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    Colaspoides

    From Wikipedia - Reading time: 14 min

    Colaspoides
    Temporal range: Eocene–recent, 37.2–0 Ma[1]
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    Scientific classification Edit this classification
    Domain: Eukaryota
    Kingdom: Animalia
    Phylum: Arthropoda
    Class: Insecta
    Order: Coleoptera
    Infraorder: Cucujiformia
    Family: Chrysomelidae
    Subfamily: Eumolpinae
    Tribe: Eumolpini
    Genus: Colaspoides
    Laporte, 1833[2]
    Type species
    Cryptocephalus limbatus
    Fabricius, 1781
    Species

    See text

    Synonyms[3]
    • Amasia Dejean, 1836
      (nomen nudum)
    • Melina Chevrolat in Dejean, 1836
      (nomen nudum)
    • Colaspidoides Agassiz, 1846
    • Amasia Chapuis, 1874[4]
    • Melina Chapuis, 1874
      (preoccupied name)      [4]
    • Melinophora Lefèvre, 1885[5]
    • Tailandia Chûjô, 1964

    Colaspoides is a genus of leaf beetles in the subfamily Eumolpinae. It is one of the largest genera in the subfamily, containing over 260 species worldwide.[6] It is an extant genus but there is at least one species, C. eocenicus, found in Baltic amber from the Upper Eocene of Russia,[7] and the genus has also been reported from the Miocene of the Dominican Republic.[1]

    The Neotropical species of Colaspoides, including the type species, lack elytra-locking grooves on the pygidium, unlike in the Asian species and in similar-looking Neotropical Eumolpinae genera such as Beltia. According to Flowers (2018), this suggests that the Asian species of Colaspoides should be transferred to different genera.[8]

    Species[edit]

    The genus contains the following species:[9]

    Neotropical species:[10][11]

    • Colaspoides abdominalis Jacoby, 1900[12]
    • Colaspoides alcyonea (Erichson, 1847)
    • Colaspoides amabilis Lefèvre, 1876
    • Colaspoides amazona Jacoby, 1881[13]
    • Colaspoides batesi Jacoby, 1879[14]
    • Colaspoides bicolor (Olivier, 1808)
    • Colaspoides cupreipennis Jacoby, 1890
    • Colaspoides elongata Jacoby, 1880
    • Colaspoides fasciata Lefèvre, 1875
    • Colaspoides freyi Bechyné, 1955[15]
    • Colaspoides fulgida Lefèvre, 1875
    • Colaspoides hlisnikovskyi Bechyné, 1949[16]
    • Colaspoides hypoxantha Lefèvre, 1885
      • Colaspoides hypoxantha hypoxantha Lefèvre, 1885
      • Colaspoides hypoxantha nisotricollis Bechyné, 1954[17]
    • Colaspoides igneicollis Lefèvre, 1891
    • Colaspoides inedita Lefèvre, 1876
    • Colaspoides itaituba Bechyné & Bechyné, 1964[18]
    • Colaspoides limbata (Fabricius, 1781)
    • Colaspoides limbicollis Lefèvre, 1875
    • Colaspoides marginata Jacoby, 1882[19]
    • Colaspoides mera (Germar, 1824)
    • Colaspoides peruana Jacoby, 1879[14]
    • Colaspoides quadriplagiata Jacoby, 1882[19]
    • Colaspoides reticulata Lefèvre, 1876
    • Colaspoides rufitarsis Lefèvre, 1875
    • Colaspoides smaragdina Lefèvre, 1875
    • Colaspoides tibialis (Germar, 1824)
    • Colaspoides tibiella Bechyné, 1953[11]
    • Colaspoides unicolor Jacoby, 1879[14]
    • Colaspoides varicolor Lefèvre, 1878
    • Colaspoides viridicollis Jacoby, 1878[20]
    • Colaspoides viridicornis Lefèvre, 1878

    Palaearctic, Indomalaysian and Australasian species:[21][6][22]

