This is a list of fossil primates—extinct primates for which a fossil record exists. Primates are generally thought to have evolved from a small, unspecialized mammal , which probably fed on insects and fruits. However, the precise source of the primates remains controversial and even their arboreal origin has recently been questioned.[ 1] As it has been suggested, many other mammal orders are arboreal too, but they have not developed the same characteristics as primates. Nowadays, some well known genera, such as Purgatorius and Plesiadapis , thought to be the most ancient primates for a long time, are not usually considered as such by recent authors, who tend to include them in the new order Plesiadapiformes , within superorder Euarchontoglires .[ 2] Some, to avoid confusions, employ the unranked term Euprimates , which excludes Plesiadapiformes.[ 3] That denomination is not used here.
There is an academic debate on the time the first primates appeared. One of the earliest probable primate fossils is the problematic Altiatlasius koulchii , perhaps an Omomyid , but perhaps a non-Primate Plesiadapiform , which lived in Morocco, during the Paleocene , around 60 Ma .[ 1] However, other studies, including molecular clock studies, have estimated the origin of the primate branch to have been in the mid-Cretaceous period, around 85 Ma, that is to say, in the time previous to the extinction of dinosaurs and the successful mammal radiation.[ 4] [ 5] [ 6] Nevertheless, there seems to be a consensus about the monophyletic origin of the order, although the evidence is not clear.[ 7]
The order Primates, established by Linnaeus in 1758 , includes humans and their immediate ancestors. However, contrarily to the common opinion, most primates do not have especially large brains. Brain size is a derived character, which only appeared with genus Homo , and was lacking in the first hominid . In fact, hominid encephalization quotient is only 1.5 Ma more recent than that of some dolphin species. The encephalization quotient of some cetaceans is therefore higher than that of most primates, including the nearest relatives of humans, such as Australopithecus .[ 8]
This list follows partly from Walter Carl Hartwig 's 2002 book The Fossil Primate Record [ 9] and John G. Fleagle 's 2013 book Primate Adaptation and Evolution (3rd edition).[ 10] Parentheses around authors' names (and dates) indicates a change in generic name for the fossil, as stated in the International Code of Zoological Nomenclature (ICZN).[ 11] Since the publication of the book as well as the creation of this article, new fossil taxon have been discovered that has helped improved the taxonomy among primates in general.
Sulaimanius arifi (Gunnell et al ., 2008)
Smilodectes
Darwinius
Ekgmowechashalidae Szalay, 1976
Bugtilemur Marivaux et al ., 2001
Bugtilemur mathesoni Marivaux et al ., 2001
Ekgmowechashala Macdonald, 1963
Ekgmowechashala philotau Macdonald, 1963
Ekgmowechashala zancanellai Samuels, Albright & Fremd, 2015
Gatanthropus Ni et al. , 2016
Gatanthropus micros Ni et al. , 2016
Muangthanhinius Marivaux et al. , 2006
Muangthanhinius siami Marivaux et al. , 2006
