azerbaidzhanicus Serebrovsky 1940
        • A. devjatkini Kuročkin 1971
        • A. djuktaiensis Zelenkov & Kuročkin 2014
        • A. eldaricus Burchak-Abramovich & Gadzyev 1978
        • A. oeningensis (Meyer 1865) Milne-Edwards 1867b [Anas oeningensis Meyer 1865]
        • A. pratensis (Short 1970) [Heterochen pratensis Short 1970]
        • A. tchikoicus Kuročkin 1985
        • A. thompsoni Martin & Mengel 1980
        • A. thraceiensis Burchak-Abramovich & Nikolov 1984
        • A. pressa (Brodkorb 1964) [Chen pressa Brodkorb 1964] (Dwarf Snow goose)
        • A. udabnensis Burchak-Abramovich 1957
      • Aythya
      • Branta
      • Oxyura
      • Chendytes
        • O. milleri Howard 1955 (Early Pleistocene of San Nicolás Island US)
      • Lophodytes (Late Pleistocene – Recent)
      • Neochen
      • Tadorna

The group that includes domestic chickens and their relatives.

  • Placement unresolved
    • Archaealectrornis sibleyi Crowe & Short 1992 (Oligocene) - phasianid?
    • Archaeophasianus Lambrecht 1933 (Oligocene? - Late Miocene) - tetraonid or phasianid
      • A. mioceanus (Shufeldt 1915) [Phasianus mioceanus Shufeldt 1915]
      • A. roberti (Stone 1915) [Phasianus roberti Stone 1915]
    • Austinornis lentus (Marsh 1877b) Clarke 2004 [Graculavus lentus Marsh 1877b; Ichthyornis lentus (Marsh 1877b) Marsh 1880; Pedioecetes lentus (Marsh 1877b); Pedioecetes phasianellus (Linnaeus 1758) Shufeldt 1915; Tetrao phasianellus Linnaeus 1758] - (Late Cretaceous) tentatively placed here;
    • Chambiortyx cristata Mourer-Chauviré et al. 2013
    • "Cyrtonyx" tedfordi (Miller 1952) (Late Miocene)
    • Linquornis gigantis Yeh 1980 (middle Miocene)
    • Namaortyx sperrgebietensis Mourer-Chauviré, Pickford & 2011
    • Palaealectoris incertus Wetmore 1930 (Early Miocene) - tetraonid?
    • Palaeoalectoris songlinensis Hou 1987 (middle Miocene)
    • Palaeonossax senectus Wetmore 1956 (Late Oligocene) - cracid?
    • Palaeortyx Milne-Edwards 1869 [Palaeoperdix Milne-Edwards 1869; Proalector Brodkorb 1964] (middle Eocene -? Early Pliocene) - phasianid or odontophorid
      • P. media Milne-Edwards 1871 (nomen nudum)
      • P. volans Göhlich & Pavia 2008
      • P. gallica Milne-Edwards 1869 non Lydekker 1891 [Rallus dasypus Milne-Edwards 1892; Quercyrallus dasypus (Milne-Edwards) Lambrecht 1933; Taoperdix gallica (Milne-Edwards) Brodkorb 1964; ?Palaeortyx intermedia Ballmann 1969b; Coturnix gallica (Milne-Edwards) Mlýkovský 2002a]
      • P. brevipes Milne-Edwards 1869 emend. Paris 1912 [Palaeortyx ocyptera Milne-Edwards 1892; Palaeortyx grivensis Lydekker 1893 non Ballmann 1969a; Plioperdix grivensis (Lydekker 1893) Brodkorb 1964; Palaeortyx cayluxensis Lydekker 1891 non Milne-Edwards 1892 emend. Gaillard 1908; Taoperdix brevipes (Milne-Edwards 1869) Brodkorb 1964]
      • P. prisca (Milne-Edwards 1869) [Alectoris prisca (Milne-Edwards 1869) Mlýkovský 2002a; Palaeoperdix prisca Milne-Edwards 1869 emend. Göhlich & Mourer-Chauviré 2005; Palaeoperdix sansaniensis Milne-Edwards 1869; Palaeocryptonyx grivensis Ennouchi 1930]
      • P. phasianoides Milne-Edwards 1869 [Palaeoperdix longipes Milne-Edwards 1869; Palaeortyx longipes (Milne-Edwards 1869) Mlíkovský 2000e; Coturnix longipes (Milne-Edwards 1869) Mlíkovský 2002; Palaeocryptonyx gaillardia Ennouchi 1930; Proalector gaillardia (Ennouchi 1930) Brodkorb 1964; Palaeortyx phasianoides grivensis Ballmann 1969a non Lydekker 1893b; Taoperdix phasianoides (Milne-Edwards 1869) Brodkorb 1964]
      • P. joleaudi Ennouchi 1930 [Plioperdix joleaudi (Ennouchi 1930) Brodkorb 1964; Palaeortyx depereti Ennouchi 1930; Plioperdix depereti (Ennouchi 1930) Brodkorb 1964]
    • Procrax brevipes Tordoff & Macdonald 1957 (middle Eocene? - Early Oligocene) - cracid? gallinuloidid?
    • Shandongornis Yeh 1997 (middle Miocene)
    • Sobniogallus albinojamrozi Tomek et al. 2014
    • Taoperdix Milne-Edwards 1869 (Late Oligocene) - gallinuloidid?
      • T. pesseiti (Gervais 1862) [Tetrao pessieti Gervais 1862]
      • T. miocaena (Gaillard 1939) Ballmann 1969 [Palaeortyx miocaena Gaillard 1939]
    • Galliformes gen. et sp. indet. MCZ 342506 (Oligocene) - formerly in Gallinuloides; phasianid?
  • Gallinuloididae Lucas 1900
    • Gallinuloides wyomingensis Eastman 1900 (Early/middle Eocene)
    • Paraortygoides Mayr 2000 (London Clay Early Eocene of Walton-on-the-Naze, England – middle Eocene of Messel, Germany)
      • P. messelensis Mayr 2000
      • P. radagasti Dyke & Gulas 2002
  • Paraortygidae Mourer-Chauviré 1992
    • Pirortyx major (Gaillard 1939) Brodkorb 1964 [Palaeortyx major Gaillard 1939]
    • Scopelortyx klinghardtensis Mourer-Chauviré, Pickford & Senut 2015
    • Paraortyx Gaillard 1908 sensu Brodkorb 1964
      • P. lorteti Gaillard 1908 [Palaeortyx cayluxensis Lydekker 1891; Palaeortyx gaillardi Lambrecht 1933; Ludiortyx gaillardi (Lambrecht 1933) Brodkorb 1964]
      • P. brancoi Gaillard 1908
  • Quercymegapodiidae Mourer-Chauviré 1992
    • Taubacrex granivora dAlvarenga 1988 (Late Oligocene/Early Miocene of Brazil)
    • Ameripodius Alvarenga 1995 (Late Oligocene – Early Miocene of Brazil and France)
      • A. silvasantosi Alvarenga 1995
      • A. alexis Mourer-Chauviré 2000
    • Quercymegapodius Mourer-Chauviré 1992 (middle Eocene – Early Oligocene)
      • Q. depereti (Gaillard 1908) [Palaeocryptonyx depereti Gaillard 1908]
      • Q. brodkorbi Mourer-Chauviré 1992
  • Megapodidae – megapodes
    • Ngawupodius minya Boles & Ivison 1999
    • Prehistoric species of extant genera
      • Leipoa gallinacea – formerly Chosornis, Palaeopelargus, Progura
  • Cracidae – guans and curassows
    • Boreortalis laesslei Brodkorb 1954 (Early Miocene) – may be same as Ortalis
    • Extant genera present in the fossil record
  • Odontophoridae – New World quails
    • Nanortyx inexpectatus Weigel 1963 (Cypress Hills Early Oligocene of North Calf Creek, Canada)
    • Neortyx peninsularis Holman 1961 (Early Pleistocene of Reddick, US)
    • Miortyx Miller 1944 (Rosebud Early Miocene of Flint Hill, US)
      • M. teres Miller 1944
      • M. aldeni Howard 1966
    • Placement unresolved
      • Odontophoridae gen. et sp. indet. KUVP 9393 (White River Early/middle Oligocene of Logan County, US)
    • Prehistoric species of extant genera
      • Cyrtonyx cooki Gutierrez et al. 1981 (Late Miocene? of Upper Sheep Creek, US)
      • Callipepla shotwelli (Brodkorb 1958) [Lophortyx shotwelli Brodkorb 1958] (Middle Pliocene of McKay Reservoir, US)
      • Colinus hibbardi Wetmore 1944 (Rexroad Late Pliocene of Rexroad, US)
      • Colinus suilium Brodkorb 1959 (Early Pleistocene of SE US)
      • Colinus sp. (Late Pliocene of Benson, US)
      • Dendrortyx? sp. (Late Pleistocene of San Josecito Cavern, Mexico)
  • Phasianidae – pheasants, quails, partridges, grouse and turkeys
    • Bantamyx georgicus Kuročkin 1982
    • Centuriavis lioae Ksepka et al., 2022
    • Diangallus mious Hou 1985
    • Lophogallus naranbulakensis Zelenkov & Kuročkin 2010
    • Megalocoturnix cordoni Sánchez Marco 2009 (Early Pliocene of Layna, Spain)
    • Miophasianus Brodkorb 1952 [Miophasianus Lambrecht 1933 nomen nudum; Miogallus Lambrecht 1933]
      • M. maxima (Lydekker 1893) Brodkorb 1964 [Palaeortyx maxima Lydekker 1893]
      • M. altus (Milne-Edwards 1869) Villalta & Crusafont 1950 [Phasianus altus Milne-Edwards 1869; Miogallus altus (Milne-Edwards 1869) Mlíkovský 2002; Phasianus Desnoyersii Milne-Edwards 1869; Ardea similis Fraas 1870; Tantalus milne-edwardsii Shufeldt 1896; Pseudotantalus Milne-Edwardsi (Shufeldt 1896) Sharpe; Gallus longaevus Ammon 1918; Phasianus augustus Ammon 1918; Botaurites similis (Fraas 1870) Lambrecht 1933; Miophasianus augustus (Ammon 1918) Lambrecht 1933; Miophasianus desnoyersii (Milne-Edwards 1869) Lambrecht 1933; Miogallus longaevus (Ammon 1918) Lambrecht 1933; Proardea similis (Ammon 1870) Gaillard 1939; Ibis milne-edwardsi (Shufeldt 1896) Brodkorb 1963c]
      • M. medius (Milne-Edwards 1869) Lambrecht 1933 [Phasianus medius Milne-Edwards 1869; Palaeoperdix medius (Milne-Edwards 1869) Cheneval 2000]
    • Proagriocharis kimballensis Martin & Tate 1970 (Kimball Late Miocene/Early Pliocene of Lime Creek, US)
    • Palaeocryptonyx Depéret 1892 [Chauvireria Boev 1997; Pliogallus Tugarinov 1940b non Gaillard 1939; Lambrechtia Janossy 1974] (Late Pliocene of SW Europe)
      • P. donnezani Depéret 1892 [Francolinus Čapeki Lambrecht 1933; Pliogallus coturnoides Tugarinov 1940b; Plioperdix coturnoides (Tugarinov 1940b) Kretzoi 1955b; Francolinus capeki wenzensis Jánossy 1974a; Francolinus capeki villányiensis Jánossy 1974a; Francolinus (Lambrechtia) capeki (Lambrecht 1933) Jánossy 1974a; Alectoris baryosefi Tchernov 1980; Francolinus wezensis (Jánossy 1974a) Jánossy 1981; Plioperdix capeki (Lambrecht 1933) Mlíkovský 1995b; Alectoris donnezani (Depéret 1892) Mlíkovský 2002; Chauvireria balcanica Boev 1997a]
      • P. grivensis Ennouchi 1930
      • P. novaki Sánchez Marco 2009
      • P. edwardsi (Depéret 1887) Ballmann 1969a [Palaeortyx edwardsi Depéret 1887; Palaeortyx miocaena Gaillard 1939; Taoperdix miocaena (Gaillard 1939) Ballmann 1969; Palaeoperdix edwardsi (Depéret 1887) Brodkorb 1964; Alectoris edwardsi (Depéret 1887) Mlíkovský 2002]
      • P. minor (Jánossy 1974) [Francolinus minor Jánossy 1974; Ammoperdix ponticus Tugarinov 1940b; Pliogallus ponticus (Tugarinov 1940); Plioperdix ponticus (Tugarinov 1940) Kuročkin 1985; Francolinus (Lambrechtia) minor Jánossy 1974a]
      • P. subfrancolinus (Jánossy 1976b) [Francolinus subfrancolinus Jánossy 1976b; Plioperdix subfrancolinus (Jánossy 1976b) Mlíkovský 1995b]
    • Pliogallus Běljajeva 1948
      • P. coturnoides Běljajeva 1948
      • P. csarnotanus Kessler & Horváth 2022
    • Plioperdix Kretzoi 1955 [Pliogallus Gaillard 1939 non Tugarinov 1940]
      • P. crassipes (Gaillard 1939) [Pliogallus crassipes Gaillard 1939]
      • P. kormosi (Gaillard 1939) [Pliogallus kormosi Gaillard 1939]
      • P. africana Mourer-Chauviré & Geraads 2010
      • P. hungarica (Jánossy 1991) Zelenkov & Panteleyev 2014 [Palaeocryptonyx hungaricus Jánossy 1991]
    • Rustaviornis georgicus Burchak-Abramovich & Meladze 1972
    • Rhegminornis calobates Wetmore 1943 (Early Miocene of Bell, US)
    • Schaubortyx keltica (Eastman 1905) Brodkorb 1964 [Taoperdix keltica Eastman 1905] (middle Eocene – Early Oligocene)
    • Shandongornis Yeh 1997
    • Shanxiornis fenyinis Wang et al. 2006
    • Tologuica Zelenkov & Kuročkin 2009 (Middle Miocene of Sharga, Mongolia)
      • T. aurorae Zelenkov & Kuročkin 2009
      • T. karhui Zelenkov & Kuročkin 2009
    • Placement unresolved
      • Tetraoninae gen. et sp. indet. (Sajóvölgyi Middle Miocene of Mátraszõlõs, Hungary)
      • Meleagridae gen. et sp. indet. (Late Miocene of Westmoreland County, US)
      • "Tympanchus" stirtoni Miller 1944 (Early Miocene of South Dakota)
      • "Tympanuchus" lulli Shuefeldt 1915 (Pleistocene? of New Jersey)
    • Extant genera present in the fossil record

Gulls, auks, shorebirds

  • Basal and unresolved taxa
    • Charadriiformes gen. et sp. indet. (Late Cretaceous) – burhinid? basal?
    • "Morsoravis" (Late Paleocene/Early Eocene) – a nomen nudum?
    • Jiliniornis (middle Eocene) – charadriid?
    • Boutersema (Early Oligocene) – glareolid?
    • Turnipax (Early Oligocene) – turnicid?
    • Elorius (Early Miocene)
    • "Larus desnoyersii (Early Miocene of SE France) – larid? stercorarid?
    • "Larus pristinus (John Day Early Miocene of Willow Creek, US) – larid?
    • Charadriiformes gen. et spp. indet. (Early/Middle Miocene) – several species, 1 probably larid
    • Charadriiformes gen. et sp. indet. (Middle Miocene)
    • "Totanus" teruelensis (Late Miocene of Los Mansuetos, Spain) – scolopacid? larid?
    • "Actitis" balcanica (Late Pliocene of Varshets, Bulgaria) – scolopacid? charadriid?
  • Scolopacidae – waders and snipes
    • Paractitis (Early Oligocene)
    • Mirolia (Middle Miocene)
    • Placement unresolved
      • Scolopacidae gen. et sp. indet. (Middle – Late Miocene)
      • Scolopacidae gen. et sp. indet. (Early Pliocene)
    • Extant genera present in the fossil record
      • Limosa (Late Eocene? – Recent)
      • Tringa (Late Eocene/Early Oligocene? – Recent) – includes Totanus
      • Gallinago (Late Miocene/Early Pliocene – Recent)
      • Scolopax (Early/Middle Pliocene? – Recent)
      • Phalaropus (Middle Pliocene – Recent)
      • Actitis (Late Pliocene – Recent)
      • Numenius (Late Pleistocene – Recent) – includes Palnumenius
  • Jacanidae – jacanas
    • Nupharanassa (Early Oligocene)
    • Janipes
    • Prehistoric species of extant genera
      • Jacana farrandi
  • Laridae – gulls
    • Laridae gen. et sp. indet. (Early Oligocene)
    • Laricola (Late Oligocene/Early Miocene) – larid? Formerly "Larus" elegans and "L." totanoides
    • Gaviota Miller & Sibley 1941 non Fischer 1983 (Middle/Late Miocene)
    • Extant genera present in the fossil record
      • Larus (Middle Miocene – Recent)
  • Alcidae – auks
    • Hydrotherikornis (Late Eocene)
    • Pseudocepphus (Middle – Late Miocene)
    • Petralca (Early –? Late Oligocene)
    • Miocepphus (Middle Miocene)
    • Alcodes (Late Miocene)
    • Praemancalla (Late Miocene – Early Pliocene)
    • Mancalla (Late Miocene – Early Pleistocene)
    • Extant genera present in the fossil record
      • Cepphus (Late Miocene – Recent)
      • Cerorhinca (Late Miocene – Recent)
      • Uria (Late Miocene – Recent)
      • Aethia (Late Miocene – Recent)
      • Alca (Late Miocene/Early Pliocene – Recent)
      • Synthliboramphus (Late Miocene/Early Pliocene – Recent)
      • Fratercula (Early Pliocene – Recent)
      • Pinguinus (Early Pliocene – Recent)
      • Brachyramphus (Late Pliocene – Recent)
      • Ptychoramphus (Late Pliocene – Recent)
  • Stercorariidae – skuas and jaegers
    • Prehistoric species of extant genera
      • Stercorarius sp. (Middle Miocene)
      • Stercorarius shufeldti (Fossil Lake Middle Pleistocene of WC US)
      • Prehistoric subspecies of extant species
        • Stercorarius pomarinus philippi
  • Glareolidae – pratincoles
    • Paractiornis (Agate Fossil Beds Early Miocene of Sioux County, US)
    • Mioglareola (Early Miocene of Czech Republic) - formerly "Larus" dolnicensis
    • Prehistoric species of extant genera
      • Glareola neogena
  • Burhinidae – thick-knees
    • Prehistoric species of extant genera
      • Burhinus lucorum (Early Miocene)
      • Burhinus aquilonaris
      • Burhinus sp. (Cuba, West Indies)
      • Burhinus sp. (Late Pleistocene of Las Higueruelas, Spain)
      • Prehistoric subspecies of extant species
        • Burhinus bistriatus nanus (Bahamas, West Indies)
  • Charadriidae – plovers
    • Limicolavis (John Day Early Miocene of Malheur County, US)
    • Viator (Late Pleistocene of Talara, Peru) – may be synonym of Vanellus (or Belanopteryx if valid)
    • Extant genera present in the fossil record
      • Vanellus (Middle/Late Pleistocene – Recent) – includes Belanopteryx
    • Additional prehistoric species of extant genera
      • Oreopholus orcesi
  • Recurvirostridae – avocets
    • Extant genera present in the fossil record
      • Himantopus (Late Miocene – Recent)
    • Additional prehistoric species of extant genera
      • Recurvirostra sanctaeneboulae

Gastornithiformes (extinct)

The diatrymas, a group of huge flightless Paleogene birds of unclear affinities. Traditionally placed within the Gruiformes, they are usually considered a distinct order nowadays and appear closer to the Anseriformes.

