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The serpent seed doctrine is the belief that Eve literally had sexual intercourse with the serpent in the Garden of Eden, leading to offspring.
How can sexual intercourse between a human and a snake take place, you ask? Ah, well, it happened before The Fall, so God hadn't gotten mad at the serpent yet… so the serpent wasn't "crawling in the dust" yet… so the serpent was pretty much human-shaped, like you or I. So Eve bangs the serpent and produces Cain — the spawn of Satan (although nowhere in the Old Testament does it say that the serpent was Satan). Abel, on the other hand (and Seth later), would have been a child of Adam (and thus a grandchild of God, we suppose). Similar to the "Curse of Ham" doctrine, some believers also hold that the "serpent-seed" were non-white people.
Serpent seed believers differ on the story from that point on. Did Cain's offspring get wiped out in the Great Flood? Were they the Kenites, a tribe living in the greater Canaan metropolitan area during the Exodus? Did they interbreed with their more godly neighbors so everyone alive today has a little piece of original sin in them? Or are only some of us *cough*Democrats*cough* Satan-spawn, and others (say, Southern Baptists) pure sons of Adam?
Anti-semites sometimes use verses like Matthew 12:34 and John 8:44 to claim that the descendants of the serpent/Satan are the Jews.