Notharctidae Trouessart, 1879
Notharctinae Trouessart, 1879
Cantius Simons, 1962
Cantius abditus Gingerich & Simmons, 1977
Cantius angulatus Cope, 1875
Cantius antediluvius Kihm, 1992
Cantius eppsi Cooper, 1932
Cantius frugivorus Cope, 1875
Cantius lohseorum Robinson, 2016
Cantius mckennai Gingerich & Simons, 1977
Cantius nuniensis Cope, 1881
Cantius ralstoni Matthew, 1915
Cantius savagei Gingerich, 1977
Cantius torresi Gingerich, 1986
Copelemur Gingerich & Simons, 1977
Copelemur australotutus Beard, 1988
Copelemur praetutus Gazin, 1962
Copelemur tutus Cope, 1877
Hesperolemur Gunnell, 1995
Hesperolemur actius Gunnell, 1995
Megaceralemur Robinson, 2016
Megaceralemur trigonodus (Matthew, 1915)
Megaceralemur matthewi Robinson, 2016
Notharctus Leidy, 1870
Notharctus pugnax Granger & Gregory, 1917
Notharctus robustior Leidy, 1870
Notharctus tenebrosus Leidy, 1870
Notharctus venticolus Osborn, 1902
Pelycodus Cope, 1875
Pelycodus danielsae Froehlich & Lucas, 1991
Pelycodus jarrovii Cope, 1874
Pinolophus Robinson, 2016
Pinolophus meikei Robinson, 2016
Smilodectes Wortman, 1903
Smilodectes gingerichi Beard, 1988
Smilodectes gracilis Marsh, 1871
Smilodectes mcgrewi Gingerich, 1979
Cercamoniinae Gingerich, 1975
Agerinia Crusafont-Pairo & Golpe-Posse, 1973
Agerinia marandati Femenias-Gual, Minwer-Barakat, Marigó, Poyatos-Moré, and Moyà-Solà, 2017
Agerinia roselli Crusafont-Pairo & Golpe-Posse, 1973
Agerinia smithorum Femenias-Gual, Minwer-Barakat, Marigó, and Moyà-Solà, 2016
Anchomomys Stehlin, 1916
Anchomomys crocheti Godinot, 1988
Anchomomys gaillardi Stehlin, 1916
Anchomomys pygmaeus Rütimeyer, 1890
Anchomomys quercy Stehlin, 1916
Barnesia Thalmann, 1994
Barnesia hauboldi Thalmann, 1994
Buxella Godinot, 1988
Buxella magna Godinot, 1988
Buxella prisca Godinot, 1988
Donrussellia Szalay, 1976
Donrussellia gallica Russell, Louis & Savage, 1967
Donrussellia louisi
Donrussellia magna
Donrussellia provincialis
Donrussellia russelli
Mazateronodon Marigó, Minwer-Barakat, & Moyà-Solà, 2010
Mazateronodon endemicus Marigó, Minwer-Barakat, & Moyà-Solà, 2010
Panobius Russell & Gingerich, 1987
Panobius afridi Russell & Gingerich, 1987
Periconodon Stehlin, 1916
Periconodon helleri Schwartz et al., 1983
Periconodon helveticus Rütimeyer, 1891
Periconodon huerzeleri Gingerich, 1977
Periconodon jaegeri Godinot, 1988
Periconodon lemoinei Gingerich, 1977
Protoadapis Lemoine, 1878
Protoadapis angustidens Filhol, 1888
Protoadapis brachyrhynchus Stehlin, 1912
Protoadapis curvicuspidens Lemoine, 1878
Protoadapis ignoratus Thalmann, 1994
Protoadapis muechelnensis Thalmann, 1994
Protoadapis recticuspidens Lemoine, 1878
Protoadapis weigelti Gingerich, 1977
Pronycticebus Grandidier, 1904
Pronycticebus gaudryi Grandidier, 1904
Pronycticebus neglectus Thalmann et al. , 1989
Asiadapidae Rose et al ., 2009
Anthrasimias Bajpai et al. , 2008
Anthrasimias gujaratensis Bajpai et al. , 2008
Asiadapis Rose et al ., 2007
Asiadapis cambayensis Rose et al ., 2007
Asiadapis tapiensis Rose et al. , 2018 [ 12]
Marcgodinotius Bajpai et al ., 2005
Marcgodinotius indicus Bajpai et al ., 2005
Adapidae Trouessart, 1879
Adapinae Trouesart, 1879