  • Gastornithidae
    • Ornitholithes biroi Dughi & Sirugue 1969 [ootaxa- Gastornis?]
    • Ornithoformipes controversus [ichnotaxa]
    • Rivavipes giantess [ichnotaxa]
    • †Gasthornithidae gen. et sp. indet. (Paleocene) – possibly Gastornis
    • †Gasthornithidae gen. et sp. indet. YPM PU 13258 (Early Eocene) – possibly juvenile Gastornis giganteus
    • "Diatryma" cotei Galliard 1936 (middle-Late Eocene)
    • Gastornis Hébert 1855 [Diatryma Cope 1876; Barornis Marsh 1894; Omorhamphus Sinclair 1928; Palaeornis Bonaparte 1856; Macrornis Seeley 1866; Zhongyuanus Hou 1980] (Late Paleocene – middle Eocene)
      • G. parisiensis Hébert 1855 [Gastornis edwardsii Lemoine 1878; Gastornis klaasseni Newton 1885]
      • G. giganteus (Cope 1876) [Diatryma gigantea Cope 1876; Barornis regens Marsh 1894; Diatryma regens (Marsh 1894); Omorhamphus storchii Sinclair 1928 (juvenile); Diatryma steini Matthew & Granger 1917; Diatryma ajax Shufeldt 1913; Gastornis ajax (Shufeldt 1913) Brodkorb 1967]
      • G. sarasini (Schaub 1929) Mlíkovský 2002 [Diatryma sarasini Schaub, 1929b; Diatryma geiselensis Fischer 1978; Gastornis geiselensis (Fischer 1978)]
      • G. russeli Martin 1992
      • G. xichuanensis (Hou 1980) Buffetaut 2013 [Zhongyuanus xichuanensis Hou 1980]

The group that includes modern rails and cranes. Probably paraphyletic.