    • Colaspoides acervata Lea, 1915[23]
    • Colaspoides ahmadi Medvedev, 2010[24]
    • Colaspoides amamianus Komiya, 1998
    • Colaspoides angusticollis Jacoby, 1898[25]
    • Colaspoides annamita Medvedev, 2004[6]
    • Colaspoides annandalei Jacoby, 1905
    • Colaspoides anomogastra Lea, 1915[23]
    • Colaspoides apicata Medvedev, 2004[6]
    • Colaspoides apicicornis Jacoby, 1884
    • Colaspoides armata Medvedev, 2004[6]
    • Colaspoides bacboensis Medvedev, 2004[6]
    • Colaspoides bakeri Medvedev, 2006[26]
    • Colaspoides balyana Medvedev, 2007[27]
    • Colaspoides basilana Medvedev, 2006[26]
    • Colaspoides bengalensis Duvivier, 1892
    • Colaspoides bicarinata Lea, 1915[23]
    • Colaspoides bicoloricollis Medvedev & Romantsov, 2014[28]
    • Colaspoides bicoloripes Medvedev, 2019[29]
    • Colaspoides bidentatus Medvedev, 2004[6]
    • Colaspoides biplagiata Baly, 1867[30]
    • Colaspoides bolmi Medvedev, 2010[24]
    • Colaspoides borneoensis Jacoby, 1898[25]
    • Colaspoides brancuccii Medvedev, 2004[6]
    • Colaspoides brevicollis Jacoby, 1898[25]
    • Colaspoides brunnea Bryant, 1957[31]
    • Colaspoides buechi Medvedev, 2008[32]
    • Colaspoides buonloicus Medvedev, 2004[6]
    • Colaspoides caledonica Medvedev, 2007[27]
    • Colaspoides cantonensis Medvedev, 2004[6]
    • Colaspoides cariniventris Lea, 1926[33]
    • Colaspoides cechovskyi Medvedev, 2016[34]
    • Colaspoides chakratongii (Chûjô, 1964)[3]
    • Colaspoides chapuisi Medvedev, 2004[6]
    • Colaspoides cheni Medvedev, 2004[6]
    • Colaspoides chinensis Jacoby, 1888[35]
    • Colaspoides ciliaticornis Medvedev, 2019[36]
    • Colaspoides ciliatipes Lefèvre, 1890
    • Colaspoides circumdatus Medvedev & Romantsov, 2014[28]
    • Colaspoides clavipes Medvedev, 2004[6]
    • Colaspoides coerulescens Baly, 1867[30]
    • Colaspoides coerulipes Baly, 1867[30]
    • Colaspoides confusa Gressitt, 1957[31]
    • Colaspoides cognata Baly, 1867[30]
    • Colaspoides cognatella Medvedev, 2004[6]
    • Colaspoides cognatomima Medvedev, 2004[6]
    • Colaspoides complicata Lea, 1915[23]
    • Colaspoides costalis Medvedev, 2004[6]
    • Colaspoides crassifemur Tan & Wang, 1984
    • Colaspoides crassipes Lea, 1915[23]
    • Colaspoides cuprea Baly, 1867[30]
    • Colaspoides cupreicollis Jacoby, 1908[37]
    • Colaspoides cupreoviridis Lea, 1922[38]
    • Colaspoides curticornis Medvedev, 2010[24]
    • Colaspoides curvipes Medvedev, 2004[6]
    • Colaspoides cyaneipennis Medvedev, 2010[24]
    • Colaspoides daccordii Medvedev, 2004[6]
    • Colaspoides dapi Medvedev, 2004[6]
    • Colaspoides diffinis Lefèvre, 1893[39]
    • Colaspoides dimorphus Medvedev, 2004[6]
    • Colaspoides doddi Lea, 1915[23]
    • Colaspoides dohertii Jacoby, 1908[37]
    • Colaspoides dongnaicus Medvedev, 2012[40]
    • Colaspoides elegans Baly, 1867[30]
    • Colaspoides elegantula Lea, 1915[23]
    • Colaspoides elenae Medvedev & Romantsov, 2014[28]
    • †Colaspoides eocenicus Moseyko & Kirejtshuk, 2013[7]
    • Colaspoides excavativentris Lea, 1926[33] (excaviventris?)
