Charis is a genus of the Riodininitribe of metalmark butterflies (familyRiodinidae). Nineteen species have been identified within the Charis cleonus complex[3] (however, further analysis has revised this clade to contain 22 species[4]) and eight species within the Charis gynaea group (clade).[5]Charis butterflies are common in the Neotropics and often live in primary and secondary growth.[5] The Charis cleonus group exhibits contemporary parapatric distributions throughout Amazonia and are thought to have speciated allopatrically; residing in "areas of endemism".[3] Some evidence suggests that Charis (among other Riodininae groups) are reproductively isolated by mating preferences for different topographic areas and different times—of which, may have promoted speciation between the various groups.[6]
^Jason P. W. Hall and Donald J. Harvey (2002), "Phylogenetic revision of the Charis cleonus complex (Lepidoptera: Riodinidae)", Systematic Entomology, 27: 265–300, doi:10.1046/j.1365-3113.2002.00173.x Full text: [1]
^ abJason P. W. Hall and Donald J. Harvey (2001), "Phylogenetic Revision of the Charis gynaea Group (Lepidoptera: Riodinidae) with Comments on Historical Relationships Among Neotropical Areas of Endemism", Annals of the Entomological Society of America, 94 (5): 631–647, doi:10.1603/0013-8746(2001)094[0631:protcg]2.0.co;2
^Curtis J. Callaghan (1983), "A study of isolating mechanisms among neotropical butterflies of the subfamily Riodininae", Journal of Research on the Lepidoptera, 21 (3): 159–176