  • Placement unresolved
    • Rupelrallus Fischer 1997 (Early Oligocene)– rallid? parvigruid?
      • R. belgicus Mayr 2012
      • R. saxoniensis Fischer 1997
    • †"Gruiformes" gen. et sp. indet. MNZ S42623 (Early/Middle Miocene)– Aptornithidae?
  • Songziidae Hou 1990 possibly a nomen nudum
  • Eogruidae Wetmore 1934
    • Sonogrus gregalis Kuročkin 1981 (Ergilin Dzo Late Eocene/Early Oligocene of Khor Dzan, Mongolia)
    • Eogrus Wetmore 1932 [Progrus Bendukidze 1971] (Irdin Manha Middle/Late Eocene – Tung Gur Late Miocene/Early Pliocene of Mongolia)
      • E. aeola Wetmore 1932
      • E. crudus Kuročkin 1981
      • E. turanicus (Bendukidze 1971) [Progrus turanicus Bendukidze 1971]
      • E. wetmorei Brodkorb 1967
    • Ergilornithinae
    • Proergilornis minor Kozlova 1960 (Early/middle Oligocene of Ergil-Obo, Mongolia)
    • Ergilornis Kozlova 1960(Early/middle Oligocene of Ergil-Obo, Mongolia)
      • E. rapidus
      • E. minor
    • Amphipelargus Lydekker 1891 [Urmiornis Harrison 1981]
      • A. brodkorbi (Karkhu 1997) [Urmiornis brodkorbi Karkhu 1997]
      • A. majori Lydekker 1891 [Amphipelargus maraghanus (Mecquenem 1925) Kuročkin 1985; Urmiornis maraghanus Mecquenem 1925] [treated as ciconiid by Louchart et al. 2005]
      • A. dzabghanensis Kuročkin 1985
      • A. cracrafti (Harrison & Walker 1982) Kuročkin 1985 [Urmiornis cracrafti Harrison & Walker 1982]
      • A. ukrainus (Kuročkin 1981) Kuročkin 1985 [Urmiornis ukrainus Kuročkin 1981]
      • A. orientalis (Kuročkin 1981) Kuročkin 1985 [Urmiornis orientalis Kuročkin 1981]
  • Geranoididae Wetmore 1933
    • Eogeranoides campivagus Cracraft 1969 (Willwood Early Eocene of Foster Gulch, US)
    • Geranoides jepseni Wetmore 1933 (Willwood Early Eocene of South Elk Creek, US)
    • Geranodornis aenigma Cracraft 1969 (Bridger middle Eocene of Church Buttes, US)
    • Galligeranoides boriensis Bourdon, Mourer-Chauviré & Laurent 2014
    • Palaeophasianus Shufeldt 1913 (Willwood Early Eocene of Bighorn County, US)
      • P. meleagroides Shufeldt 1913
      • P. incompletus Cracraft 1969
    • Paragrus Lambrecht 1933 (Early Eocene of WC US)
      • P. prentici (Loomis 1906) Lambrecht 1933 [Gallinuloides prentici Loomis 1906; Grus prentici (Loomis 1906)]
      • P. shufeldti Cracraft 1969
  • Parvigruidae Mayr 2005
  • Aramidae Bonaparte 1854
    • Badistornis aramus Wetmore 1940 (middle Oligocene)– aramid?
    • Extant genera present in the fossil record
  • Gruidae – cranes
    • Camusia quintanai Seguí 2002 (Late Miocene of Menorca, Mediterranean)
    • Aramornis Wetmore 1926 (Middle Miocene) – gruid? aramid?
      • A. longurio Wetmore 1926
      • A. robalearica
      • A. crataegensis
    • Palaeogrus Portis 1885 (middle Eocene of Germany and Italy – Middle Miocene of France)
      • P. princeps Portis 1884 [Ornitocnemus robustus (sic) Zigno 1875 nomen nudum; Grus princeps (Portis 1885) Lydekker 1891a]
      • P. hordwelliensis (Lydekker 1891) Lambrecht 1933 [Grus hordwellianus Lydekker 1891; Ornitocnemus hordwelliensis (Lydekker) Brodkorb 1967]
      • P. mainburgensis Göhlich 2003
    • Geranopsis hastingsiae Lydekker 1871 Hordwell Late Eocene– Early Oligocene of England)– gruiform (gruid?) or anseriform (anseranatid?)?
    • Eobalearica tugarinovi Gureev 1949 (Ferghana Late? Eocene of Ferghana, Uzbekistan)
    • Pliogrus (Early Pliocene of Eppelsheim, Germany)
    • Probalearica mongolica Kurochkin 1985 (Late Oligocene? – Middle Pliocene) gruid? A nomen dubium?
    • Gruidae gen. et sp. indet. – formerly Grus conferta Miller & Sibley 1942 (Late Miocene/Early Pliocene of Contra Costa County, US)
    • Extant genera present in the fossil record
      • Balearica (Early Miocene – Recent)
        • B. crataegensis (Brodkorb 1963) [Probalearica crataegensis Brodkorb 1963]
        • B. excelsa (Milne-Edwards 1871) Mlíkovský 2002 [Palaeogrus excelsa (Milne-Edwards 1871) Lambrecht 1933; Grus excelsa Milne-Edwards 1871; Ornitocnemus excelsus (Milne-Edwards 1871) Brodkorb 1967]
        • B. exigua Feduccia & Voorhies 1992
        • B. rummeli (Mlíkovský 1998) Mourer-Chauviré 2001 [Basityto rummeli Mlíkovský 1998] (giant barn-owl)
      • Grus (Middle/Late Miocene – Recent)
        • G. bogatshevi
        • G. haydeni Marsh 1870
        • G. latipes [Baeopteryx latipes]
        • G. moldavica (Kuročkin & Ganya 1972) Mlíkovský 2002 [Probalearica moldavica Kuročkin & Ganya 1972]
        • G. nannodes Wetmore & Martin 1930
        • G. pagei Campbell 1995
        • G. pentelici Gaudry 1862 [Pliogrus pentelici (Gaudry 1862) Lambrecht 1933; G. afghana Mourer-Chauviré et al. 1985]
        • G. primigenia Milne-Edwards 1869
        • G. melitensis Lydekker 1890 (Maltese crane)
  • Rallidae – rails
    • Aletornis Marsh 1872 [Protogrus Lambrecht 1933] (middle Eocene)
      • A. marshi (Shufeldt 1915) Lambrecht 1933 [Grus marshi Shufeldt 1915; Protogrus marshi (Shufeldt 1915)]
      • A. bellus Marsh 1872
      • A. gracilis Marsh 1872
      • A. nobilis Marsh 1872 [Grus nobilis (Marsh 1872) Shufeldt; Protogrus nobilis (Marsh 1872) Lambrecht 1933]
      • A. pernix Marsh 1872
    • Australlus Worthy & Boles 2011
      • A. disneyi (Boles 2005) Worthy & Boles 2011 [Gallinula disneyi Boles 2005]
      • A. gagensis Worthy & Boles 2011
    • Baselrallus intermedius De Pietri & Mayr 2014
    • Belgirallus Mayr & Smith 2001 (Early Oligocene)
      • B. minutus Mayr & Smith 2001
      • B. oligocaenus Mayr & Smith 2001
    • Creccoides osbornii Shufeldt 1892 (Late Pliocene/Early Pleistocene)
    • Eocrex Wetmore 1931 (Early Eocene)
      • E. primus Wetmore 1931
      • E. tagusevae Panteleyev 2001
    • Euryonotus Mercerat 1897 (Pleistocene)– rallid?
      • E. uspallatensis Mercerat 1897
      • E. brachypterus Mercerat 1897
      • E. argentinus Mercerat 1897
    • Fulicaletornis venustus (Marsh 1872) Lambrecht 1933 [Aletornis venustus Marsh 1872] (middle Eocene)
    • Ibidopsis hordwelliensis Lydekker 1891 (Late Eocene)
    • Latipons Harrison & Walker 1979 (middle Eocene)
      • L. gardneri Harrison & Walker 1979
      • L. robinsoni Harrison & Walker 1979
    • Miofulica dejardini (Van Beneden 1871) Lambrecht 1933 [Fulica dejardinii Van Beneden 1871] (Middle Miocene)
    • Miorallus major (Milne-Edwards 1869) Lambrecht 1933 [Rallus major Milne-Edwards 1869] (Middle – Late Miocene)
    • Palaeoaramides Lambrecht 1933 [Pararallus Lambrecht 1933; Tertiariaporphyrula Kuročkin & Ganya 1972] (Late Oligocene/Early Miocene – Late Miocene)
      • P. lungi (Kuročkin & Ganya 1972) Olson 1977 [Tertiariaporphyrula lungi Kuročkin & Ganya 1972; Mioporphyrula lungi (Kurochkin &Ganya 1972)]
      • P. minutus Cracraft 1973
      • P. tugarinovi Kuročkin 1980 [Tertiariaporphyrula tugarinovi (Kurochkin, 1980)]
      • P. christyi (Milne-Edwards 1869) Lambrecht 1933 [Rallus christyi Milne-Edwards 1869; Rallus eximius Milne-Edwards 1869; Palaeoaramides eximius (Milne-Edwards 1869) Brodkorb 1967]
      • P. minor [Rallus minor]
      • P. beaumontii (Milne-Edwards 1869) Cracraft 1973 [Rallus beaumontii Milne-Edwards 1869; Pararallus beaumontii (Milne-Edwards 1869) Brodkorb 1967); Tertiariaporphyrula beaumonti (Milne-Edwards) Kuročkin & Ganea 1972; Rallus dispar Milne-Edwards 1869; Pararallus dispar (Milne-Edwards 1869) Lambrecht 1933; Palaeoaramides dispar (Milne-Edwards 1869)]
    • Palaeorallus Wetmore 1931 (Early Eocene)
      • P. troxelli Wetmore 1931
      • P. brodkorbi Cracraft 1973
    • Paraortygometra porzanoides (Milne-Edwards 1871) Lambrecht 1933 [Microrallus Švec 1983; Rallus porzanoides Milne-Edwards 1871; Microrallus fejfari Švec 1983] (Late Oligocene/?Early Miocene –? Middle Miocene)
    • Parvirallus Harrison & Walker 1979 (Early – middle Eocene)
      • P. gracilis Harrison & Walker 1979
      • P. bassetti Harrison 1984
      • P. gassoni Harrison 1984
      • P. medius Harrison 1984
    • Pastushkinia zazhigini (Kuročkin 1980) Zelenkov 2013 [Crex zazhigini Kuročkin 1980]
    • Pleistorallus flemingi Worthy 1997 (Fleming's rails)
    • Quercyrallus (Late Eocene –? Late Oligocene)
      • Q. ludianus Brodkorb 1963 [Quercyrallus intermedius Milne-Edwards 1868; Rallus adelus Oberholser 1917; Ludiortyx adelus (Oberholser 1917)]
      • Q. arenarius (Milne-Edwards 1891) Lambrecht 1933 [Rallus arenarius Milne-Edwards 1891]
      • Q. quercy Cracraft 1973
    • Rallicrex (Middle/Late Oligocene)
      • R. litkensis Kessler & Hir 2012
      • R. polgardiensis Jánossy 1991
      • R. kolozsvarensis Lambrecht 1933
    • Rhenanorallus rhenanus Mayr 2010
    • Wanshuina lii Hou 1994
    • Youngornis (Middle Miocene)
      • Y. gracilis Yeh 1981
      • Y. qiluensis Yeh & Bo 1989
    • Placement unresolved
      • Rallidae gen. et sp. indet. (Late Oligocene)
      • Rallidae gen. et spp. indet. (Early/Middle Miocene)– several species
      • Rallidae gen. et sp. indet. (Middle Miocene)
      • Rallidae gen. et sp. indet. (Late Miocene)
      • Rallidae gen. et sp. indet. UMMP V55013/-14; UMMP V55012/V45750/V45746 (Late Pliocene)
      • Rallidae gen. et sp. indet. UMMP V29080 (Late Pliocene)
      • Rallidae gen. et sp. indet. (Bermuda, West Atlantic)
    • Extant genera present in the fossil record
  • Itaboravis elaphrocnemoides Mayr, Alvarenga & Clarke 2011
  • Walbeckornis creber Mayr 2007
  • Qianshanornithidae Mayr et al. 2013
  • Salmilidae Mayr 2002
    Salmila
    • Salmila robusta Mayr 2000
  • Ameghinornithidae Mourer-Chauviré 1981
    • Strigogyps Gaillard 1908 [Aenigmavis Peters 1987; Ameghinornis Mourer-Chauviré 1981; Eocathartes Lambrecht 1935 non Wetmore 1944; Geiseloceros Lambrecht 1935]
      • S. dubius Gaillard 1908
      • S. robustus (Lambrecht 1935) [Eocathartes robustus Lambrecht 1935; Geiseloceros robustus Lambrecht 1935] (middle Eocene of Germany)
      • S. sapea (Peters 1987) [Aenigmavis sapea Peters 1987; Ameghinornis minor Mourer-Chauviré 1981]
  • Bathornithidae Wetmore 1933
    • Eutreptornis uintae Cracraft 1971 (Uinta Late Eocene of Ouray Agency, US)
    • Neocathartes grallator (Wetmore 1944) Wetmore 1950 [Eocathartes Wetmore 1944 non Lambrecht 1935; Eocathartes grallator Wetmore 1944; Bathornis grallator (Wetmore 1944) Olson 1985] (Late Eocene)
    • Paracrax Brodkorb 1964 (Early/middle Oligocene of Gerry's Ranch, US – Brule Late Oligocene of South Dakota, US)
      • P. antiqua (Marsh 1871) Cracraft 1971 [Meleagris antiquus Marsh 1871; Oligocorax mediterraneus (Shufeldt 1915); Phalacrocorax mediterraneus Shufeldt 1915]
      • P. wetmorei Cracraft 1968
      • P. gigantea Cracraft 1968
    • †Bathornithidae gen. nov. (Early – middle Oligocene of C US) – formerly Bathornis celeripes Wetmore 1933 and B. cursor Wetmore 1933
    • Bathornis Wetmore 1927 (Early Oligocene – Early Miocene of C US)
      • B. minor Cracraft 1971
      • B. veredus Wetmore 1927 [Palaeogyps prodromus Wetmore 1927] (Early Oligocene of WC North America)
      • B. fax (Wetmore 1927) Cracraft 1968 [Palaeocrex fax Wetmore 1927]
      • B. geographicus Wetmore 1942
      • B. fricki Cracraft 1968
Paraphysornis
  • Idiornithidae Brodkorb 1965 [Orthocnemidae]
    • Dynamopterus Milne-Edwards 1892 [Idiornis Oberholser 1899; Orthocnemus Milne-Ewards 1892 non Jekel 1854] (middle Eocene of Messel, Germany –? Quercy Phosphorites middle Oligocene of Quercy, France)
      • I. cursor (Milne-Edwards 1892) Oberholser 1899 [Orthocnemus cursor Milne-Edwards 1892; Orthocnemus major Milne-Edwards 1892; Idiornis major (Milne-Edwards 1892) Oberholser 1899]
      • I. gallicus (Milne-Edwards 1892) Oberholser 1899 [Filholornis gravis Milne-Edwards 1892; Orthocnemus gallicus Milne-Edwards 1892; Agnopterus gallicus (Milne-Edwards 1892) Mlíkovský 2002]
      • I. gracilis (Milne-Edwards 1892) Mourer-Chauviré 1983 [Elaphrocnemus gracilis Milne-Edwards 1892; Agnopterus gracilis (Milne-Edwards 1892) Mlíkovský 2002]
      • I. minor (Milne-Edwards 1892) Oberholser 1899 [Orthocnemus minor Milne-Edwards 1892; Agnopterus minor (Milne-Edwards 1892) Mlíkovský 2002]
      • D. anthracinus (Mayr 2002) Mourer-Chauviré 2013 [Idiornis anthracinus Mayr 2002]
      • D. tuberculata (Peters 1995) Mourer-Chauviré 2013 [Idiornis tuberculata Peters 1995]
      • D. gaillardi (Cracraft 1973) Mourer-Chauviré 2013 [Idiornis gaillardi Cracraft 1973]
      • D. itardiensis (Mourer-Chauviré 1983) Mourer-Chauviré 2013 [Idiornis itardiensis Mourer-Chauviré 1983; Agnopterus itardiensis (Mourer-Chauviré 1983) Mlíkovský 2002]
    • Talantatos Reichenbach 1852 [Elaphrocnemus Milne-Edwards 1892; Filholornis Milne-Edwards 1892] (Late Eocene/Early Oligocene of France)
      • T. bordkorbi (Mourer-Chauviré 1983) Mlíkovský 2002 [Elaphrocnemus bordkorbi Mourer-Chauviré 1983]
      • T. crex (Milne-Edwards 1892) [Elaphrocnemus crex Milne-Edwards 1892]
      • T. phasianus (Milne-Edwards 1892) [Elaphrocnemus phasianus Milne-Edwards 1892; Filholornis debilis Milne-Edwards 1892; Filholornis paradoxa Milne-Edwards 1892]
      • T. fossilis (Giebel 1847) Reichenbach 1852 [Tantalus fossilis Giebel 1847; Orthocnemus major Milne-Edwards 1892; Idiornis major (Milne-Edwards 1892) Oberholser 1899; Telecrex peregrinus Mlíkovský 1989b]
      • T. proudlocki (Harrison & Walker 1979) Mlíkovský 2002 [Percolinus proudlocki Harrison & Walker 1979]
    • Gypsornis Milne-Edwards 1869 [Percolinus Harrison & Walker 1977] (Montmartre Late Eocene of Montmartre, France)
      • G. cuvieri Milne-Edwards 1869
      • G. venablesi (Harrison & Walker 1977) Mlíkovský 2002 [Percolinus venablesi Harrison & Walker 1977]
    • Propelargus Lydekker 1891 [Oblitavis Mourer-Chauviré 1983] (Late Eocene/Early Oligocene)
      • P. edwardsi
      • P. insolitus (Mourer-Chauviré 1983) Mlíkovský 2002 [Oblitavis insolitus Mourer-Chauviré 1983]
      • P. cayluxensis Lydekker 1891
    • Occitaniavis elatus (Milne-Edwards 1892) Mourer-Chauvire 1983 [Geranopsis Milne-Edwards 1892 non Lydekker 1891a; Geranopsis elatus Milne-Edwards 1892; Propelargus elatus (Milne-Edwards 1892) Mlíkovský 2002]
    • Ibidopodia Milne-Edwards 1868 (Late Oligocene/Early Miocene of France) – threskiornithid?
      • I. chavrochensis De Pietri & Mayr 2014
      • I. minuta De Pietri & Mayr 2014
      • I. palustris Milne-Edwards 1868
  • Phorusrhacidae Ameghino 1899 – Terror birds
    • Lavocatavis africana Mourer-Chauviré et al. 2011
    • Patagorhacos terrificus Agnolin & Chafrat 2015
    • Eleutherornis Schaub 1940 (Late Eocene)
      • E. helveticus Schaub 1940
      • E. cotei Gaillard 1936
    • Paleopsilopterus itaboraiensis Alvarenga 1985 (Middle Paleocene)
    • Procariama simplex Rovereto 1914 (Late Miocene – Early Pliocene)
    • Psilopterus Moreno & Mercerat 1891 [Pelecyornis Ameghino 1891; Staphylornis Mercerat 1897] (middle Oligocene – Late Miocene)
      • P. bachmanni (Moreno & Mercerat 1891) [Patagornis bachmanni Moreno & Mercerat 1891; Psilopterus communis Moreno & Mercerat 1891; Psilopterus intermedius Moreno & Mercerat 1891; Phororhacos delicatus Ameghino 1891; Pelecyornis puerredonensis Sinclair & Farr 1932]
      • P. lemoinei (Moreno & Mercerat 1891) [Patagornis lemoinei Moreno & Mercerat 1891; Psilopterus australis Moreno & Mercerat 1891; Pelecyornis tubulatus Ameghino 1895; Phorohacos modicus Ameghino 1895; Staphylornis gallardoi Mercerat 1897; Staphylornis erythacus Mercerat 1897; Pelecyornis tenuirostris Sinclair & Farr 1932]
      • P. affinis (Ameghino 1899) [Phororhacos affinis Ameghino 1899]
      • P. colzecus Tonni & Tambussi 1988
    • Llallawavis scagliai Degrange et al. 2015
    • Mesembriornis Moreno 1889 [Paleociconia Moreno 1889; Prophorohacos Rovereto 1914; Hermosiornis Rovereto 1914] (Late Miocene – Late Pliocene)
      • M. milneedwardsi Moreno 1889 [Paleociconia australis Moreno 1889; Driornis pampeanus Moreno & Mercerat 1891; Hermosiornis milneedwardsi Rovereto 1914; Hermosiornis rapax Kraglievich 1946; Prophororhacos australis Brodkorb 1967]
      • M. incertus (Rovereto 1914) [Prophororhacos incertus Rovereto 1914]
    • Patagornis marshi Moreno & Mercerat 1891 [Morenomercatia Lambrecht 1933; Tolmodus Ameghino 1891; Tolmodus inflatus Ameghino 1891; Phorusrhacos inflatus Ameghino 1891; Paleociconia cristata (Lambrecht 1933) Brodkorb 1967; Morenomerceraria cristata Lambrecht 1933] (Early – Middle Miocene)
    • Andrewsornis abbotti Paterson 1941 (middle – Late Oligocene)
    • Andalgalornis steulleti (Kraglievich 1931) Patterson & Kraglievich 1960 [Phororhacos steulleti Kraglijevič 1931; Phororhacos deautieri Kraglijevič 1931; Andalgalornis ferox Patterson & Kraglijevič 1960] (Late Miocene – Early Pliocene)
    • Phorusrhacos longissimus Ameghino 1887 [Stereornis Moreno & Mercerat 1891; Darwinornis Moreno & Mercerat 1891; Owenornis Moreno & Mercerat 1891; Titanornis Mercerat 1893; Callornis Ameghino 1895 non Calornis Dalman 1820; Eucallornis Ameghino 1901; Liornis Ameghino 1895; Phororhacos longissimus Ameghino 1889; Phororhacos platygnathus Ameghino 1891; Phororhacos sehuensis Ameghino 1891; Stereornis rollieri Moreno & Mercerat 1891; Stereornis gaundryi Moreno & Mercerat 1891; Mesembriornis studeri Moreno & Mercerat 1891; Mesembriornis quatrefragesi Moreno & Mercerat 1891; Darwinornis copei Moreno & Mercerat 1891; Darwinornis zittelli Moreno & Mercerat 1891; Darwinornis socialis Moreno & Mercerat 1891; Owenornis affinis Moreno & Mercerat 1891; Owenornis lydekkeri Moreno & Mercerat 1891; Titanornis mirabilis Mercerat 1893; Callornis giganteus Ameghino 1895; Eucallornis giganteus Ameghino 1901; Liornis floweri Ameghino 1895; Liornis minor Dolgopol de Saez 1927] (Early – Middle Miocene)
    • Devincenzia pozzi (Kraglievich 1931) Kraglievich 1932 [Onactornis Cabrera 1939; Phorohacos pozzi Kraglievich 1931; Phorohacos longissimus mendocinus Kraglievich 1931; Devincenzia gallinali Kraglievich 1932; Onactornis depressus Cabrera 1939; Onactornis pozzi (Kraglievich 1931) Brodkorb 1967; Onactornis mendocinus (Kraglievich 1931) Brodkorb 1967] (Late Miocene – Early Pliocene)
    • Titanis walleri Brodkorb 1963 (Early – Late Pliocene)
    • Kelenken guillermoi Bertelli, Chiappe & Tambussi 2007 (Early – Late Pliocene)
    • Physornis fortis Ameghino 1895 [Aucornis Ameghino 1898; Aucornis euryrhynchus Ameghino 1898] (middle – Late Oligocene)
    • Paraphysornis brasiliensis (Alvarenga 1982) Alvarenga 1993 [Physornis brasiliensis Alvarenga 1982] (Late Oligocene/Early Miocene)
    • Brontornis burmeisteri Moreno & Mercerat 1891 [Rostrornis Moreno & Mercerat 1891; Rostrornis floweri Moreno & Mercerat 1891; Brontornis platyonyx Ameghino 1895] (Early – Middle Miocene)
  • Cariamidae Bonaparte 1850 – seriemas
    • Riacama caliginea Ameghino 1899
    • Pseudolarus guaraniticus Ameghino 1899 [Pseudogavia Sharpe 1892; Ameghinia Sharpe 1899]
    • Smiliornis penetrans Ameghino 1899
    • Noriegavis santacrucensis (Noriega, Vizcaino & Bargo 2009) Agnolin 2009 [Cariama santacrucensis Noriega, Vizcaino & Bargo 2009]
    • Prehistoric species of extant genera
      • Chunga
        • C. incerta Tonni 1974
  • Otididae – bustards
    • Gryzaja odessana Zubareva 1939 [Chlamydotis pliodeserti Serebrovskyj 1941; Otis gryzaja Vojinstvens'kyj 1967]
    • Ioriotis gabunii Burchak-Abramovich & Vekua 1981
    • Miootis compactus Umanskaya 1979
    • Pleotis liui Hou 1982
    • Prehistoric species of extant genera
  • Placement unresolved
  • Palaelodidae Stejneger 1885 – Swimming-flamingos
    • Adelalopus hoogbutseliensis Mayr & Smith 2002 (stout-legged flamingos) - (Borgloon Early Oligocene of Hoogbutsel, Belgium)
    • Palaelodus Milne-Edwards 1863 [Pliogrus Lambrecht 1933] (middle Oligocene –? Middle Pleistocene)
      • P. ambiguus Milne-Edwards 1863 [Grus problematica Milne-Edwards 1871; Probalearica problematica (Milne-Edwards 1871); Palaelodus gracilipes Milne-Edwards 1863; Palaelodus crassipes Milne-Edwards 1863; Paloelodus minutus (sic) Milne-Edwards 1868, Grus miocenicus Grigorescu & Kessler 1977]
      • P. aotearoa Worthy et al. 2010
      • P. germanicus (Lambrecht 1933) [Pliogrus germanicus Lambrecht 1933]
      • P. kurochkini Zelenkov 2013
      • P. pledgei Baird & Vickers-Rich 1998
      • P. wilsoni Baird & Vickers-Rich 1998
    • Megapaloelodus Miller 1944 (Late Oligocene – Early Pliocene)
  • Phoenicopteridae – flamingos
    • Harrisonavis croizeti (Gervais 1852) Kashin 1978 [Gervaisia Harrison & Walker 1976 non Bonaparte 1854 non Waga 1858 non Robineau-Desvoidy 1863; Phoenicopterus croizeti Gervais 1852; Gervaisia croizeti (Gervais 1852) Harrison & Walker 1976]
    • Leakeyornis aethiopicus (Harrison & Walker 1976) Vickers-Rich & Walker 1983 [Phoenicopterus aethiopicus Harrison & Walker 1976]
    • Elornis Milne-Edwards 1868 [Elornis Aymard 1856 nomen nudum; Helornis Lydekker 1891; Actiornis] (Middle? Eocene – Early Oligocene)
      • E. anglicus Aymard 1856 nomen nudum [Helornis anglicus; Actiornis anglicus Lydekker 1891]
      • E. grandis Milne-Edwards 1868 [Elornis grandis1856 Aymard nomen nudum]
      • E. littoralis Milne-Edwards 1868 [Elornis littoralis Aymard 1856 nomen nudum; Elornis antiquus Aymard 1856 nomen nudum; Elornis antiquus Milne-Edwards 1868; Helornis littoralis (Milne-Edwards 1868) Lydekker 1891]
    • Placement unresolved
      • Phoenicopteridae gen. et sp. indet. (Middle? – Late Miocene)
    • Extant genera present in the fossil record
  • Podicipedidae – grebes
    • Miobaptus Švec 1982 (Early Miocene)
      • M. huzhiricus Zelenkov 2015
      • M. walteri Švec 1982 [Podiceps walteri (Švec 1984) Mlíkovský 2000]
    • Miodytes serbicus Dimitreijevich, Gál & Kessler 2002
    • Pliolymbus baryosteus Murray 1967 (Late Pliocene – Early? Pleistocene)
    • Thiornis sociata Navás 1922 [Podiceps sociatus (Navás 1922) Olson] (Late Miocene –? Early Pliocene)
    • Placement unresolved
      • Podicipedidae gen. et sp. indet. (Late Pliocene) – formerly included in Podiceps parvus
      • Podicipedidae gen. et sp. indet. UMMP 49592, 52261, 51848, 52276, KUVP 4484 (Late Pliocene)
      • Podicipedidae gen. et sp. indet. (Late Pliocene/Early Pleistocene)
    • Extant genera present in the fossil record
  • Prophaethontidae Harrison & Walker 1976
    • Lithoptila abdounensis Bourdon, Bouya & Iarochène 2005 (Late Paleocene – Early Eocene of Ouled Abdoun Basin, Morocco)– including Abdounornis
    • Prophaethon shrubsolei Andrews 1899 (Late Paleocene ?– Early Eocene)
  • Phaethontidae Brandt 1831 – tropicbirds
    • ?†Proplegadis fisheri Harrison & Walker 1971 (London Clay Early Eocene of England) – Threskiornithidae?
    • Phaethusavis pelagicus Bourdon, Amaghzaz & Bouya 2008 (Early Eocene of Ouled Abdoun Basin, Morocco)
    • Heliadornis Olson 1985 (Miocene of North America and Europe)
      • H. ashbyi Olson 1985
      • H. minor Kessler 2009
      • H. paratethydicus Mlíkovský 1997

The diverse group that includes storks, herons and New World vultures. Paraphyletic as listed here.

  • Ciconiidae – storks
    • Ciconiopsis antarctica Ameghino 1899
    • Palaeoephippiorhynchus dietrichi Lambrecht 1930 (Early Oligocene)
    • Grallavis edwardsi (Lydekker 1891c) Cheneval 1984 [Propelargus edwardsi Lydekker 1891c; Palaeoephippiorhynchus edwardsi (Lydekker 1891)] (Early Miocene) – may be same as Prociconia
    • Pelargosteon tothi Kretzoi 1962 (Early Pleistocene)
    • Sanshuiornis zhangi Wang et al. 2012
    • Placement unresolved
      • Ciconiidae gen. et sp. indet. – formerly Aquilavus bilinicus (Laube 1909) Brodkorb 1964; Cygnus bilinicus Laube 1909 (Early Miocene)
      • Ciconiidae gen. et sp. indet. (Late Miocene)
      • cf. Leptoptilos gen. et sp. indet. – formerly Leptoptilos siwalicensis Harrison 1974 (Late Miocene? - Late Pliocene)
      • Ciconiidae gen. et sp. indet. (Late Pleistocene) – Ciconia or Mycteria?
    • Extant genera present in the fossil record

The group that includes modern pelicans and cormorants. As presented here paraphyletic; the tropicbird lineage is not part of this group and relationships with Procellariiformes and Sphenisciformes require more research. Also, as the pelicans are at least as close to the Ciconiiformes as to cormorants, the latter group is being recognized as Phalacrocoraciiformes by some recent authors and the core Pelecaniformes are occasionally merged into the Ciconiiformes.