    • Colaspoides fasciculata Lea, 1921[41]
    • Colaspoides feae Jacoby, 1882
    • Colaspoides fedorenkoi Medvedev, 2012[40]
    • Colaspoides femoralis Lefèvre, 1885[5]
    • Colaspoides flavimana Medvedev, 2004[6]
    • Colaspoides fontis Jolivet, Verma & Mille, 2008[42]
    • Colaspoides foveiventris Lea, 1915[23]
    • Colaspoides frenchi Lea, 1915[23]
    • Colaspoides fulvicornis (Baly, 1865)
    • Colaspoides fulvimana Jacoby, 1908[37]
    • Colaspoides fulvipes Lefèvre, 1889
    • Colaspoides fulvitarsis Jacoby, 1899[43]
    • Colaspoides fulvus (Chûjô, 1935)
    • Colaspoides fulvus Medvedev, 2015[44] (junior homonym of above)
    • Colaspoides fuscoaenea Baly, 1867[30]
    • Colaspoides geminata Weise, 1908
    • Colaspoides geniculatus Medvedev, 2004[6]
    • Colaspoides glabrata Jacoby, 1898
    • Colaspoides glabricollis Jacoby, 1908[37]
    • Colaspoides gorbunovi Medvedev, 2010[24]
    • Colaspoides gratiosa (Baly, 1864)
    • Colaspoides gressetti Medvedev, 2004[6]
    • Colaspoides haemorrhoidalis Lea, 1915[23]
    • Colaspoides hainanensis Gressitt & Kimoto, 1961
    • Colaspoides hageni Lefèvre, 1887
    • Colaspoides hagiangi Medvedev, 2004[6]
    • Colaspoides heroni Lea, 1915[23]
    • Colaspoides hoblerae Lea, 1915[23]
    • Colaspoides howensis Lea, 1915[23]
    • Colaspoides icterica Weise, 1922[45]
    • Colaspoides imasakai Komiya, 1991
    • Colaspoides imitans Medvedev, 2004[6]
    • Colaspoides inconspicua Jacoby, 1896
    • Colaspoides inornata Baly, 1867[30]
    • Colaspoides insignis Baly, 1867[30]
    • Colaspoides iridipennis Jacoby, 1896
    • Colaspoides jacobyi Clavareau, 1914
    • Colaspoides jacobyi Medvedev, 2004[6] (junior homonym of above)
    • Colaspoides japanus Chûjô, 1956
    • Colaspoides javana Weise, 1924
    • Colaspoides kabakovi Medvedev, 2004[6]
    • Colaspoides kantneri Medvedev, 2004[6]
    • Colaspoides kasaharai Komiya, 1991
    • Colaspoides kelantana Medvedev, 2019[36]
    • Colaspoides kimotoi Medvedev, 2004[6]
    • Colaspoides klimenkoi Medvedev & Romantsov, 2014[28]
    • Colaspoides kubani Medvedev, 2004[6]
    • Colaspoides laeta Medvedev, 2004[6]
    • Colaspoides laevicollis Lefèvre, 1887
    • Colaspoides lamellatus Medvedev, 2004[6]
    • Colaspoides langbianicus Kimoto & Gressitt, 1982[3]
    • Colaspoides laosensis Kimoto & Gressitt, 1982[3]
    • Colaspoides laotica Medvedev, 2004[6]
    • Colaspoides laportei Baly, 1867[30] (laportii?)
    • Colaspoides lateralis Jacoby, 1900
    • Colaspoides laticollis Jacoby, 1900
    • Colaspoides laysi Medvedev, 2006[26]
    • Colaspoides leai Medvedev, 2001[46]
    • Colaspoides lefevrei Jacoby, 1889
    • Colaspoides lobatus Medvedev, 2004[6]
    • Colaspoides macgregori Medvedev, 2006[26]
    • Colaspoides malayana Jacoby, 1894[47]
    • Colaspoides malayensis Medvedev, 2004[6]
    • Colaspoides martini Lefèvre, 1885[48]
      • Colaspoides martini martini Lefèvre, 1885[48]
      • Colaspoides martini spinigerus Lefèvre, 1893[39][6]
    • Colaspoides medogensis Tan, 1989
    • Colaspoides melanocephala Jacoby, 1908[37]