Adapis Cuvier, 1821
Adapis bruni
Adapis collinsonae Hooker, 1986
Adapis parisiensis de Blainville, 1841
Adapis sudrei Gingerich, 1977
Cryptadapis Godinot, 1984
Cryptadapis laharpei Godinot, 1984
Cryptadapis tertius Godinot, 1984
Leptadapis Gervais, 1876
Leptadapis assolicus
Leptadapis filholi Godinot & Couette, 2008
Leptadapis capellae Crusafont-Pairo, 1967
Leptadapis leenhardti Stehlin, 1912
Leptadapis magnus Filhol, 1874
Leptadapis ruetimeyeri Stehlin, 1912
Magnadapis Godinot & Couette, 2008
Magnadapis quercyi Godinot & Couette, 2008
Magnadapis fredi Godinot & Couette, 2008
Magnadapis laurenceae Godinot & Couette, 2008
Magnadapis intermedius Godinot & Couette, 2008
Microadapis Szalay, 1974
Microadapis lynnae
Microadapis sciureus Stehlin, 1916
Palaeolemur Delfortrie, 1873
Palaeolemur betillei Delfortrie, 1873
Paradapis Tattersall & Schwartz 1983
Paradapis ruetimeyeri Stehlin 1912
Paradapis priscus Stehlin, 1916
Caenopithecinae Szalay & Delson 1979
Adapoides Beard et al. , 1994
Adapoides troglodytes Beard et al. , 1994
Afradapis Seiffert et al ., 2009
Afradapis longicristatus Seiffert et al ., 2009
Aframonius Simons et al. , 1995
Aframonius diedes Simons et al. , 1995
Caenopithecus Rütimeyer, 1862
Caenopithecus lemuroides Rütimeyer, 1862
Darwinius Franzen et al. , 2009
Darwinius masillae Franzen et al. , 2009
Europolemur Weigelt, 1933
Europolemur dunaifi Tattersall & Schwartz, 1983
Europolemur klatti Weigelt, 1933
Europolemur koenigswald Franzen, 1987
Godinotia Franzen, 2000
Godinotia neglecta Thalmann, Haubold & Martin, 1989
Mahgarita Wilson & Szalay, 1976
Mahgarita stevensi Wilson & Szalay, 1976
Masradapis Seiffert, Boyer, Fleagle, Gunnell, Heesy, Perry, Sallam, 2017
Masradapis tahai Seiffert, Boyer, Fleagle, Gunnell, Heesy, Perry, and Sallam, 2017
Mescalerolemur Kirk & Williams, 2011
Mescalerolemur horneri Kirk & Williams, 2011
Sivaladapidae Thomas & Verma, 1979
Ramadapis Gilbert, Patel, Singh, Campisano, Fleagle, Rust, and Patnaik, 2017
Ramadapis sahnii Gilbert, Patel, Singh, Campisano, Fleagle, Rust, and Patnaik, 2017
Sivaladapinae Thomas & Verma, 1979
Indraloris Lewis, 1933
Indraloris himalayensis Pilgrim, 1932
Indraloris kamlialensis Flynn and Morgan, 2005
Sinoadapis Wu & Pan, 1985
Sinoadapis carnosus Wu & Pan, 1985
Sivaladapis Gingerich & Sahni, 1979
Sivaladapis nagrii Prasad, 1970
Sivaladapis palaendicus Pilgrim, 1932
Hoanghoniinae Gingerich et al ., 1994
Hoanghonius Zdansky, 1930
Hoanghonius stehlini Zdansky, 1930
Lushius Chow, 1961
Lushius qinlinensis Chow, 1961
Rencunius Gingerich et al. , 1994
Rencunius zhoui Gingerich et al. , 1994
Wailekia Ducrocq et al. , 1995
Wailekia orientale Ducrocq et al. , 1995
incertae sedis
Guangxilemur Qi & Beard, 1998
Guangxilemur tongi Qi & Beard, 1998
Kyitchaungia Beard et al . 2007
Kyitchaungia takaii Beard et al . 2007
Laomaki Ni et al. 2016
Laomaki yunnanensis Ni et al. 2016
Paukkaungia Beard et al. 2007
Paukkaungia parva Beard et al. 2007
Siamoadapis Chaimanee et al. , 2007
Siamoadapis maemohensis Chaimanee et al. , 2007
Yunnanadapis Ni et al. 2016
Yunnanadapis folivorus Ni et al. 2016
Yunnanadapis imperator Ni et al. 2016
Plesiopithecus teras Simons, 1992