  • Basal and unresolved forms
    • Piscator tenuirostris Harrison & Walker 1976 (Early-Late Eocene of England) (Auk98:199)– basal phalacrococacoid?
    • "Prophalacrocorax ronzoni (Gervais 1849) Harrison 1975 [Sula ronzoni (Gervais 1849) Milne-Edwards 1897; Mergus ronzoni ronzoni Gervais 1849] (Early Oligocene of Ronzon, France)
    • † "Pelecaniformes" gen. et sp. indet. (Jebel Qatrani Early Oligocene of Fayum, Egypt) basal phalacrococacoid (similar to Piscator?)?
  • Fregatidae – frigatebirds
    • Limnofregata Olson 1977 (Early Eocene)
      • L. azygosternon Olson 1977
      • L. hutchisoni Stidham 2014
      • L. hasegawai Olson & Matsuoka 2005
  • Sulidae – gannets and boobies
    • Bimbisula melanodactylus Benson & Erickson 2013
    • Eostega lebedinskyi Lambrecht 1929 (Middle/Late Eocene)
    • Empheresula arvenensis (Milne-Edwards 1867) Harrison 1975 [Sula arvernensis Milne-Edwards 1867; Parasula] (Late Oligocene – Middle Miocene)
    • Masillastega rectirostris Mayr 2002 (middle Eocene)
    • Microsula pygmaea (Milne-Edwards 1874) Wetmore 1938 [Sula pygmaea; Microsula pygmaea; Sula avita; Enkurosula; Pseudosula] (Late Oligocene – Middle Miocene)
    • Miosula media Miller 1925 (Late Miocene)
    • Palaeosula stocktoni (Miller 1935) Howard 1958 (?Early Pliocene)
    • Rhamphastosula Stucchi & Urbina 2004 (Early Pliocene)
      • R. ramirezi Stucchi & Urbina 2004
      • R. aguirrei Stucchi, Varas-Malca & Urbina-Schmitt 2015
    • Sarmatosula dobrogensis Grigorescu & Kessler 1977 (Middle Miocene)
    • Placement unresolved
      • Sulidae gen. et sp. indet. (Late Oligocene)
      • Sulidae gen. et sp. indet. (Late Pliocene)
    • Extant genera present in the fossil record
      • Morus (Early Miocene – Recent)
        • M. avitus (Wetmore 1938) [Sula (Microsula) avita (Wetmore 1938); Microsula avitus Wetmore 1938]
        • M. lompocanus Domning 1978 [Sula lompocana (Domning 1978)]
        • M. loxostylus (Cope 1870) Brodkorb 1963 [Sula loxostyla Cope 1870; Morus atlanticus (Shufeldt 1915); Sula atlantica Shufeldt 1915]
        • M. olsoni Grigorescu & Kessler 1988
        • M. peninsularis Brodkorb 1955 [Sula peninsularis (Brodkorb 1955)]
        • M. peruvianus Stucchi 2003 [Sula peruviana (Stucchi 2003)]
        • M. recentior Howard 1949
        • M. reyanus Howard 1936 (Del Rey gannet)
        • M. vagabundus Wetmore 1930 [Sula vagabundus (Wetmore 1930)]
        • M. willetti (Miller 1925) [Sula willetti Miller 1925]
      • Sula (Middle Pliocene – Recent)
        • S. brandi Stucchi, Varas-Malca & Urbina-Schmitt 2015
        • S. figueroae Stucchi, Varas-Malca & Urbina-Schmitt 2015
        • S. clarki Chandler 1990
        • S. humeralis Miller & Bowman 1958 [Morus humeralis Miller & Bowman 1958]
        • S. magna (Howard 1978) Stucchi 2003 [Morus magnus Howard 1978]
        • S. pohli Howard 1958
        • S. sulita Stucchi 2003
        • S. universitatis Brodkorb 1963Lua error: Internal error: The interpreter has terminated with signal "24".
Copepteryx
  • Phalacrocoracidae – cormorants and shags
    • Borvocarbo Mourer-Chauviré, Berthet & Hugueney 2004Lua error: Internal error: The interpreter has terminated with signal "24". (Late Oligocene of C Europe) – phalacrocoracid? basal phalacrococacoid?
      • B. guilloti Mourer-Chauviré, Berthet & Hugueney 2004Lua error: Internal error: The interpreter has terminated with signal "24".
      • B. tardatus Göhlich & Mourer-Chauviré 2010Lua error: Internal error: The interpreter has terminated with signal "24".
    • Limicorallus Kuročkin 1968Lua error: Internal error: The interpreter has terminated with signal "24". (middle Oligocene)
      • L. saiensis Kuročkin 1968Lua error: Internal error: The interpreter has terminated with signal "24".
      • L. carbunculus Mayr 2009Lua error: Internal error: The interpreter has terminated with signal "24".
    • Nambashag Worthy 2011Lua error: Internal error: The interpreter has terminated with signal "24".
      • N. billerooensis Worthy 2011Lua error: Internal error: The interpreter has terminated with signal "24".
      • N. microglaucus Worthy 2011Lua error: Internal error: The interpreter has terminated with signal "24".
    • Nectornis Cheneval 1984Lua error: Internal error: The interpreter has terminated with signal "24". (Late Oligocene?/Early Miocene – Middle Miocene)
      • N. africanus Mayr 2014Lua error: Internal error: The interpreter has terminated with signal "24".
      • N. miocaenus (Milne-Edwards 1868)Lua error: Internal error: The interpreter has terminated with signal "24". [?Phalacrocorax miocaenus (Milne-Edwards 1868)Lua error: Internal error: The interpreter has terminated with signal "24".]
      • N. anatolicus (Mourer-Chauviré 1978) Mlíkovský 1998Lua error: Internal error: The interpreter has terminated with signal "24". [Phalacrocorax anatolicus Mourer-Chauviré 1978Lua error: Internal error: The interpreter has terminated with signal "24".]
    • Valenticarbo praetermissus Harrison 1979Lua error: Internal error: The interpreter has terminated with signal "24". (Late Pliocene/Early Pleistocene) – a nomen dubium
    • Oligocorax Lambrecht 1933
      • O. littoralis Milne-Edwards 1868Lua error: Internal error: The interpreter has terminated with signal "24". [Phalacrocorax littoralis (Milne-Edwards 1868)Lua error: Internal error: The interpreter has terminated with signal "24".]
      • O. stoeffelensis (Mayr 2007) Mayr 2014Lua error: Internal error: The interpreter has terminated with signal "24". [Oligocorax sp. Mayr 2001Lua error: Internal error: The interpreter has terminated with signal "24".; Borvocarbo stoeffelensis Mayr 2007Lua error: Internal error: The interpreter has terminated with signal "24".]
    • Placement unresolved
      • †Phalacrocoracidae gen. et sp. indet. (Late Eocene ?–? mid-Oligocene)
    • Extant genera present in the fossil record
      • Phalacrocorax (Oligocene –? Recent) – may be several genera. Includes Australocorax, Miocorax
        • P. gregorii De Vis 1905Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. gregorii (De Vis 1905)Lua error: Internal error: The interpreter has terminated with signal "24". [Australocorax vetustus De Vis 1905Lua error: Internal error: The interpreter has terminated with signal "24".]
        • P. chapalensis Alvarez 1977Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. destefani (Regalia 1902)Lua error: Internal error: The interpreter has terminated with signal "24". [Paracorax destefani Regalia 1902Lua error: Internal error: The interpreter has terminated with signal "24".]
        • P. femoralis (Miller 1929)Lua error: Internal error: The interpreter has terminated with signal "24". [Miocorax femoralis Miller 1929Lua error: Internal error: The interpreter has terminated with signal "24".]
        • P. filyawi Emslie 1995bLua error: Internal error: The interpreter has terminated with signal "24".
        • P. goletensis Howard 1965Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. ibericus Villalta 1963Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. idahensis (Marsh 1870)Lua error: Internal error: The interpreter has terminated with signal "24". [Graculus idahensis Marsh 1870Lua error: Internal error: The interpreter has terminated with signal "24".]
        • P. intermedius (Milne-Edwards 1867)Lua error: Internal error: The interpreter has terminated with signal "24". [Phalacrocorax praecarbo, Ardea brunhuberi; Phalacrocorax brunhuberi (Milne-Edwards)Lua error: Internal error: The interpreter has terminated with signal "24".; Botaurites avitus]
        • P. kennelli (Howard 1949)Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. kuehneanus Schlüter 1991Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. longipes (Tugarinov 1940)Lua error: Internal error: The interpreter has terminated with signal "24". [Pliocarbo longipes Tugarinov 1940Lua error: Internal error: The interpreter has terminated with signal "24".]
        • P. leptopus Brodkorb 1961Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. macer BrodkorbLua error: Internal error: The interpreter has terminated with signal "24".
        • P. macropus (Cope 1878)Lua error: Internal error: The interpreter has terminated with signal "24". [Graculus macropus Cope 1878Lua error: Internal error: The interpreter has terminated with signal "24".]
        • P. marinavis (Shufeldt 1915)Lua error: Internal error: The interpreter has terminated with signal "24". [Oligocorax marinavis Shufeldt 1915Lua error: Internal error: The interpreter has terminated with signal "24".]
        • P. mongoliensis Kuročkin 1971Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. owrei Brodkorb & Mourer-Chauviré 1984Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. reliquus Kuročkin 1976Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. rogersi (Howard 1932)Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. serdicensis Burčak-Abramovič & Nikolov 1984Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. tanzaniae Harrison & Walker 1979Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. wetmorei Brodkorb 1955Lua error: Internal error: The interpreter has terminated with signal "24".
  • Plotopteridae – diving-"boobies"
    • Copepteryx Olson & Hasegawa 1996Lua error: Internal error: The interpreter has terminated with signal "24".
      • C. hexeris Olson & Hasegawa 1996Lua error: Internal error: The interpreter has terminated with signal "24".
      • C. titan Olson & Hasegawa 1996Lua error: Internal error: The interpreter has terminated with signal "24".
    • Hokkaidornis abashiriensis Sakurai, Kimura & Katoh 2008Lua error: Internal error: The interpreter has terminated with signal "24".
    • Phocavis maritimus Goedert 1988Lua error: Internal error: The interpreter has terminated with signal "24".
    • Plotopterum joaquinensis Howard 1969Lua error: Internal error: The interpreter has terminated with signal "24".
    • Tonsala Olson 1980Lua error: Internal error: The interpreter has terminated with signal "24".
      • T. hildegardae Olson 1980Lua error: Internal error: The interpreter has terminated with signal "24".
      • T. buchanani Dyke, Xia & Habib 2011 Lua error: Internal error: The interpreter has terminated with signal "24".
  • Protoplotidae
    • Protoplotus beauforti Lambrecht 1931Lua error: Internal error: The interpreter has terminated with signal "24". (Paleocene? – middle Eocene of Sumatra)
  • Anhingidae – darters
    • Meganhinga chilensis Alvarenga 1995Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene)
    • Macranhinga Noriega 1992Lua error: Internal error: The interpreter has terminated with signal "24". (giant snake-birds) (Middle/Late Miocene –? Early Pliocene)
      • M. paranensis Noriega 1992Lua error: Internal error: The interpreter has terminated with signal "24". [Meganhinga paranensis (Noriega 1992)Lua error: Internal error: The interpreter has terminated with signal "24".]
      • M. ranzii Alvarenga & Guilherme 2003Lua error: Internal error: The interpreter has terminated with signal "24".
    • †"Paranavis" (Middle/Late Miocene) – a nomen nudum
    • Giganhinga kiyuensis Inderknecht & Noriega 2002Lua error: Internal error: The interpreter has terminated with signal "24". (Late Pliocene/Early Pleistocene)
    • Extant genera present in the fossil record
      • Anhinga (Early Miocene – Recent)
        • A. beckeri Emslie 1998Lua error: Internal error: The interpreter has terminated with signal "24".
        • A. hadarensis Brodkorb & Mourer-Chauviré 1982Lua error: Internal error: The interpreter has terminated with signal "24".
        • A. laticeps (De Vis 1906)Lua error: Internal error: The interpreter has terminated with signal "24". [Plotus laticeps De Vis 1906Lua error: Internal error: The interpreter has terminated with signal "24".]
        • A. malagurala Mackness 1995Lua error: Internal error: The interpreter has terminated with signal "24".
        • A. pannonica Lambrecht 1916Lua error: Internal error: The interpreter has terminated with signal "24".
        • A. subvolans (Brodkorb 1956) Becker 1986Lua error: Internal error: The interpreter has terminated with signal "24". [Phalacrocorax subvolans Brodkorb 1956Lua error: Internal error: The interpreter has terminated with signal "24".]
        • A. walterbolesi Worthy 2012Lua error: Internal error: The interpreter has terminated with signal "24".
        • A. fraileyi Campbell 1996Lua error: Internal error: The interpreter has terminated with signal "24". [?Macranhinga fraileyi (Campbell 1996)Lua error: Internal error: The interpreter has terminated with signal "24".]
        • A. grandis Martin & Mengel 1975Lua error: Internal error: The interpreter has terminated with signal "24".
        • A. minuta Alvarenga & Guilherme 2003Lua error: Internal error: The interpreter has terminated with signal "24".
  • Pelecanidae – pelicans
    • Protopelicanus cuvieri Reichenbach 1852Lua error: Internal error: The interpreter has terminated with signal "24".
    • Miopelecanus Cheneval 1984
      • M. gracilis (Milne-Edwards 1863) Cheneval 1984Lua error: Internal error: The interpreter has terminated with signal "24". [Pelecanus gracilis Milne-Edwards 1863Lua error: Internal error: The interpreter has terminated with signal "24".]
      • M. intermedius (Fraas 1870) Cheneval 1984Lua error: Internal error: The interpreter has terminated with signal "24". [Pelecanus intermedius Fraas 1870Lua error: Internal error: The interpreter has terminated with signal "24".]
    • Extant genera present in the fossil record
      • Pelecanus (Late Pliocene – Recent)
        • P. aethiopicus Harrison & Walker 1976Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. cautleyi Davies 1880Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. fraasi Lydekker 1891Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. halieus Wetmore 1933Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. odessanus Widhalm 1886Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. schreiberi Olson 1999Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. sivalensis Davies 1880Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. tirarensis Miller 1966Lua error: Internal error: The interpreter has terminated with signal "24".
Osteodontornis
  • Pelagornithidae Fürbringer 1888 – pseudotooth birds
    • Pseudodontornis Lambrecht 1930 [Neodontornis Harrison & Walker 1976; Macrodontopteryx Harrison & Walker 1976; Palaeochenoides Schufeldt 1916] (Late Paleocene – London Clay Early Eocene of England)
      • P. longidentata Harrison & Walker 1976 [Odontopteryx oweni (Harrison & Walker 1976); Macrodontopteryx oweni Harrison & Walker 1976]
      • P. tenuirostris Harrison 1985
      • P. tschulensis (Averianov et al. 1991) [Odontopteryx tschulensis]
      • P. longirostris (Spulski 1910) Lambrecht 1930 [Odontopteryx longirostris Spulski 1910; Palaeochenoides miocaenus Schufeldt 1916; Pelagornis longirostris (Spulski 1910)]
      • P. stirtoni Howard & Warter 1969 [Neodontornis stirtoni (Howard & Warter 1969) Harrison & Walker 1976; Pelagornis stirtoni (Howard & Warter 1969)]
    • Dasornis Owen 1870 [Megalornis Seeley 1866 non Gray 1841; Argillornis Owen 1878] (London Clay Early Eocene of England)
      • D. abdoun Bourdon 2010
      • D. emuinus (Bowerbank 1854) Owen 1870 [Lithornis emuinus Bowerbank 1854; Megalornis emuianus (sic) (Bowerbank 1854) Seeley 1866; Neptuniavis miranda Harrison & Walker 1977; Argillornis longipennis Owen 1878; Dasornis londinensis Owen 1869 nomen nudum; Dasornis londinensis Owen 1870; Argillornis emuinus (Bowerbank 1854) Brodkorb 1963; Argillornis longipes Lambrecht 1933 (lapsus)]
      • D. gigas [Odontopteryx gigas Bourdon 2006 nomen nudum]
    • Macrodontopteryx (London Clay Early Eocene of England)
    • Odontopteryx toliapica Owen 1873 [Odontornis (sic) Owen 1873; Neptuniavis minor Harrison & Walker 1977; Dasornis toliapica (Owen 1873)] (London Clay Early Eocene of England)
    • Gigantornis eaglesomei Andrews 1916 (middle Eocene of Nigeria)
    • Cyphornis magnus Cope 1894 (Eocene of Vancouver, Canada)
    • Osteodontornis orri Howard 1957 [Pelagornis orri Howard 1957] (Early Oligocene – Pliocene)
    • Pelagornis (Middle Miocene of France – Late Pliocene of Morocco)
      • P. miocaenus Lartet 1857 (Miocene false-tooothed pelican)
      • P. mauretanicus Mourer-Chauviré & Geraads 2008
      • P. chilensis Mayr & Rubilar-Rogers 2010
      • P. sandersi Ksepka 2014
    • Caspiodontornis kobystanicus Aslanova & Burčak-Abramovič 1982 [?Guguschia nailiae Aslanova & Burčak-Abramovič 1968]
    • Tympanoneisiotes wetmorei Hopson 1964 [Pelagornis wetmorei (Hopson 1964)]
    • Lutetodontopteryx tethyensis Mayr & Zvonok 2012
    • Aequornis traversei Bourdon 2006 nomen nudum
    • Placement unresolved
      • ?Pelagornithidae gen. et sp. indet. (middle Eocene)
  • Xenerodiopidae Rasmussen 1987Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene)
    • Xenerodiopus mycter Rasmussen, Olson & Simons 1987Lua error: Internal error: The interpreter has terminated with signal "24".
  • Ardeidae – herons
    • Ardeagrandis arborea Kurockin & Ganya 1972Lua error: Internal error: The interpreter has terminated with signal "24".
    • Calcardea junnei Gingerich 1987Lua error: Internal error: The interpreter has terminated with signal "24". (Paleocene)
    • Matuku otagoense Scofield, Worthy & Tennyson 2010Lua error: Internal error: The interpreter has terminated with signal "24".
    • Nyctisoma robusta Elżanowski & Zelenkov 2015Lua error: Internal error: The interpreter has terminated with signal "24".
    • Palaeophoyx columbiana McCoy 1963Lua error: Internal error: The interpreter has terminated with signal "24".
    • Pikaihao bartlei Worthy et al. 2013Lua error: Internal error: The interpreter has terminated with signal "24".
    • Proardea amissa (Milne-Edwards 1892) Lambrecht 1933Lua error: Internal error: The interpreter has terminated with signal "24". [Ardea amissa Milne-Edwards 1892Lua error: Internal error: The interpreter has terminated with signal "24".; Egretta amissa (Milne-Edwards 1892) Mlíkovský & Švec 1989Lua error: Internal error: The interpreter has terminated with signal "24".] (Late Eocene –? Late Oligocene)
    • Proardeola walkeri Harrison 1979)Lua error: Internal error: The interpreter has terminated with signal "24". [Ardea Formosa Milne-Edwards 1871 (nomen nudum)Lua error: Internal error: The interpreter has terminated with signal "24".; Ardeola walker (Harrison 1979) Mlíkovský & Švec 1989Lua error: Internal error: The interpreter has terminated with signal "24".] – possibly same as Proardea
    • Zeltornis ginsburgi Balouet 1981Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene)
    • Placement unresolved
      • Anas basaltica Bayer 1882 (Late Oligocene)
    • Extant genera present in the fossil record
      • Nyctanassa
        • N. kobdoena Kuročkin 1976Lua error: Internal error: The interpreter has terminated with signal "24".
      • Ardea (Middle Miocene – Recent)
        • A. aurelianensis Milne-Edwards 1871Lua error: Internal error: The interpreter has terminated with signal "24". [Proardea aurelianensis (Milne-Edwards 1871)Lua error: Internal error: The interpreter has terminated with signal "24".]
        • A. polkensis Brodkorb 1955Lua error: Internal error: The interpreter has terminated with signal "24".
        • A. howardae Brodkorb 1980Lua error: Internal error: The interpreter has terminated with signal "24".
        • A. paloccidentalis Shufeldt 1892Lua error: Internal error: The interpreter has terminated with signal "24".
        • A. sytchevskayae Zelenkov 2011Lua error: Internal error: The interpreter has terminated with signal "24".
      • Egretta (Late Miocene/Early Pliocene – Recent)
        • E. polgardiensis Kessler 2009Lua error: Internal error: The interpreter has terminated with signal "24".
        • E. subfluvia Becker 1985Lua error: Internal error: The interpreter has terminated with signal "24".
      • Butorides (Early Pleistocene – Recent)
        • B. validipes Campbell 1976Lua error: Internal error: The interpreter has terminated with signal "24".
      • Botaurus
        • B. hibbardi Moseley & Feduccia 1975Lua error: Internal error: The interpreter has terminated with signal "24".
      • Syrigma
        • S. sanctimartini Campbell 1979Lua error: Internal error: The interpreter has terminated with signal "24".
  • Scopidae – hammerkop
    • Prehistoric species of extant genera
      • Scopus xenopus Olson 1984Lua error: Internal error: The interpreter has terminated with signal "24".
  • Threskiornithidae – ibises
    • Threskiornithidae gen. et sp. indet. NMMP-KU 1301 (Pondaung middle Eocene of Paukkaung, Myanmar) – Threskiornithinae?
    • Gerandibis paganus (Milne-Edwards 1868) De Pietri 2013Lua error: Internal error: The interpreter has terminated with signal "24". [Milnea Lydekker 1891a non Reichenbach 1866Lua error: Internal error: The interpreter has terminated with signal "24".; Ibis pagana Milne-Edwards 1868Lua error: Internal error: The interpreter has terminated with signal "24".; Eudocimus paganus (Milne-Edvards 1868) Sharpe 1899Lua error: Internal error: The interpreter has terminated with signal "24".; Plegadis paganus (Milne-Edvards 1868) Olson 1981Lua error: Internal error: The interpreter has terminated with signal "24".; Actiornis pagana (Milne-Edwards 1868) Harrison 1976Lua error: Internal error: The interpreter has terminated with signal "24".; Milnea gracilis Lydekker 1891aLua error: Internal error: The interpreter has terminated with signal "24".; Plegadis gracilis (Lydekker 1891) Cheneval 1984 non Miller & Bowman 1956Lua error: Internal error: The interpreter has terminated with signal "24".]
    • Rhynchaeites messelensis Wittich 1898Lua error: Internal error: The interpreter has terminated with signal "24". [Plumumida Hoch 1980Lua error: Internal error: The interpreter has terminated with signal "24".; Plumumida lutetialis Hoch 1980Lua error: Internal error: The interpreter has terminated with signal "24".] (middle Eocene of Messel, Germany) – may include Mopsitta
    • Vadaravis brownae Smith, Grande & Clarke 2013Lua error: Internal error: The interpreter has terminated with signal "24".
    • Extant genera present in the fossil record
      • Geronticus (Middle Miocene – Recent)
        • G. perplexus (Milne-Edwards 1869) Cheneval 2000Lua error: Internal error: The interpreter has terminated with signal "24". [Ardea perplexa Milne-Edwards 1869Lua error: Internal error: The interpreter has terminated with signal "24".; Proardea perplexus (Milne-Edwards 1869) Gaillard 1939Lua error: Internal error: The interpreter has terminated with signal "24".]
        • G. apelex Olson 1985Lua error: Internal error: The interpreter has terminated with signal "24".
        • G. olsoni Mourer-Chauviré & Geraads 2010Lua error: Internal error: The interpreter has terminated with signal "24".
      • Platalea
        • P. chione (Emslie 1995)Lua error: Internal error: The interpreter has terminated with signal "24". [Ajaia chione Emslie 1995Lua error: Internal error: The interpreter has terminated with signal "24".]
        • P. tiangangensis Hou 1987Lua error: Internal error: The interpreter has terminated with signal "24". (Xiacaowan Middle Miocene of Sihong, China)
      • Plegadis
        • P. pharangites Olson 1981Lua error: Internal error: The interpreter has terminated with signal "24". [Plegadis gracilis Miller & Bowman 1956 non (Lydekker 1891)Lua error: Internal error: The interpreter has terminated with signal "24".] (Late Pliocene of WC US)
      • Theristicus
        • T. wetmorei Campbell 1979Lua error: Internal error: The interpreter has terminated with signal "24". (Late Pleistocene of Peru)
      • Eudocimus
        • E. leiseyi Emslie 1995aLua error: Internal error: The interpreter has terminated with signal "24". (Early Pleistocene of Florida)
        • E. peruvianus Campbell 1979Lua error: Internal error: The interpreter has terminated with signal "24". (Late Pleistocene of Peru; may be living species)
        • Eudocimus sp. (Middle Pliocene of Florida)
  • Balaenicipitidae – shoebills
    • Goliathia andrewsi Lambrecht 1930Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene/Early Oligocene of Egypt)
    • Paludiavis richae Harrison & Walker 1982Lua error: Internal error: The interpreter has terminated with signal "24". (Late Miocene of Tunisia and Pakistan)