    • Colaspoides mentaweica Medvedev, 2003[49]
    • Colaspoides micans Baly, 1867[30]
    • Colaspoides microdentata Medvedev, 2004[6]
    • Colaspoides mimeta Lea, 1915[23]
    • Colaspoides mimica Medvedev, 2004[6]
    • Colaspoides mindorensis Medvedev, 2006[26]
    • Colaspoides minimus Kimoto & Gressitt, 1982[3]
    • Colaspoides minuta Medvedev, 1995[50]
    • Colaspoides mirabilis Medvedev, 2010[24]
    • Colaspoides miyatakei Kimoto, 1967[51]
    • Colaspoides modesta Baly, 1867[30]
    • Colaspoides modiglianii Jacoby, 1896[52]
    • Colaspoides montana Jacoby, 1900
    • Colaspoides morimotoi Kimoto, 1967[51]
    • Colaspoides multicostata Medvedev, 2007[27]
    • Colaspoides napolovi Medvedev, 2012[53]
    • Colaspoides negrosana Medvedev, 2006[26]
    • Colaspoides nepalensis Kimoto, 2001[54]
    • Colaspoides nitidicollis Medvedev, 2010[24]
    • Colaspoides nigricornis Jacoby, 1884
    • Colaspoides nigripes Jacoby, 1884
    • Colaspoides nigritarsis Jacoby, 1885
    • Colaspoides nigrotibialis Medvedev, 2004[6]
    • Colaspoides norfolcensis Lea, 1915[23]
    • Colaspoides obscuripes Medvedev, 2010[24]
    • Colaspoides okinawanus Komiya, 1991[55]
    • Colaspoides okumai Komiya, 1991
    • Colaspoides olegi Medvedev, 2004[6]
    • Colaspoides opaca Jacoby, 1888[35]
    • Colaspoides ovalis Lefèvre, 1890
    • Colaspoides paddis Aslam, 1968[56]
    • Colaspoides pallidicornis Tan & Zhou, 1997
    • Colaspoides parrotti Medvedev, 2006[26]
    • Colaspoides parvidens Lea, 1915[23]
    • Colaspoides parvula Baly, 1867[30]
    • Colaspoides patrikeevi Moseyko & Romantsov, 2022[57]
    • Colaspoides persicariae Chûjô, 1956
    • Colaspoides persimilis Kimoto & Gressitt, 1982[3]
    • Colaspoides philippinensis Baly, 1867[30]
    • Colaspoides picea Baly, 1867[30]
    • Colaspoides piceana Medvedev, 2004[6]
    • Colaspoides picipes Weise, 1908
    • Colaspoides picticornis Lea, 1915[23]
    • Colaspoides pictipes Lea, 1915[23]
    • Colaspoides pilicornis Lefèvre, 1882
    • Colaspoides planifrons Jacoby, 1900
    • Colaspoides pleuralis Medvedev, 2010[24]
    • Colaspoides poeciloderma Lea, 1915[23]
    • Colaspoides polilovi Medvedev, 2010[58]
    • Colaspoides prasinella Medvedev, 2004[6]
    • Colaspoides prasinus Lefèvre, 1890
      • Colaspoides prasinus occidentalis Medvedev, 2004[6]
      • Colaspoides prasinus prasinus Lefèvre, 1890
    • Colaspoides pseudodiffinis Medvedev, 2004[6]
    • Colaspoides pseudofemoralis Romantsov & Moseyko, 2019[59]
    • Colaspoides pseudomodesta Medvedev, 2010[24]
    • Colaspoides pseudorufa Medvedev, 2015[44]
    • Colaspoides pulchella Clark, 1865[60]
    • Colaspoides puncticeps Baly, 1867[30]
    • Colaspoides punctipleuris Medvedev, 2003[49]
    • Colaspoides purpurascens Medvedev, 2015[61]
    • Colaspoides purpurata Medvedev, 2006[26]
    • Colaspoides quadripartita Baly, 1867[30]
    • Colaspoides quieta Lea, 1915[23]
    • Colaspoides rafflesii Baly, 1867[30]
    • Colaspoides rara Lea, 1915[23]
    • Colaspoides rectilatera Lea, 1915[23]