Propotto leakeyi Simpson, 1967
Algeripithecus minutus Godinot & Mahboubi, 1992
Azibius trerki Sudre, 1975
Djebelemur martinezi Hartenberger & Marandat, 1992
Namaia bogenfelsi Pickford et al ., 2008
Omanodon minor Gheerbrant et al. , 1993
Shizarodon dhofarensis Gheerbrant et al. , 1993
Plesiopithecus teras Simons, 1992
Subfossil lemurs :
Babakotia radofilai
Archaeolemur edwardsi Filhol, 1895
Archaeolemur majori Filhol, 1895
Hadropithecus stenognathus Lorenz von Liburnau, 1899
Mesopropithecus dolichobrachion Simons et al. , 1995
Mesopropithecus globiceps Lamberton, 1936
Mesopropithecus pithecoides Standing, 1905
Babakotia radofilai Godfrey et al. , 1990
Palaeopropithecus ingens G. Grandidier, 1899
Palaeopropithecus kelyus Gommery et al., 2010
Palaeopropithecus maximus Standing, 1903
Archaeoindris fontoynontii Standing, 1909
Megaladapis (Megaladapis) grandidieri Standing, 1903
Megaladapis (Megaladapis) madagascariensis Forsyth-Major, 1894
Megaladapis (Peloriadapis) edwardsi Grandidier, 1899
Pachylemur insignis Filhol, 1895
Pachylemur jullyi Lamberton, 1948
Karanisia clarki Seiffert et al., 2003
Mioeuoticus bishopi Leakey, 1962
Mioeuoticus kichotoi Kunimatsu, Tsujikawa, Nakatsukasa, Shimizu, Ogihara, Kikuchi, Nakano, Takano, Morimoto, and Ishida, 2017 [ 19]
Mioeuoticus shipmani Phillips & Walker, 2000
Nycticeboides simpsoni Jacobs, 1981
Komba minor Le Gros Clark & Thomas, 1952
Komba robustus Le Gros Clark & Thomas, 1952
Komba winamensis McCrossin, 1992
Progalago dorae MacInnes, 1943
Progalago songhorensis Simpson, 1967
Saharagalago misrensis Seiffert et al ., 2003
Wadilemur elegans Simons, 1997 [ 17]
Teilhardina asiatica Ni et al. , 2004
Teilhardina belgica (Teilhard de Chardin, 1927)
Altanius orlovi Dashzeveg & McKenna, 1977
Altiatlasius koulchii Sigé et al. , 1990
An artist's reconstruction of Archicebus achilles .
Archicebus achilles Ni et al., 2013 [ 20]
Baataromomys ulaanus Ni, Beard, Meng, Wang, and Gebo, 2007
Kohatius coppensi Russell & Gingerich, 1980
Melaneremia bryanti Hooker, 2007
Microchoerus creechbarrowensis Hooker, 1986
Microchoerus edwardsi Filhol, 1880
Microchoerus erinaceus Wood, 1846
Microchoerus hookeri Minwer-Barakat, Marigó, Femenias-Gual, Costeur, Esteban-Trivigno, and Moyà-Solà, 2017
Microchoerus ornatus Stehlin, 1916
Microchoerus wardi Hooker, 1986
Necrolemur anadoni Minwer-Barakat, Marigó & Moyà-Solà, 2015
Necrolemur antiquus Filhol, 1873
Necrolemur zitteli Schlosser, 1887
Nannopithex filholi Chantre & Gaillard, 1897
Nannopithex humilidens Thalmann, 1994
Nannopithex quaylei Hooker, 1986
Nannopithex raabi Heller, 1930
Nannopithex zuccolae Godinot et al. , 1992
Pseudoloris crusafonti Louis & Sudre, 1975
Pseudoloris godinoti Köhler & Moyà-Solà, 1999
Pseudoloris isabenae Crusafont-Pairo, 1967
Pseudoloris parvulus Filhol, 1890
Anaptomorphus aemulus Cope, 1872
Anaptomorphus westi Szalay, 1976
Tetonius homunculus Cope, 1882
Tetonius matthewi Bown & Rose, 1987
Tetonius mckennai Bown & Rose, 1987
Absarokius abbotti Loomis, 1906
Absarokius australis Bown & Rose, 1987
Absarokius nocerai Robinson, 1966
Absarokius metoecus Bown & Rose, 1987
Absarokius witteri Morris, 1954