The group that includes modern albatrosses, petrels and storm-petrels.

  • Placement unresolved
    • Eopuffinus kazachstanensis Nesov 1986Lua error: Internal error: The interpreter has terminated with signal "24". (Late Paleocene of Zhylga, Kazakhstan)
    • Makahala mirae Mayr 2015Lua error: Internal error: The interpreter has terminated with signal "24".
    • Proaestrelata Imber 1985Lua error: Internal error: The interpreter has terminated with signal "24".
  • Marinavidae Harrison & Walker 1977Lua error: Internal error: The interpreter has terminated with signal "24".
    • Marinavis longirostris Harrison & Walker 1977 Lua error: Internal error: The interpreter has terminated with signal "24".
  • Tytthostonychidae Olson & Parris 1987Lua error: Internal error: The interpreter has terminated with signal "24".
  • Diomedeoididae Fischer 1985Lua error: Internal error: The interpreter has terminated with signal "24".
    • Rupelornis van Beneden 1871 [Gaviota Fischer 1983 non Miller & Sibley 1941Lua error: Internal error: The interpreter has terminated with signal "24".; Ardeita Cheneval 1996Lua error: Internal error: The interpreter has terminated with signal "24".; Diomedeoides Fischer 1985Lua error: Internal error: The interpreter has terminated with signal "24".; Frigidafrons Cheneval 1995Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Oligocene– Early Miocene of C Europe and Iran)
      • R. harmati (Kessler 2009) Mayr & Smith 2012Lua error: Internal error: The interpreter has terminated with signal "24". [Diomedeoides harmati Kessler 2009Lua error: Internal error: The interpreter has terminated with signal "24".]
      • R. definitus van Beneden 1871Lua error: Internal error: The interpreter has terminated with signal "24". [Vanellus selysii van Beneden 1871Lua error: Internal error: The interpreter has terminated with signal "24".; Anas creccoides van Beneden 1871Lua error: Internal error: The interpreter has terminated with signal "24".; Gaviota lipsiensis Fischer 1983Lua error: Internal error: The interpreter has terminated with signal "24".; Diomedeoides minimus Fischer 1985Lua error: Internal error: The interpreter has terminated with signal "24".; Frigidafrons brodkorbi Cheneval 1995Lua error: Internal error: The interpreter has terminated with signal "24".; Ardeita gracilis Cheneval 1996 (nomen nudum)Lua error: Internal error: The interpreter has terminated with signal "24".; Diomedeoides lipsiensis (Fischer 1983) Mayr et al. 2002Lua error: Internal error: The interpreter has terminated with signal "24".; Diomedeoides brodkorbi (Cheneval 1995) Fischer 2003Lua error: Internal error: The interpreter has terminated with signal "24".]
      • R. babaheydariensis (Peters & Hamedani 2000) Mayr & Smith 2012Lua error: Internal error: The interpreter has terminated with signal "24". [Frigidafons babaheydariensis Peters & Hamedani 2000Lua error: Internal error: The interpreter has terminated with signal "24".; Diomedeoides babaheydariensis (Peters & Hamedani 2000) Mayr, Peters & Rietschel 2002Lua error: Internal error: The interpreter has terminated with signal "24".]
  • Oceanitidae
    • Extant genera present in the fossil record
  • Diomedeidae – albatrosses
    • Murunkus subitus Panteleyev & Nesov 1993Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene)
    • Plotornis Milne-Edwards 1874Lua error: Internal error: The interpreter has terminated with signal "24". [Chenornis Portis 1884Lua error: Internal error: The interpreter has terminated with signal "24".] (Early – Middle Miocene)
      • P. arvernensis (Milne-Edwards 1871) Cheneval 1984aLua error: Internal error: The interpreter has terminated with signal "24". [Puffinus arvernensis Milne-Edwards 1871 nomen nudumLua error: Internal error: The interpreter has terminated with signal "24".; Puffinus arvernensis Shufeldt 1896Lua error: Internal error: The interpreter has terminated with signal "24".]
      • P. deltortrii Milne-Edwards 1874Lua error: Internal error: The interpreter has terminated with signal "24".
      • P. graculoides (Portis 1884) Mayr & Pavia 2014Lua error: Internal error: The interpreter has terminated with signal "24". [Chenornis graculoides Portis 1884Lua error: Internal error: The interpreter has terminated with signal "24".]
    • Tydea septentrionalis Mayr & Smith 2012Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene of the North Sea basin)
    • †Diomedeidae gen. et sp. indet. (Late Oligocene of South Carolina)
    • Extant genera present in the fossil record
      • Diomedea (Middle Miocene – Recent)
        • D. anglica Lydekker 1891aLua error: Internal error: The interpreter has terminated with signal "24".
        • D. howardae Chandler 1990Lua error: Internal error: The interpreter has terminated with signal "24".
        • D. milleri Howard 1966Lua error: Internal error: The interpreter has terminated with signal "24".
        • D. rumana Grigorescu & Kessler 1988Lua error: Internal error: The interpreter has terminated with signal "24".
        • D. tanakai Davis 2003 Lua error: Internal error: The interpreter has terminated with signal "24".
        • D. thyridata Wilkinson 1969Lua error: Internal error: The interpreter has terminated with signal "24".
      • Phoebastria (Middle Miocene – Recent)
        • P. rexsularum Olson & Rasmussen 2001Lua error: Internal error: The interpreter has terminated with signal "24".
      • Thalassarche (Late Miocene – Recent)
  • Hydrobatidae – storm-petrels
    • Primodroma bournei Harrison & Walker 1977Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene)
    • Prehistoric species of extant genera
      • "Oceanodroma" hubbsi Miller 1951Lua error: Internal error: The interpreter has terminated with signal "24". (Hubbs' Storm-petrel)(Capistrano Middle/Late Miocene of Orange County, US)
      • Pelagodroma sp. 1
      • Pelagodroma sp. 2
  • Procellariidae – petrels
    • Argyrodyptes microtarsus Ameghino 1905Lua error: Internal error: The interpreter has terminated with signal "24". (San Julián Late Eocene/Early Oligocene of Chubut, Argentina)
    • Pterodromoides minoricensis Sequí et al. 2001Lua error: Internal error: The interpreter has terminated with signal "24".
    • Extant genera present in the fossil record
      • Calonectris (Early Pliocene – Recent)
        • C. krantzi Olson & Rasmussen 2001 Lua error: Internal error: The interpreter has terminated with signal "24".
        • C. kurodai Olson 2009Lua error: Internal error: The interpreter has terminated with signal "24".
        • C. wingatei Olson 2008Lua error: Internal error: The interpreter has terminated with signal "24".
      • Fulmarus (Middle Miocene – Recent)
        • F. hammeri Howard 1968Lua error: Internal error: The interpreter has terminated with signal "24".
        • F. miocaenus Howard 1984Lua error: Internal error: The interpreter has terminated with signal "24".
      • Pachyptila (Late Miocene– Recent)
        • P. salax Olson 1985Lua error: Internal error: The interpreter has terminated with signal "24".
      • Pelecanoides (Early/Middle Miocene – Recent)
        • P. cymatotrypetes Olson 1985Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. miokuaka Worthy et al. 2007Lua error: Internal error: The interpreter has terminated with signal "24". (Miocene diving petrel)
        • P. urinatrix Gmelin 1789Lua error: Internal error: The interpreter has terminated with signal "24".
      • "Pterodroma" (Pleistocene – Recent)
        • P. kurodai Harrison & Walker 1978Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. rupinarum Olson 1975Lua error: Internal error: The interpreter has terminated with signal "24".
      • Puffinus (Early Oligocene – Recent)
        • P. aquitanicus (Milne-Edwards) Brodkorb 1963Lua error: Internal error: The interpreter has terminated with signal "24". Procellaria aquitanica Milne-Edwards 1874Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. antiquus (Milne-Edwards) Brodkorb 1963cLua error: Internal error: The interpreter has terminated with signal "24". Procellaria antiqua Milne-Edwards 1874Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. barnesi Howard 1978Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. calhouni Howard 1968Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. conradi Marsh 1870Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. diatomicus Miller 1925Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. felthami Howard 1949Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. gilmorei Chandler 1990Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. inceptor Wetmore 1930Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. kanakoffi Howard 1949Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. micraulax Brodkorb 1963Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. mitchelli Miller 1961Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. nestori Alcover 1989Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. priscus Miller 1961Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. raemdonckii (Van Beneden 1871) Brodkorb 1962Lua error: Internal error: The interpreter has terminated with signal "24". [Larus Raemdonckii Beneden 1871]
        • P. tedfordi Howard 1971Lua error: Internal error: The interpreter has terminated with signal "24".
Waimanu
Icadyptes
  • Unresolved and basal forms
    • Waimanu Jones, Ando & Fordyce 2006 (Early – Late Palaeocene)
      • W. manneringi Jones, Ando & Fordyce 2006 (Mannering's penguin)
      • W. tuatahi Jones, Ando & Fordyce 2006 (Waipara penguin)
    • Perudyptes devriesi Clarke et al. 2007 – basal? (middle Eocene)
    • Sphenisciformes gen. et sp. indet. CADIC P 21 (middle Eocene)
  • Spheniscidae – penguins
    • Kaiika maxwelli Fordyce & Tomas 2011
    • Palaeoapterodytes ictus (Ameghino 1891) Ameghino 1905 (nomen dubium) [Apterodytes Ameghino 1891 non Hermann 1783; Apterodytes ictus Ameghino 1891]
    • Tereingaornis moisleyi Scarlett 1983 (Moisley's penguin)
    • Crossvallia unienwillia Tambussi et al. 2005 (Late Paleocene)
    • Anthropornis Wiman 1905 (mid-Eocene –? Early Oligocene)
      • A. nordenskjoeldii Wiman 1905 (Nordenskjoeld's giant penguin)
      • A. grandis (Wiman 1905) [Arthrodytes grandis (Ameghino 1901) Ameghino 1905]
    • Archaeospheniscus Marples 1952 (mid-/Late Eocene – Late Oligocene)
    • Delphinornis Wiman 1905 [Notodyptes Marples 1953] (mid-/Late Eocene –? Early Oligocene)
      • D. gracilis Myrcha et al. 2002
      • D. larseni Wiman 1905
      • D. arctowskii Myrcha et al. 2002
      • D. wimani (Marples 1953) Ksepka & Clarke 2010 [Notodyptes wimani Marples 1953; Archaeospheniscus wimani (Marples 1953) Simpson 1971a]
    • Palaeeudyptes Huxley 1859 [Eosphaeniscus Wiman 1905] (mid-/Late Eocene – Late Oligocene)
    • Icadyptes salasi Clarke et al. 2007 (Late Eocene)
    • Inkayacu paracasensis Clarke et al. 2010 (Late Eocene)
    • Pachydyptes ponderosus Oliver 1930 (Late Eocene)
    • Marambiornis exilis Myrcha et al. 2002 (Late Eocene–? Early Oligocene)
    • Mesetaornis polaris Myrcha et al. 2002 (Late Eocene –? Early Oligocene)
    • Tonniornis Tambussi et al. 2006 (Late Eocene –? Early Oligocene)
      • T. mesetaensis Tambussi et al. 2006
      • T. minimum Tambussi et al. 2006
    • Wimanornis seymourensis Simpson 1971 (Late Eocene –? Early Oligocene)
    • Arthrodytes andrewsi Ameghino 1901 (Late Eocene/Early Oligocene – Early Miocene)
    • Duntroonornis parvus Marples 1953 (Duntroon penguin) (Late Oligocene)
    • Kairuku Ksepka et al. 2012 (Late Oligocene)
      • K. grebneffi Ksepka et al. 2012
      • K. waitaki Ksepka et al. 2012
    • Korora oliveri Marples 1952 (Oliver's penguin) (Late Oligocene)
    • Platydyptes Marples 1952 (Late Oligocene)
      • P. marplesi (Brodkorb 1963) Simpson 1971 [Palaeeudyptes marplesi Brodkorb 1963] (Simpson's/Marples' penguin)
      • P. novaezealandiae(Oliver 1930) Marples 1952 (wide-flippered penguin)
      • P. amiesi Marples 1952 (Amies' penguin)
    • Eretiscus tonnii (Simpson 1981) Olson 1986 [Microdytes Simpson 1981 non Balfour-Browne 1949; Microdytes tonnii Simpson 1981] (Patagonia Early Miocene)
    • Palaeospheniscus Moreno & Mercerat 1891 [Neculus Ameghino 1905; Paraspheniscus Ameghino 1905 nec Hendel 1927; Perispheniscus Ameghino 1905; Pseudospheniscus Ameghino 1905; Chubutodyptes Simpson 1970] (Early? – Late Miocene/Early Pliocene)
      • P. huxleyorum Simpson 1973
      • P. wimani (Ameghino 1905) [Perispheniscus wimani Ameghino 1905; Palaeospheniscus robustus (partim) Simpson 1946]
      • P. gracilis Ameghino 1899 [?Delphinornis gracilis; Palaeospheniscus nereius Ameghino 1901; Neculus rothi Ameghino 1905; Palaeospheniscus rothi Ameghino 1905; Paraspheniscus nereius (Ameghino 1901) Ameghino 1905; Palaeospheniscus medianus Ameghino 1905]
      • P. patagonicus Moreno & Mercerat 1891 [Palaeospheniscus menzbieri Moreno & Mercerat 1891; Palaeospheniscus interruptus Ameghino 1905; Palaeospheniscus intermedius Ameghino 1905; Palaeospheniscus affinis Ameghino 1905]
      • P. bergi Moreno & Mercerat 1891 [Paraspheniscus bergi (Moreno & Mercerat 1891) Ameghino 1905; Palaeospheniscus planus Ameghino 1905; Palaeospheniscus interplanus Ameghino 1905; Pseudospheniscus interplanus Ameghino 1905; Pseudospheniscus planus Ameghino 1905 nom rej.; Pseudospheniscus concavus Ameghino 1905; Pseudospheniscus convexus Ameghino 1905 nom rej.]
      • P. biloculata (Simpson 1970) Acosta Hospitaleche 2007 [Chubutodyptes biloculata Simpson 1970]
    • Paraptenodytes Ameghino 1891 [Isotremornis Ameghino 1905; Metancylornis Ameghino 1905; Treleudytes Ameghino 1905] (Early – Late Miocene/Early Pliocene)
      • P. brodkorbi Simpson 1972 [Isotremornis nordenskjöldi Ameghino 1905 (partim)]
      • P. robustus (Ameghino 1895) [Palaeospheniscus robustus Ameghino 1895; Perispheniscus robustus (Ameghino 1895) Brodkorb 1963; Paraptenodytes curtus Ameghino 1901; Metancylornis curtus (Ameghino 1901) Ameghino 1905; Paraptenodytes grandis Ameghino 1901; Arthrodytes grandis (Ameghino 1901) Ameghino 1905; Paraptenodytes andrewsi Ameghino 1901; Arthrodytes andrewsi (Ameghino 1901) Ameghino 1905; Treleudytes crassus Ameghino 1905]
      • P. antarcticus (Huxley 1859) Ameghino 1891 [Palaeeudyptes antarcticus Huxley 1859 non Hector 1873; Palaeospheniscus antarcticus (Huxley 1859) Moreno & Mercerat 1891] (narrow-flippered penguin)
    • Anthropodyptes gilli Simpson 1959 (Middle Miocene)
    • Madrynornis mirandus Hospitaleche et al. 2007 (Late Miocene)
    • Pseudaptenodytes Simpson 1970 (Late Miocene/Early Pliocene)
    • Dege hendeyi Simpson 1979 (Early Pliocene)
    • Marplesornis novaezealandiae (Marples 1960) Simpson 1972 [Paleospheniscus novaezealandiae Marples 1960] (Harris penguins) (Early Pliocene)
    • Nucleornis insolitus Simpson 1979 (Early Pliocene)
    • Inguza predemersus (Simpson 1971) Simpson 1975 [Spheniscus predemersus Simpson 1971] (Late Pliocene)
    • Placement unresolved
      • Spheniscidae gen. et sp. indet (Late Oligocene/Early Miocene of Hakataramea, New Zealand)
    • Extant genera present in the fossil record
      • Pygoscelis (Middle/Late Miocene – Recent)
      • Spheniscus (Middle/Late Miocene – Recent)
        • S. muizoni Göhlich 2007
        • S. chilensis Emslie & Correa 2003
        • S. megaramphus Stucchi, Urbina & Giraldo 2003
        • S. urbinai Stucchi 2002
      • Aptenodytes (Early? Pliocene – Recent)
        • A. ridgeni Simpson 1972 [Arthrodytes ridgeni (Simpson 1972)] (Ridgen's Penguin)
      • Eudyptes