    • Colaspoides regalini Medvedev, 2004[6]
    • Colaspoides regularis Baly, 1867[30]
    • Colaspoides riedeli Medvedev, 2010[24]
    • Colaspoides robusta Baly, 1867[30]
    • Colaspoides rubra Medvedev, 2010[24]
    • Colaspoides rufa Gressitt & Kimoto, 1961
    • Colaspoides rufipes Kimoto & Gressitt, 1982[3]
    • Colaspoides rufiventris Medvedev, 2006[26]
    • Colaspoides rufofulva Medvedev, 2004[6]
    • Colaspoides rugipennis (Motschulsky, 1860)[62]
    • Colaspoides rugipennis Lefèvre, 1893[39] (junior homonym of above)
    • Colaspoides rugulosus Lefèvre, 1889
    • Colaspoides sabahensis Medvedev, 2010[24]
    • Colaspoides sandakana Medvedev, 2010[24]
    • Colaspoides sarawacensis Medvedev, 2010[24]
    • Colaspoides sarrameae Jolivet, Verma & Mille, 2008[42]
    • Colaspoides sauteri Chûjô, 1956
    • Colaspoides schawalleri Medvedev, 2010[24]
    • Colaspoides schulzi Medvedev, 2007[63]
    • Colaspoides sculpturata Medvedev, 2006[26]
    • Colaspoides semipicea Jacoby, 1895[64]
    • Colaspoides serratipes Medvedev, 2013[65]
    • Colaspoides seticornis Medvedev, 2004[6]
    • Colaspoides shapaensis Medvedev, 2004[6]
    • Colaspoides shuteae Medvedev, 2010[24]
    • Colaspoides siamensis Jacoby, 1905
    • Colaspoides similis Lea, 1915[23]
    • Colaspoides simillima Baly, 1867[30]
    • Colaspoides simplicipennis Jacoby, 1885
    • Colaspoides speciosa Lefèvre, 1887
    • Colaspoides staudingeri Medvedev, 2010[24]
    • Colaspoides striatopunctata (Boisduval, 1835)
    • Colaspoides suavis Lea, 1915[23]
    • Colaspoides subfasciata Medvedev, 2019[29]
    • Colaspoides sublaevicollis Duvivier, 1892
    • Colaspoides submetallica Medvedev, 2010[24]
    • Colaspoides subovata Medvedev, 2004[6]
    • Colaspoides subtuberculata Medvedev, 2006[26]
    • Colaspoides suginoi Komiya, 1988
    • Colaspoides sulawensis Medvedev, 2008[32]
    • Colaspoides sumatrensis Jacoby, 1884
    • Colaspoides taiwanus Chûjô, 1956
    • Colaspoides takizawai Medvedev, 2010[24]
    • Colaspoides tamdaoensis Medvedev, 2004[6]
    • Colaspoides tarsalis Lea, 1915[23]
    • Colaspoides tenenbaumi Pic, 1942
    • Colaspoides tenuicornis Medvedev, 2010[24]
    • Colaspoides thailandicus Kimoto & Gressitt, 1982[3]
    • Colaspoides tridentata Medvedev, 2004[6]
    • Colaspoides trusmadiensis Medvedev & Romantsov, 2014[28]
    • Colaspoides tuberculata Baly, 1867[30]
    • Colaspoides varians Baly, 1867[30]
    • Colaspoides ventralis Medvedev, 2018[66]
    • Colaspoides venusta Lefèvre, 1889
    • Colaspoides vietnamicus Kimoto & Gressitt, 1982[3]
    • Colaspoides violacea Baly, 1867[30]
    • Colaspoides viridana Baly, 1867[30]
    • Colaspoides viridimarginata Baly, 1867[30]
    • Colaspoides viridipennis Weise, 1923[67]
    • Colaspoides viridiventris Medvedev, 2010[24]
    • Colaspoides vitiensis Bryant, 1938[68]
    • Colaspoides vocki Medvedev, 2007[63]
    • Colaspoides volkovi Medvedev & Romantsov, 2014[28]
    • Colaspoides weisei Medvedev, 2006[26]
    • Colaspoides yunnanicus Medvedev, 2004[6]
    • Colaspoides zoiai Medvedev, 2004[6]

    Renamed species:

    • Colaspoides malayana Medvedev, 1998 (preoccupied by C. malayana Jacoby, 1894): renamed to Colaspoides malayensis Medvedev, 2004[6]
    • Colaspoides pallidula Lea, 1915[23] (preoccupied by C. pallidula Jacoby, 1889): renamed to Colaspoides leai Medvedev, 2001[46]
    • Colaspoides piceus Kimoto & Gressitt, 1982[3] (preoccupied by C. picea Baly, 1867): renamed to Colaspoides piceana Medvedev, 2004[6]
    • Colaspoides purpurata Medvedev & Takizawa, 2011[69] (preoccupied by C. purpurata Medvedev, 2006) renamed to Colaspoides purpurascens Medvedev, 2015[61]
    • Colaspoides tarsalis Chen, 1935 (preoccupied by C. tarsalis Lea, 1915): renamed to Colaspoides paddis Aslam, 1968[56]
    • Colaspoides tibialis Lefèvre, 1875 (preoccupied by C. tibialis (Germar, 1824)): renamed to Colaspoides tibiella Bechyné, 1953[11]

    Synonyms:

    • Colaspoides discoidea Lefèvre, 1891: synonym of Colaspoides viridicollis Jacoby, 1878[70]
    • Colaspoides fruhstorferi Jacoby, 1898:[25] synonym of Colaspoides robusta Baly, 1867 [24]
    • Colaspoides imasakai yakuanus Komiya, 1991: synonym of Colaspoides imasakai Komiya, 1991[55]
    • Colaspoides opulenta Jacoby, 1900:[12] synonym of Colaspoides fasciata Lefèvre, 1875[71]
    • Colaspoides paviei Lefèvre, 1890: synonym of Colaspoides rugipennis (Motschulsky, 1860)[62]
    • Colaspoides phalerata Weise, 1922:[45] synonym of Colaspoides philippinensis Baly, 1867[26]
    • Colaspoides subrugosa Jacoby, 1908:[37] synonym of Colaspoides bengalensis Duvivier, 1892[72]

    Species moved to Beltia:[8]

    • Colaspoides chanchamayensis Bechyné, 1950:[73] now a synonym of Beltia weyrauchi (Bechyné, 1950)
    • Colaspoides chiriquensis Jacoby, 1882:[19] now Beltia chiriquensis (Jacoby, 1882)
    • Colaspoides placidula Bechyné, 1950:[73] now Beltia placidula (Bechyné, 1950)
    • Colaspoides placidula angustomarginata Bechyné, 1953:[11] now Beltia angustomarginata (Bechyné, 1953)
    • Colaspoides turialbana Bechyné, 1950:[74] now a synonym of Beltia chiriquensis (Jacoby, 1882)
    • Colaspoides weyrauchi Bechyné, 1950:[73] now Beltia weyrauchi (Bechyné, 1950)