Pterocliformes

  • Pteroclidae – sandgrouse
    • Gerandia calcaria (Milne-Edwards 1869) Lambrecht 1933 [Columba calcaria Milne-Edwards 1869] (Early Miocene)
    • Leptoganga sepultus (Milne-Edwards 1869) Mourer-Chauviré 1993 [Pterocles sepultus Milne-Edwards 1869]
    • Archaeoganga Mourer-Chauviré 1992
      • A. validus(Milne-Edwards 1892) [Pterocles validus Milne-Edwards 1892]
      • A. pinguis Mourer-Chauviré 1992
      • A. larvatus (Milne-Edwards 1892) [Pterocles larvatus Milne-Edwards 1892]
    • Extant genera present in the fossil record

Columbiformes

  • Columbidae – doves and pigeons
    • Arenicolumba (Early Miocene) – doubtfully distinct from Patagioenas
    • Rupephaps (Early Miocene)
    • Placement unresolved
      • Columbidae gen. et sp. indet. (Early/Middle Miocene)
    • Extant genera present in the fossil record

Unresolved and basal fossil parrots:

  • Psittacopes (Early/middle Eocene)
  • Serudaptus – pseudasturid or psittacid?
  • Pseudasturidae (= Halcyornithidae?)
  • Messelasturidae Mayr 2005Lua error: Internal error: The interpreter has terminated with signal "24".
    • Tynskya eocaena Mayr 2000Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene of N America and England)
    • Messelastur gratulator Peters 1994Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene of Messel, Germany)
    • Pseudasturidae FU 125 gen. et sp. indet. (Early Eocene)
    • Pseudasturides – formerly Pseudastur
  • Vastanavidae
  • Quercypsittidae
  • Cacatuidae
    • Extant genera present in the fossil record
      • Cacatua (Early Miocene – Recent)
  • Psittacidae – parrots, parakeets and lories
    • Archaeopsittacus (Late Oligocene/Early Miocene)
    • Xenopsitta (Early Miocene)
    • Psittacidae gen. et spp. indet. (Early/Middle Miocene) – several species
    • Bavaripsitta (Middle Miocene)
    • Psittacidae gen. et sp. indet. (Middle Miocene) – erroneously placed in Pararallus dispar, includes "Psittacus lartetianus
    • Extant and recently extinct genera present in the fossil record
  • Strigopidae – New Zealand parrots, kakapo

Hoatzins

  • Opisthocomidae Swainson 1837Lua error: Internal error: The interpreter has terminated with signal "24". [Foratidae Olson 1992Lua error: Internal error: The interpreter has terminated with signal "24".; Onychopterygidae Cracraft 1971Lua error: Internal error: The interpreter has terminated with signal "24".]
    • Foro panarium Olson 1992Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene) - cuculiform?
    • Hoatzi panarium Thomas 1996Lua error: Internal error: The interpreter has terminated with signal "24".
    • Onychopteryx simpsoni Cracraft 1971Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene of Argentina) – falconid? A nomen dubium
    • Protoazin parisiensis Mayr & De Pietri 2014Lua error: Internal error: The interpreter has terminated with signal "24".
    • Namibiavis senutae Mourer-Chauviré 2003Lua error: Internal error: The interpreter has terminated with signal "24". (Late Oligocene of Namibia)
    • Hoazinavis lacustris Alvarenga, Mayr & Mourer-Chauviré 2011Lua error: Internal error: The interpreter has terminated with signal "24". (Miocene of Brazil)
    • Hoazinoides magdalenae Miller 1953Lua error: Internal error: The interpreter has terminated with signal "24". (Late Miocene of Colombia)
  • Musophagidae – turacos
    • Placement unresolved
      • Musophagidae gen. et sp. indet. (Late Oligocene – Middle Miocene of WC Europe)
      • Musophagidae gen. et sp. indet. (Egypt)
    • Veflintornis Kashin 1976Lua error: Internal error: The interpreter has terminated with signal "24". [Apopempsis Brodkorb 1971 non Schenkling 1903Lua error: Internal error: The interpreter has terminated with signal "24".] (Middle Miocene)
      • V. meini (Ballmann 1969)Lua error: Internal error: The interpreter has terminated with signal "24". [Musophaga meini Ballmann 1969Lua error: Internal error: The interpreter has terminated with signal "24".]
      • V. africanus (Harrison 1980)Lua error: Internal error: The interpreter has terminated with signal "24". [Musophaga africanus Harrison 1980Lua error: Internal error: The interpreter has terminated with signal "24".; Apopempsis africanus (Harrison 1980)Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Miocene)

Cuckoos, turacos and allies.

  • Placement unresolved
    • Cuculiformes gen. et sp. indet. (Early Eocene)
  • Cuculidae – cuckoos
    • Eocuculus (Late Eocene)
    • Dynamopterus (Late Eocene/Early Oligocene)
    • Neococcyx (Early Oligocene)
    • Cursoricoccyx (Early Miocene)
    • Placement unresolved
      • Cuculidae gen. et sp. indet. (Early Pliocene)
Teratornis
  • Teratornithidae Miller 1909Lua error: Internal error: The interpreter has terminated with signal "24".– teratorns
    • Argentavis magnificens Campbell & Tonni 1980Lua error: Internal error: The interpreter has terminated with signal "24". (Late Miocene)
    • Aiolornis incredibilis (Howard 1952) Campbell, Scott & Springer 1999Lua error: Internal error: The interpreter has terminated with signal "24". [Teratornis incredibilis Howard 1952Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Pliocene – Late Pleistocene)
    • Oscaravis olsoni (Arredondo & Arredondo 2002) Suarez & Olson 2009Lua error: Internal error: The interpreter has terminated with signal "24". [Teratornis olsoni Arredondo & Arredondo 2002Lua error: Internal error: The interpreter has terminated with signal "24".] (Cuban teratorn)(Pleistocene)
    • Cathartornis gracilis Miller 1910Lua error: Internal error: The interpreter has terminated with signal "24".
    • Taubatornis campbelli Olson & Alvarenga 2002Lua error: Internal error: The interpreter has terminated with signal "24".
    • Teratornis Miller 1909Lua error: Internal error: The interpreter has terminated with signal "24". (Early Pleistocene – Late Pleistocene)
      • T. woodburnensis Campbell & Stenger 2002Lua error: Internal error: The interpreter has terminated with signal "24".
      • T. merriami Miller 1909Lua error: Internal error: The interpreter has terminated with signal "24". (Merriam's teratorn)
  • Cathartidae – New World vultures
    • Brasilogyps faustoi Alvarenga 1985Lua error: Internal error: The interpreter has terminated with signal "24". (Late Oligocene – Early Miocene)
    • Kuntur cardenasi Stucchi et al. 2015Lua error: Internal error: The interpreter has terminated with signal "24".
    • Pleistovultur nevesi Alvarenga et al. 2008Lua error: Internal error: The interpreter has terminated with signal "24".
    • Tapinopus ellioti Milne-Edwards 1891Lua error: Internal error: The interpreter has terminated with signal "24".
    • Diatropornis ellioti (European vulture) (Late Eocene/Early Oligocene –? middle Oligocene)
    • Phasmagyps patritus Wetmore 1927Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene)
    • Hadrogyps aigialeus Emslie 1988Lua error: Internal error: The interpreter has terminated with signal "24". (American dwarf vulture) (Middle Miocene)
    • Pliogyps Becker 1986Lua error: Internal error: The interpreter has terminated with signal "24". (Miocene vulture) (Late Miocene – Late Pliocene)
      • P. charon Becker 1986Lua error: Internal error: The interpreter has terminated with signal "24".
      • P. fisheri Tordoff 1959Lua error: Internal error: The interpreter has terminated with signal "24".
    • Perugyps diazi Stucchi & Emslie 2005Lua error: Internal error: The interpreter has terminated with signal "24". (Peruvian vulture) (Late Miocene/Early Pliocene)
    • Dryornis (Argentinean vulture) Moreno & Mercerat, 1891Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene – Late Pliocene) – similar to the extant genus Vultur
      • D. pampeanus Moreno & Mercerat, 1891Lua error: Internal error: The interpreter has terminated with signal "24".
      • D. hatcheri Degrange 2022Lua error: Internal error: The interpreter has terminated with signal "24".
    • Aizenogyps toomeyae Emslie 1998Lua error: Internal error: The interpreter has terminated with signal "24". (South American vulture) (Late Pliocene)
    • Breagyps clarki Miller 1938Lua error: Internal error: The interpreter has terminated with signal "24". (long-legged vulture) (Late Pleistocene)
    • Geronogyps reliquus Campbell 1979Lua error: Internal error: The interpreter has terminated with signal "24". (Late Pleistocene)
    • Wingegyps cartellei Alvarenga & Olson 2004Lua error: Internal error: The interpreter has terminated with signal "24". (Amazonian vulture) (Late Pleistocene)
    • Parasarcoramphus milneedwardsi Mourer-Chauviré 2002Lua error: Internal error: The interpreter has terminated with signal "24".
    • Placement unresolved
      • Cathartidae gen. et sp. indet. (Late Oligocene of Mongolia)
      • Cathartidae gen. et sp. indet. (Late Miocene/Early Pliocene of Lee Creek Mine, US)
      • Cathartidae gen. et sp. indet. (Middle Pliocene of Argentina)
      • Cathartidae gen. et sp. indet. (Cuba)
    • Extant genera present in the fossil record
      • Sarcoramphus (Middle Pliocene –? Recent)
        • S. kernense' (Kern vulture)
        • S. fischeri Campbell 1979Lua error: Internal error: The interpreter has terminated with signal "24".
      • Gymnogyps (Early Pleistocene – Recent)
        • G. howardae' Campbell 1979Lua error: Internal error: The interpreter has terminated with signal "24".
        • G. kofordi' Emslie 1988Lua error: Internal error: The interpreter has terminated with signal "24".
      • Vultur (Pliocene – Recent)– distinctiveness disputed
        • V. fossilis Moreno & Mercerat 1891Lua error: Internal error: The interpreter has terminated with signal "24". [Cathartes fossilis; Sarcoramphus fossilis]
  • Horusornithidae Mourer-Chauviré 1991Lua error: Internal error: The interpreter has terminated with signal "24".
    • Horusornis vianeyliaudae Mourer-Chauviré 1991Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene)
  • Pandionidae – ospreys
    • Extant genera present in the fossil record
      • Pandion (Early Oligocene – Recent)
        • P. lovensis Becker 1985Lua error: Internal error: The interpreter has terminated with signal "24".
        • P. homalopteron Warter 1976Lua error: Internal error: The interpreter has terminated with signal "24".
  • Sagittariidae – secretarybirds
    • Amanuensis pickfordi Mourer-Chauviré 2003Lua error: Internal error: The interpreter has terminated with signal "24".
    • Pelargopappus Stejneger 1885Lua error: Internal error: The interpreter has terminated with signal "24".(Late Eocene/Early Oligocene – Late Oligocene/Early Miocene of France) – formerly Amphiserpentarius/Amynoptilon/Pelargopsis
      • P. schlosseri
      • P. magnus
  • Accipitridae – hawks, eagles and Old World vultures
    • Milvoides (Late Eocene)
    • Aquilavus (Late Eocene/Early Oligocene – Early Miocene)
    • Palaeocircus (Late Eocene/Early Oligocene)
    • Palaeastur (Early Miocene)
    • Pengana (Early Miocene)
    • Promilio (Early Miocene)
    • Proictinia (Early – Late Miocene/Early Pliocene)
    • Neophrontops (Early/middle Miocene – Late Pleistocene) – formerly in Neophron
    • Mioaegypius (middle Miocene)
    • Apatosagittarius (Late Miocene)
    • Gansugyps (Late Miocene)
    • Palaeoborus (Miocene)
    • Qiluornis (Miocene)
    • Thegornis (Miocene)
    • Garganoaetus (Early Pliocene)
    • Amplibuteo (Late Pliocene of Peru – Late Pleistocene) – Belongs to the extant genus Buteogallus[19]
    • Cryptogyps (Middle – Late Pleistocene)
    • Neogyps (Late Pleistocene)
    • Palaeohierax – includes "Aquila" gervaisii
    • Placement unresolved
      • Accipitridae gen. et sp. indet. AMNH FR 7434 (Early Eocene)
      • Accipitridae gen. et sp. indet. (Early Oligocene)
      • Accipitridae gen. et sp. indet. (Early/Middle Miocene)
      • Accipitridae gen. et sp. indet. MPEF-PV-2523 (Late Miocene)
      • Accipitridae gen. et sp. indet. (Early/Middle Pliocene) – Parabuteo?
      • Accipitridae gen. et sp. indet. (Late Pliocene/Early Pleistocene) – Buteo?
      • Accipitridae gen. et sp. indet. (Egypt)
      • "Aquila" danana (Late Miocene/Early Pliocene) – formerly also Geranoaetus or Buteo
    • Extant genera present in the fossil record
      • Haliaeetus (Early Oligocene – Recent)
      • Buteo (middle Oligocene – Recent)
      • Aquila (Middle Miocene – Recent)
      • Buteogallus (Middle Miocene – Recent) – might include Harpyhaliaetus
      • "Hieraaetus" (Middle Miocene – Recent) – doubtfully distinct from Aquila
      • Milvus (Early Pleistocene – Recent)
      • Gyps (Middle Pleistocene – Recent)
      • Aegypius (Middle Pleistocene – Recent)
    • Additional prehistoric species of extant genera
      • Spizaetus grinnelli (Rancho La Brea Late Pleistocene of California, US) – formerly Geranoaetus or Buteo
      • Spizaetus pliogryps – formerly Aquila
      • Gypaetus georgii (Late Miocene)
      • Neophron lolis (Late Miocene)
  • Masillaraptoridae Mayr 2009Lua error: Internal error: The interpreter has terminated with signal "24".
    • Masillaraptor parvunguis Mayr 2006Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene of Messel, Germany)
    • Danielsraptor phorusrhacoides Mayr & Kitchener 2022Lua error: Internal error: The interpreter has terminated with signal "24". (early Eocene of Essex, England)
  • Falconidae – falcons
    • Lagopterus minutus Moreno & Mercerat 1891Lua error: Internal error: The interpreter has terminated with signal "24".
    • Parvulivenator watteli Harrison 1982Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene)
    • Stintonornis mitchelli Harrison 1984Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene)
    • Badiostes patagonicus Ameghino 1895Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene)
    • Pediohierax ramenta (Wetmore 1936) Becker 1987Lua error: Internal error: The interpreter has terminated with signal "24". [Falco ramenta Wetmore 1936Lua error: Internal error: The interpreter has terminated with signal "24".] - (Middle Miocene)
    • Thegornis musculosus Ameghino 1895Lua error: Internal error: The interpreter has terminated with signal "24". [Thegornis debilis Ameghino 1895Lua error: Internal error: The interpreter has terminated with signal "24".; Buteo musculosus (Ameghino 1895) Agnolin 2006Lua error: Internal error: The interpreter has terminated with signal "24".]
      • Petrosushkinia pliocaena (Tugarinov 1935) Zelenkov 2015Lua error: Internal error: The interpreter has terminated with signal "24". [Sushkinia Tugarinov 1935 non Martynov 1930Lua error: Internal error: The interpreter has terminated with signal "24".; Sushkinia pliocaena Tugarinov 1935Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Pliocene)
      • †Falconidae gen. et sp. indet. (Late Miocene)
    • Extant genera present in the fossil record
      • Milvago (Late Pleistocene -–Recent)
        • M. alexandri Olson 1976Lua error: Internal error: The interpreter has terminated with signal "24".
        • M. brodkorbi Campbell 1979Lua error: Internal error: The interpreter has terminated with signal "24".
      • Falco (Late Miocene – Recent)
        • F. antiquus Mourer-Chauviré 1975Lua error: Internal error: The interpreter has terminated with signal "24".
        • F. bakalovi Boev 1999Lua error: Internal error: The interpreter has terminated with signal "24".
        • F. bulgaricus Boev 2011Lua error: Internal error: The interpreter has terminated with signal "24".
        • F. chowi Hou 1982Lua error: Internal error: The interpreter has terminated with signal "24".
        • F. hezhengensis Li et al. 2014Lua error: Internal error: The interpreter has terminated with signal "24".
        • F. medius Umanskaya 1981Lua error: Internal error: The interpreter has terminated with signal "24".
        • F. oregonus Brodkorb 1964Lua error: Internal error: The interpreter has terminated with signal "24".
        • F. pisanus Portis 1887Lua error: Internal error: The interpreter has terminated with signal "24".
        • F. umanskajae Sobolev 2003Lua error: Internal error: The interpreter has terminated with signal "24".
  • Paraprefica Mayr 1999Lua error: Internal error: The interpreter has terminated with signal "24". (Early Eocene?)
    • P. kelleri Mayr 1999Lua error: Internal error: The interpreter has terminated with signal "24".
    • P. major Mayr 1999Lua error: Internal error: The interpreter has terminated with signal "24".
  • Steatornithidae – oilbirds
    • Prefica nivea Olson 1987Lua error: Internal error: The interpreter has terminated with signal "24".
    • Prehistoric species of extant genera
  • Nyctibiidae – potoos
    • Euronyctibius kurochkini Mourer-Chauviré 1989Lua error: Internal error: The interpreter has terminated with signal "24".
  • Fluvioviridavidae Mayr 2005Lua error: Internal error: The interpreter has terminated with signal "24".
    • Fluvioviridavis platyrhamphus Mayr & Daniels 2001Lua error: Internal error: The interpreter has terminated with signal "24". (Green River Early Eocene of N America)
  • Podargidae – frogmouths
    • Masillapodargus longipes Mayr 1999Lua error: Internal error: The interpreter has terminated with signal "24".
    • Quercypodargus olsoni Mourer-Chauvire 1989Lua error: Internal error: The interpreter has terminated with signal "24".
  • Caprimulgidae – nightjars
    • Ventivorus ragei Mourer-Chauviré 1988Lua error: Internal error: The interpreter has terminated with signal "24".
    • Prehistoric species of extant genera
      • Caprimulgus
        • C. piurensis Campbell 1979Lua error: Internal error: The interpreter has terminated with signal "24".