    References[edit]

    1. ^ a b "Colaspoides Laporte 1833". Paleobiology Database. Fossilworks. Retrieved 25 June 2018.
    2. ^ Laporte, F. L. N. de Caumont (1833). "Mémoire sur les divisions du genre Colaspis". Revue Entomologique. 1: 18–25.
    3. ^ a b c d e f g h i j Kimoto, S.; Gressitt, J. L. (1982). "Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos and Vietnam. III. Eumolpinae" (PDF). Esakia. 18: 1–141. doi:10.5109/2421. hdl:2324/2421. S2CID 83265328.
    4. ^ a b Chapuis, F. (1874). "Tome dixième. Famille des phytophages". In Lacordaire, J.T.; Chapuis, F. (eds.). Histoire naturelle des Insectes. Genera des coléoptères. Vol. 10. Paris: Librairie Encyclopédique de Roret. pp. i–iv, 1–455.
    5. ^ a b Lefèvre, É. (1885). "Eumolpidarum hucusque cognitarum catalogus, sectionum conspectu systematico, generum sicut et specierum nonnullarum novarum descriptionibus adjunctis". Mémoires de la Société Royale des Sciences de Liège. 2. 11 (16): 1–172.
    6. ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc Medvedev, L.N. (2004). "Revision of the genus Colaspoides Laporte, 1833 (Chrysomelidae: Eumolpinae) from continental Asia" (PDF). Russian Entomological Journal. 12 (3): 257–297.
    7. ^ a b Moseyko, Alexey G.; Kirejtshuk, Alexander G. (2013). "On the taxonomic position of new taxa of the subfamily Eumolpinae (Coleoptera, Chrysomelidae) from Baltic amber". Terrestrial Arthropod Reviews. 6 (1–2): 163–172. doi:10.1163/18749836-06021063.
    8. ^ a b Flowers, R. Wills (2018). "A review of the genus Beltia Jacoby (Chrysomelidae: Eumolpinae: Eumolpini), with descriptions of fourteen new species from Costa Rica, Panama, and northwestern South America". Insecta Mundi (672): 1–43.
    9. ^ Clavareau, H. (1914). "Chrysomelidae: 11. Eumolpinae". In Junk, W.; Schenkling, S. (eds.). Coleopterorum Catalogus. Vol. 59. Berlin: W. Junk. pp. 1–215.
    10. ^ Chaboo, Caroline S.; Flowers, R. Wills (2015). "Beetles (Coleoptera) of Peru: A Survey of the Families. Chrysomelidae: Eumolpinae Hope, 1840". Journal of the Kansas Entomological Society. 88 (3): 375–379. doi:10.2317/kent-88-03-375-379.1. S2CID 87900852.
    11. ^ a b c d Bechyné, J. (1953). "Katalog der neotropischen Eumolpiden (Col. Phytoph. Chrysomeloidea)". Entomologische Arbeiten aus dem Museum G. Frey (in German). 4: 26–303.
    12. ^ a b Jacoby, M. (1900). "Descriptions of New Species and a New Genus of South American Eumolpidæ, with Remarks on some of the Genera". Transactions of the Entomological Society of London. 48 (4): 453–510. doi:10.1111/j.1365-2311.1900.tb02719.x.
    13. ^ Jacoby, M. (1881). "Descriptions of new genera and species of phytophagous Coleoptera". Proceedings of the Zoological Society of London. 1881 (2): 439–450. doi:10.1111/j.1096-3642.1881.tb01300.x.
    14. ^ a b c Jacoby, M. (1879). "Descriptions of new species of Phtytophaga (Coleoptera)". Proceedings of the Zoological Society of London. 1879: 773–793.
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