Owlet-nightjars

  • Aegothelidae
    • Quipollornis koniberi Rich & McEvey 1977Lua error: Internal error: The interpreter has terminated with signal "24". (Early/Middle Miocene)
    • Extant genera present in the fossil record
      • Aegotheles (Early/Middle Miocene – Recent)
        • A. savesi Layard & Layard 1881Lua error: Internal error: The interpreter has terminated with signal "24". (New Caledonian owlet-nightjar)
        • A. zealandivetus Worthy et al., 2022Lua error: Internal error: The interpreter has terminated with signal "24".

Swifts and hummingbirds.

  • Wyomingcypselus pohli Weidig 2003 nomen nudumLua error: Internal error: The interpreter has terminated with signal "24".
  • Procypseloides mourerchauvireae (Mlíkovský) Harrison & Walker 1977Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene/Early Oligocene – Early Miocene)
  • Laputavis robusta (Dyke 2001a) Dyke 2001bLua error: Internal error: The interpreter has terminated with signal "24". [Laputa Dyke 2001a nonLua error: Internal error: The interpreter has terminated with signal "24".; Laputa robusta Dyke 2001aLua error: Internal error: The interpreter has terminated with signal "24".]
  • Palescyvus escampensis Karhu 1988Lua error: Internal error: The interpreter has terminated with signal "24".
  • Scaniacypselus Harrison 1984Lua error: Internal error: The interpreter has terminated with signal "24". (Early – middle Eocene)
    • S. wardi Harrison 1984Lua error: Internal error: The interpreter has terminated with signal "24".
    • S. szarskii (Peters 1985) Meyr & Peters 1999Lua error: Internal error: The interpreter has terminated with signal "24". [Aegialornis szarskii Peters 1985Lua error: Internal error: The interpreter has terminated with signal "24".; Primapus szarskii (Peters 1985) Mourer-Chauviré 1988Lua error: Internal error: The interpreter has terminated with signal "24".]
  • Aegialornithidae Lydekker 1891Lua error: Internal error: The interpreter has terminated with signal "24".
    • Primapus lacki Harrison & Walker 1975Lua error: Internal error: The interpreter has terminated with signal "24". [?Procuculus minutus Harrison & Walker 1977Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Eocene) ?apodid
    • Aegialornis Lydekker 1891Lua error: Internal error: The interpreter has terminated with signal "24". [Tachyornis Milne-Edwards 1892Lua error: Internal error: The interpreter has terminated with signal "24".; Belornis Milne-Edwards 1893Lua error: Internal error: The interpreter has terminated with signal "24".; Mesogiornis Mlíkovský 2002Lua error: Internal error: The interpreter has terminated with signal "24".](Early? – Late Eocene)
      • A. leenhardti
      • A. germanicus Mlíkovský 2002Lua error: Internal error: The interpreter has terminated with signal "24".
      • A. gallicus Lydekker 1891Lua error: Internal error: The interpreter has terminated with signal "24". [Tachyornis hirundo Milne-Edwards 1891Lua error: Internal error: The interpreter has terminated with signal "24".]
      • A. wetmorei Collins 1976Lua error: Internal error: The interpreter has terminated with signal "24". [Mesogiornis wetmorei (Collins 1976) Mlíkovský 2002Lua error: Internal error: The interpreter has terminated with signal "24".]
      • A. broweri Collins 1976Lua error: Internal error: The interpreter has terminated with signal "24". [Mesogiornis broweri (Collins 1976) Mlíkovský 2002Lua error: Internal error: The interpreter has terminated with signal "24".]
  • Eocypselidae Harrison 1984Lua error: Internal error: The interpreter has terminated with signal "24". – apodiform (hemiprocnid?)? caprimulgiform? basal Cypselomorphae?
    • Eocypselus Harrison 1984Lua error: Internal error: The interpreter has terminated with signal "24". (Late Paleocene ?- Early Eocene of NC Europe)
      • E. vincenti Harrison 1984 Lua error: Internal error: The interpreter has terminated with signal "24".
      • E. rowei Ksepka et al. 2013Lua error: Internal error: The interpreter has terminated with signal "24".
  • Cypselavidae Mourer-Chauviré 2006Lua error: Internal error: The interpreter has terminated with signal "24".
    • Parargornis messelensis Mayr 2003Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene) jungornithid, trochilid, basal as Argornis?
    • Argornis caucasicus Karhu 1999Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene) ?Trochilidae
    • Cypselavus gallicus Gaillard 1908Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene Early Oligocene) aegialornithid or hemiprocnid
  • Jungornithidae Karhu 1988Lua error: Internal error: The interpreter has terminated with signal "24".
    • Jungornis Karhu 1988Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene of N Caucasus, Russia)
      • J. tesselatus Karhu 1988Lua error: Internal error: The interpreter has terminated with signal "24".
      • J. geraldmayri Mourer-Chauviré & Sigé 2006Lua error: Internal error: The interpreter has terminated with signal "24".
  • Trochilidae hummingbirds
    • Eurotrochilus Mayr 2004Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene)
      • E. inexpectatus Mayr 2004Lua error: Internal error: The interpreter has terminated with signal "24".
      • E. noniewiczi Bochenski & Bochenski 2008Lua error: Internal error: The interpreter has terminated with signal "24".
    • Placement unresolved
      • Trochilidae sp. et gen. indet. (Bahamas, West Indies)
      • Trochilidae sp. et gen. indet. (Brazil)
  • Apodidae – swifts
    • Extant genera present in the fossil record

Mousebirds and relatives

  • Unresolved and basal forms
    • Botauroides parvus Shufeldt 1915Lua error: Internal error: The interpreter has terminated with signal "24". (Eocene of Wyoming, US)
    • Eobucco brodkorbi Feduccia & Martin 1976Lua error: Internal error: The interpreter has terminated with signal "24". - sandcoleid?
    • Eocolius walkeri Dyke & Waterhouse 2001Lua error: Internal error: The interpreter has terminated with signal "24". (London Clay Early Eocene of Walton-on-the-Naze, England) - sandcoleid or coliid
    • Limnatornis Milne-Edwards 1871Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene of Saint-Gérand-le-Puy, France) - coliid? (Urocolius?)
    • †Coliiformes gen. et sp. indet. (Late Miocene of Kohfidisch, Austria)[20]
    • Uintornis Marsh 1872Lua error: Internal error: The interpreter has terminated with signal "24". - sandcoleid?
  • Family †Chascacocoliidae Zelenkov & Dyke 2008Lua error: Internal error: The interpreter has terminated with signal "24".
    • Genus †Chascacocolius Houde & Olson 1992Lua error: Internal error: The interpreter has terminated with signal "24". (Late Paleocene ?- Early Eocene) - basal? sandcoleid?
  • Family †Selmeidae Zelenkov & Dyke 2008Lua error: Internal error: The interpreter has terminated with signal "24".
    • Selmes absurdipes Peters 1999Lua error: Internal error: The interpreter has terminated with signal "24". (Middle Eocene ?-Late Oligocene of C Europe) - coliid? (synonym of Primocolius?)
  • Family †Sandcoleidae Houde & Olson 1992 sensu Mayr & Mourer-Chauviré 2004Lua error: Internal error: The interpreter has terminated with signal "24".
    • Sandcoleus copiosus Houde & Olson 1992Lua error: Internal error: The interpreter has terminated with signal "24". (Paleocene)
    • Anneavis anneae Houde & Olson 1992Lua error: Internal error: The interpreter has terminated with signal "24".
    • Eoglaucidium pallas Fischer 1987Lua error: Internal error: The interpreter has terminated with signal "24".
  • Family Coliidae Swainson 1837 sensu Mayr & Mourer-Chauviré 2004Lua error: Internal error: The interpreter has terminated with signal "24".
    • Primocolius Mourer-Chauviré 1988Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene/Oligocene of Quercy, France)
    • Oligocolius Mayr 2000Lua error: Internal error: The interpreter has terminated with signal "24". (Early Oligocene of Frauenweiler, Germany)
    • Masillacolius brevidactylus Mayr & Peters 1998Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene of Messel, Germany)
    • Extant genera present in the fossil record
      • Colius [Necrornis Milne-Edwards 1871Lua error: Internal error: The interpreter has terminated with signal "24".]

†Zealandornithidae

  • Zealandornis relictus Worthy et al., 2022Lua error: Internal error: The interpreter has terminated with signal "24". (Early Miocene of Otago, New Zealand)

Owls and barn owls

  • Unresolved and basal forms
    • Berruornis (Late Paleocene) – basal? Sophornithidae?
    • Strigiformes gen. et sp. indet. (Late Paleocene)
    • Palaeoglaux (middle – Late Eocene) – own family Palaeoglaucidae or Strigidae?
    • Palaeobyas (Late Eocene/Early Oligocene) – Tytonidae? Sophiornithidae?
    • Palaeotyto (Late Eocene/Early Oligocene) – Tytonidae?
    • Strigiformes gen. et spp. indet. (Early Oligocene)
  • Ogygoptyngidae
  • Protostrigidae
    • Eostrix (Early – middle Eocene)
    • Minerva (middle – Late Eocene) – formerly Protostrix, includes "Aquila" ferox, "Aquila" lydekkeri, and "Bubo" leptosteus
    • Oligostrix (middle Oligocene)
  • Sophiornithidae
    • Sophiornis
  • Strigidae – typical owls
    • Mioglaux (Late Oligocene? – Early Miocene) – includes "Bubo" poirreiri
    • Intulula (Early/Middle –? Late Miocene) – includes "Strix/Ninox" brevis
    • Alasio (Middle Miocene) – includes "Strix" collongensis
    • Oraristrix (Late Pleistocene)
    • Miosurnia (Late Miocene)
    • Placement unresolved
      • "Otus" wintershofensis (Early/Middle Miocene) – may be close to extant genus Ninox
      • "Strix" edwardsi (Middle/Late? Miocene)
      • "Asio" pygmaeus (Early Pliocene)
      • Strigidae gen. et sp. indet. UMMP V31030 (Late Pliocene) – Strix/Bubo?
    • Extant genera present in the fossil record
      • Strix (Early Miocene – Recent)
      • Bubo (Late Miocene? – Recent)
      • Asio (Late Pliocene – Recent)
      • Athene (Late Pliocene – Recent)
      • Glaucidium (Late Pliocene – Recent)
      • Surnia (Late Pliocene – Recent)
      • Pulsatrix (Late Pleistocene – Recent)
  • Tytonidae – barn owls
    • Nocturnavis (Late Eocene/Early Oligocene)
    • Selenornis (Late Eocene/Early Oligocene)
    • Necrobyas (Late Eocene/Early Oligocene – Early Miocene)
    • Prosybris (Early Oligocene? – Early Miocene)
    • Placement unresolved
      • Tytonidae gen. et sp. indet. TMT 164 (Middle Miocene)
    • Extant genera present in the fossil record
      • Tyto (Late Miocene – Recent)

Rollers and allies. Probably paraphyletic.

  • Basal and unresolved forms
    • Quasisyndactylus longibrachis Mayr 1998Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene) – alcediniform, basal?
    • Paracoracias occidentalis Clarke et al. 2009Lua error: Internal error: The interpreter has terminated with signal "24".
    • Cryptornis antiquus Gervais 1848–52 [nomen nudum]Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene) – bucerotid? geranopterid?
    • Protornis glarniensis von Meyer 1856Lua error: Internal error: The interpreter has terminated with signal "24". (Oligocene) – basal to motmotids and meropids?
    • † Coraciiformes gen. et spp. indet. PQ 1216, QU 15640 (Late Eocene) – 2 species
  • Geranopteridae Mayr & Mourer-Chauviré 2000Lua error: Internal error: The interpreter has terminated with signal "24".
    • Geranopterus Milne-Edvards 1892Lua error: Internal error: The interpreter has terminated with signal "24". (Late Eocene – Early Miocene)
      • G. alatus (Mayr & Mourer-Chauvire 2000)Lua error: Internal error: The interpreter has terminated with signal "24".
      • G. bohemicus (Mlikovsky 1999) Mourer-Chauvire 1999Lua error: Internal error: The interpreter has terminated with signal "24". [Nupharanassa bohemica Mlikovsky 1999Lua error: Internal error: The interpreter has terminated with signal "24".]
      • G. milneendwardsi Mayr & Mourer-Chauvire 2000Lua error: Internal error: The interpreter has terminated with signal "24".
  • Eocoraciidae Mayr & Mourer-Chauviré 2000Lua error: Internal error: The interpreter has terminated with signal "24".
    • Eocoracias brachyptera Mayr & Mourer-Chauviré 2000Lua error: Internal error: The interpreter has terminated with signal "24". (middle Eocene)
  • Primobucconidae Feduccia & Martin 1976 sensu Mayr, Mourer-Chauviré & Weidig 2004Lua error: Internal error: The interpreter has terminated with signal "24".
    • Primobucco Brodkorb 1970Lua error: Internal error: The interpreter has terminated with signal "24". [Neanis Brodkorb 1965 non Gistl 1848Lua error: Internal error: The interpreter has terminated with signal "24".] (Early – middle Eocene)
      • P. kestneri Feduccia 1973Lua error: Internal error: The interpreter has terminated with signal "24". [Neanis kestneri (Feduccia 1973) Feduaccia 1976Lua error: Internal error: The interpreter has terminated with signal "24".]
      • P. frugilegus Mayr, Mourer-Chauviré & Weidig 2004Lua error: Internal error: The interpreter has terminated with signal "24".
      • P. mcgrewi Brodkorb 1970Lua error: Internal error: The interpreter has terminated with signal "24".
      • P. perneri Mayr, Mourer-Chauviré & Weidig 2004Lua error: Internal error: The interpreter has terminated with signal "24".
  • Todidae – todies
    • Palaeotodus Olson 1976 (Late Eocene of France – Early Oligocene of WC Europe and Wyoming)
      • P. emryi Olson 1976Lua error: Internal error: The interpreter has terminated with signal "24".
      • P. escampsiensis Mourer-Chauviré 1985Lua error: Internal error: The interpreter has terminated with signal "24".
      • P. itardiensis Mourer-Chauviré 1985Lua error: Internal error: The interpreter has terminated with signal "24".
  • Motmotidae – motmots
    • Placement unresolved
      • Momotidae gen. et sp. indet. (Late Miocene)
  • Coraciidae
    • Miocoracias chenevali Mourer-Chauviré, Peyrouse & Hugueney 2013Lua error: Internal error: The interpreter has terminated with signal "24".
    • Extant genera present in the fossil record
  • Messelirrisoridae
    • Messelirrisor Mayr 1998 (middle Eocene)
      • M. halcyorostris Mayr 1998Lua error: Internal error: The interpreter has terminated with signal "24".
      • M. parvus Mayr 1998Lua error: Internal error: The interpreter has terminated with signal "24".
      • M. grandis Mayr 2000Lua error: Internal error: The interpreter has terminated with signal "24".
  • Phoeniculidae (woodhoopoes)
    • Phirriculus pinicola Mlíkovský & Göhlich 2000Lua error: Internal error: The interpreter has terminated with signal "24".
  • Bucerotidae – hornbills
    • Euroceros bulgaricus Boev & Kovachev 2007Lua error: Internal error: The interpreter has terminated with signal "24". (Late Miocene of Hadzhidimovo, Bulgaria)
    • Extant genera present in the fossil record
      • Bucorvus
        • B. brailloni Brunet 1971Lua error: Internal error: The interpreter has terminated with signal "24".
  • Trogonidae – trogons
    • Foshanornis songi Zhao et al. 2015Lua error: Internal error: The interpreter has terminated with signal "24".
    • Septentrogon madseni Kristoffersen 2002 (Fur Late Paleocene/Early Eocene of Ejerslev, Denmark)
    • Paratrogon gallicus (Milne-Edwards 1871) Lambrecht 1933Lua error: Internal error: The interpreter has terminated with signal "24". [Trogon gallicus Milne-Edwards 1871Lua error: Internal error: The interpreter has terminated with signal "24".] (Early Miocene of France)
    • Primotrogon Mayr 1999Lua error: Internal error: The interpreter has terminated with signal "24". [Masillatrogon Mayr 2009Lua error: Internal error: The interpreter has terminated with signal "24".] (middle Eocene of Messel, Germany? - Early Oligocene of France)
      • P. wintersteini Mayr 1999Lua error: Internal error: The interpreter has terminated with signal "24".
      • P. pumilio Mayr 2005Lua error: Internal error: The interpreter has terminated with signal "24". [Masillatrogon pumilio (Mayr 2005) Mayr 2009Lua error: Internal error: The interpreter has terminated with signal "24".]
    • Placement unresolved
      • Trogonidae gen. et sp. indet. 1 (NW Europe)
      • Trogonidae gen. et sp. indet. 2 (NW Europe)
  • Placement unresolved
    • Piciformes gen. et sp. indet. IRScNB Av 65 (Early Oligocene)
    • Rupelramphastoides (Early Oligocene) – ramphastid?
    • Piciformes gen. et sp. indet. SMF Av 429 (Late Oligocene)
    • Capitonides (Early – Middle Miocene) – ramphastid? "capitonid" (Lybiidae, Megalaimidae)? own family Capitonididae?
    • Pici gen. et sp. indet. (Middle Miocene) – "capitonid" (Lybiidae, Megalaimidae?)
  • Miopiconidae
    • Miopico
  • Lybiidae – African barbets
    • Lybiidae gen. et sp. indet. (Late Miocene) – extant genus Pogoniulus?
  • Galbulidae
  • Picavidae
  • Picidae – woodpeckers
    • Palaeopicus (Late Oligocene)
    • Palaeonerpes (Early Pliocene)
    • Pliopicus (Early Pliocene)
    • Placement unresolved
      • Picidae gen. et sp. indet. (Middle Miocene)
      • Picidae gen. et sp. indet. (Late Miocene)
      • cf. Colaptes DMNH 1262 (Early Pliocene of Ainsworth, US)
    • Extant genera present in the fossil record
  • Placement unresolved
    • Passeriformes gen. et spp. indet. (Early Eocene) – several species, oscine?
    • Resoviaornis (Early Oligocene)
    • Wieslochia (Early Oligocene)
    • Passeriformes gen. et spp. indet. (Late Oligocene) – several suboscine and oscine species
    • Certhiops (Early Miocene of Germany) – basal Certhioidea
    • Passeriformes gen. et sp. indet. (Early/Middle Miocene) – suboscine?
    • Passeriformes gen. et spp. indet. (Early/Middle Miocene) – several species, oscine?
    • Passeriformes gen. et spp. indet. (Middle Miocene) – several species, basal?
    • Passeriformes gen. et spp. indet. (Middle Miocene) – several species, oscine?
    • Passeriformes gen. et spp. indet. (Late Miocene) – Sylvioidea
    • "Palaeostruthus" eurius (Pliocene)
  • Eurylaimidae – broadbills
    • Placement unresolved
      • Eurylaimidae gen. et sp. indet. (Early Miocene)
  • Palaeoscinidae
    • Palaeoscinis (Late Miocene)
  • Furnariidae – ovenbirds
    • Pseudoseisuropsis
    • Prehistoric species of extant genera
      • Pseudoseisura cursor (Ensenada Early/Middle Pleistocene of Anchorena, Argentina)
      • Cinclodes major (Middle Pleistocene of Buenos Aires Province, Argentina)
  • Menuridae – lyretails
    • Extant genera present in the fossil record
      • Menura (Early Miocene – Recent)
  • Meliphagidae – honeyeaters
    • Placement unresolved
      • Meliphagidae gen. et spp. indet. (Middle/Late Miocene – Pliocene of Riversleigh, Australia) – at least 7 spp., some may be from extant genera
  • Orthonychidae – logrunners
    • Extant genera present in the fossil record
      • Orthonyx (Middle/Late Miocene – Recent)
  • Oriolidae – Old World orioles
    • Longmornis (Early Miocene of Riversleigh, Australia)
  • Artamidae – woodswallows, butcherbirds, currawongs and Australian magpie
    • Placement unresolved
      • Artamidae gen. et sp. indet. (Early/Middle Miocene) – cracticine
  • Corvidae – crows, ravens, jays and magpies
    • Miocorvus (Middle Miocene)
    • Miopica (Middle Miocene)
    • Miocitta (Late Miocene)
    • Protocitta (Early Pleistocene)
    • Henocitta (Middle Pleistocene)
    • Extant genera present in the fossil record
      • Corvus (Late Miocene — Recent)
      • Pica (Late Pliocene/Early Pleistocene – Recent)
      • Pyrrhocorax
    • Placement unresolved
      • Corvidae gen. et sp. indet. (Early Pliocene)
      • Corvidae gen. et sp. indet. (Early/Middle Pleistocene) – probably belongs in extant genus
  • Laniidae – shrikes
    • Extant genera present in the fossil record
      • Lanius (Early Miocene – Recent)
  • Regulidae – kinglets
    • Extant genera present in the fossil record
      • Regulus (Late Pliocene – Recent)
  • Hirundinidae – swallows and martins
    • Placement unresolved
      • Hirundinidae gen. et spp. indet. (Early Pliocene of Langebaanweg, South Africa) – 2 species
  • Megaluridae – grass-warblers and allies
    • Extant genera present in the fossil record
  • Acrocephalidae – marsh- and tree-warblers
    • Extant genera present in the fossil record
  • Muscicapidae – Old World flycatchers and chats
    • Extant genera present in the fossil record
  • Turdidae – thrushes
    • Extant genera present in the fossil record
      • ?Turdus (Middle? Miocene – Recent)
  • Alaudidae – larks
    • Eremarida (Late Miocene of Hrabarsko, Bulgaria)
  • Motacillidae – wagtails
    • Extant genera present in the fossil record
  • Fringillidae – finches
    • Extant genera present in the fossil record
      • Loxia (Late Pliocene – Recent)
    • Additional prehistoric species of extant genera
      • Coccothraustes simeonovi (Late Pliocene of Varshets, Bulgaria)
      • Coccothraustes balcanicus
  • Icteridae – grackles and New World orioles
    • Pandanaris (Pleistocene)
    • Pyelorhamphus (Pleistocene)
    • Extant genera present in the fossil record
  • Cardinalidae – cardinals
    • Placement unresolved
      • Passerina sp. (Early Pliocene of Yepómera, Mexico)
  • Emberizidae – buntings and New World sparrows
    • Pampaemberiza (Middle Pleistocene of Necochea, Argentina)
    • Extant genera present in the fossil record
      • Ammodramus (Late Miocene – Recent) – including Palaeostruthus
    • Additional prehistoric species of extant genera
      • Pipilo angelensis (Pleistocene of Rancho La Brea, US)

These fossil taxa cannot be assigned to any major group with reasonable certainty. The "proto-birds" above are of some indeterminate basal position in the entire avialan (and paravian) radiation, but known from such diagnostic material that their relationships at the family level are known. In contrast, the taxa here have a hypodigm that is usually just sufficient for giving them a valid scientific name, but not for phylogenetic purposes beyond classing them as pygostylians or more modern birds. Some, however, are known from such fragmentary remains that the possibility that they are non-avian "reptiles" such as dinosaurs cannot be ruled out at present.

  • "Ichthyornis" minusculus Nesov 1990 (Bissekty Late Cretaceous of Kyzyl Kum, Uzbekistan) – enantiornithine?
  • Qinornis paleocenica Xue 1995 (Early/Middle Paleocene) – enantiornithine? neornithine?


See also

Footnotes

  1. Sereno (2005)
  2. Chiappe (2001, 2002)
  3. See e.g. Mortimer (2004), Sereno (2005)
  4. "Taxonomic lists- Aves". http://www.paleofile.com/. 
  5. Brodkob, Pierce (1963). "Catalogue of fossil birds 1- Archaeopterygiformes through Ardeiformes". Biological Sciences, Bulletin of the Florida State Museum 7 (4): 180–293. 
  6. Clarke et al. (2006)
  7. 7.0 7.1 Holtz, Thomas R. Jr. (2007). Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages. New York NY, US: Random House. ISBN 978-0-375-82419-7. https://archive.org/details/dinosaursmostcom00holt.  Winter 2011 Appendix.
  8. Kurochkin, E.N. et al. (2006)
  9. O'Connor, J. K.; Zhang, Y.; Chiappe, L. M.; Meng, Q.; Quanguo, L.; Di, L. (2013). "A new enantiornithine from the Yixian Formation with the first recognized avian enamel specialization". Journal of Vertebrate Paleontology 33 (1): 1–12. doi:10.1080/02724634.2012.719176. Bibcode2013JVPal..33....1O. 
  10. Li Li; En-pu Gong; Li-dong Zhang; Ya-jun Yang; Lian-hai Hou (2010). "A new enantiornithine bird (aves) from the Early Cretaceous of Liaoning, China". Acta Palaeontologica Sinica 49 (4): 524–531. http://en.cnki.com.cn/Article_en/CJFDTotal-GSWX201004012.htm. 
  11. Gareth J. Dyke, Attila Ősi (2010). "A review of Late Cretaceous fossil birds from Hungary". Geological Journal 45 (4): 434–444. doi:10.1002/gj.1209. 
  12. Fernando Emilio Novas, Federico Lisandro Agnolín and Carlos Agustín Scanferla (2010). "New enantiornithine bird (Aves, Ornithothoraces) from the Late Cretaceous of NW Argentina". Comptes Rendus Palevol 9 (8): 499–503. doi:10.1016/j.crpv.2010.09.005. Bibcode2010CRPal...9..499N. https://www.sciencedirect.com/science/article/pii/S1631068310001144. 
  13. Kurochkin (2006)
  14. O'Connor, J.K. (2012)
  15. Fide Chiappe (2002)
  16. Federico L. Agnolin and David Varricchio (2012). "Systematic reinterpretation of Piksi barbarulna Varricchio, 2002 from the Two Medicine Formation (Upper Cretaceous) of Western USA (Montana) as a pterosaur rather than a bird". Geodiversitas 34 (4): 883–894. doi:10.5252/g2012n4a10. http://www.mnhn.fr/museum/foffice/science/science/DocScientifique/publications/presentation/publicationAP/ficheContenu.xsp?CONTENU_ID=4592&PARUTION_ID=2966&PUBLICATION_ID=21&idx=6&nav=listeParution. 
  17. Tennyson, A.J.D.; Worthy, T.H.; Jones, C.M.; Scofield, R.P.; Hand, S.J. (2010). "Moa's Ark: Miocene fossils reveal the great antiquity of moa (Aves: Dinornithiformes) in Zealandia". Records of the Australian Museum 62: 105–114. doi:10.3853/j.0067-1975.62.2010.1546. 
  18. 18.0 18.1 Nikita V. Zelenkov; Thomas A. Stidham; Nicolay V. Martynovich; Natalia V. Volkova; Qiang Li; Zhuding Qiu (2018). "The middle Miocene duck Chenoanas (Aves, Anatidae): new species, phylogeny and geographical range". Papers in Palaeontology 4 (3): 309–326. doi:10.1002/spp2.1107. 
  19. Suárez, William; Olson, Storrs L. (2021). "A new fossil raptor (Accipitridae: Buteogallus) from Quaternary cave deposits in Cuba and Hispaniola, West Indies". Bulletin of the British Ornithologists' Club 141 (3). doi:10.25226/bboc.v141i3.2021.a3. 
  20. Similar to Urocolius and Limnatornis (if distinct): Mlíkovský (2002)

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References

  • Chiappe, Luis M. (2001): The rise of birds. In: Briggs, Derek E.G. & Crowther, P.R. (eds.): Palaeobiology II: A Synthesis: 102-106. Cambridge University Press, Cambridge, UK.
  • Chiappe, Luis M. (2002): Basal bird phylogeny: problems and solutions. In: Chiappe, L.M. and Witmer, L.M. (eds.): Mesozoic Birds: Above the Heads of Dinosaurs: 448-472. University of California Press, Berkeley, US.
  • Clarke, Julia A.; Zhou, Zhonghe; Zhang, Fucheng (2006). "Insight into the evolution of avian flight from a new clade of Early Cretaceous ornithurines from China and the morphology of Yixianornis grabaui". Journal of Anatomy 208 (3): 287–308. doi:10.1111/j.1469-7580.2006.00534.x. PMID 16533313.  Electronic Appendix
  • Gauthier, Jacques A. & de Queiroz, Kevin (2001): Feathered dinosaurs, flying dinosaurs, crown dinosaurs, and the name "Aves". In: Gauthier, Jacques & Gall, L.F. (eds.): New Perspectives on the Origin and Early Evolution of Birds: Proceedings of the International Symposium in Honor of John H. Ostrom: 7-41. Peabody Museum of Natural History, Yale University, New Haven, Connecticut.
  • Mortimer, Michael (2004): The Theropod Database: Phylogeny of taxa. Retrieved 2013-MAR-02.
  • Olson, Storrs L. (1985): The fossil record of birds. In: Farner, D.S.; King, J.R. & Parkes, Kenneth C. (eds.): Avian Biology 8: 79-238. Academic Press, New York. Not in copyright; PDF fulltext
  • Sereno, Paul Callistus (2005): TaxonSearch: Stem Archosauria. Version 1.0, November 7, 2005. Retrieved 2007-APR-30.

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