Teilhardina brandti Gingerich, 1993
"Teilhardina" demissa Rose, 1995
"Teilhardina" gingerichi Rose, Chew, Dunn, Kraus, Fricke, and Zack, 2012
"Teilhardina" tenuicula Jepsen, 1930
Bownomomys americanus (Bown, 1976)
Bownomomys crassidens )Bown & Rose, 1987)
Anemorhysis natronensis Beard et al. , 1992
Anemorhysis pattersoni Bown & Rose, 1984
Anemorhysis pearcei Gazin, 1962
Anemorhysis savagei Williams & Covert, 1994
Anemorhysis sublettensis Gazin, 1952
Anemorhysis wortmani Bown & Rose, 1984
Chlororhysis incomptus Bown & Rose, 1984
Chlororhysis knightensis Gazin, 1958
Pseudotetonius ambiguus Bown, 1974
Arapahovius advena Bown & Rose, 1991
Arapahovius gazini Savage & Waters, 1978
Aycrossia lovei Bown, 1979
Strigorhysis bridgerensis Bown, 1979
Strigorhysis huerfanensis Bown & Rose, 1987
Strigorhysis rugosus Bown, 1979
Gazinius amplus Bown, 1979
Gazinius bowni Gunnell, 1995
Tatmanius szalayi Bown & Rose, 1991
Trogolemur amplior Beard et al. , 1992
Trogolemur fragilis Beard et al. , 1992
Trogolemur myodes Matthew, 1909
Sphacorhysis burntforkensis Gunnell, 1995
Walshina esmaraldensis López-Torres, Silcox, and Holroyd, 2018
Walshina mcgrewi (Robinson, 1968)
Walshina shifrae (Krishtalka, 1978)
Brontomomys cerutti Atwater and Kirk, 2018
Diablomomys dalquesti Williams and Kirk, 2008
Ekwiiyemakius walshi Atwater and Kirk, 2018
Gunnelltarsius randalli Atwater and Kirk, 2018
Omomys carteri Leidy, 1869
Omomys lloydi Gazin, 1958
Steinius annectens Bown & Rose, 1991
Steinius vespertinus Matthew, 1915
Chumashius balchi Stock, 1933
Washakius insignis Leidy, 1873
Washakius izetti Honey, 1990
Washakius laurae Simpson, 1959
Washakius woodringi Stock, 1938
Shoshonius bowni Honey, 1990
Shoshonius cooperi Granger, 1910
Dyseolemur pacificus Stock, 1934
Loveina minuta Loomis, 1906
Loveina wapitiensis Gunnell et al. , 1992
Loveina zephyri Simpson, 1940
Utahia carina Muldoon and Gunnell, 2002
Utahia kayi Gazin, 1958
Stockia powayensis Gazin, 1958
Chipetaia lamporea Rasmussen, 1996
Asiomomys changbaicus Wang & Li, 1990
Wyomomys bridgeri Gunnell, 1995
Ageitodendron matthewi Gunnell, 1995
Ourayia hopsoni Robinson, 1968
Ourayia uintensis Osborn, 1895
Macrotarsius jepseni Robinson, 1968
Macrotarsius macrorhysis Beard et al. , 1994
Macrotarsius montanus Clark, 1941
Macrotarsius roederi Kelly, 1990
Macrotarsius siegerti Robinson, 1968
Hemiacodon casamissus Beard et al. , 1992
Hemiacodon gracilis Marsh, 1872
Yaquius travisi Mason, 1990
Uintanius ameghini Wortman, 1904
Uintanius rutherfurdi Robinson, 1966
Jemezius szalayi Beard, 1987
Rooneyia viejaensis Wilson, 1966
Hesperotarsius sindhensis Zijlstra, Flynn, and Wessels, 2013
Hesperotarsius thailandicus (Ginsburg & Mein, 1987)
Oligotarsius rarus Ni et al. , 2016
"Tarsius" eocaenus Beard et al. , 1994
"Tarsius" sirindhornae Chaimanee et al. , 2011 [ 25]
Xanthorhysis tabrumi Beard, 1998
Afrasia djijidae Chaimanee et al. 2012
Afrotarsius chatrathi Simons & Bown, 1985
Afrotarsius libycus Jaeger et al ., 2010
Eosimias centennicus Beard et al. , 1996
Eosimias sinensis Beard et al. , 1994
Bahinia banyueae Li et al. , 2016
Bahinia pondaungensis Jaeger et al. , 1999
Phileosimias Marivaux, Antoine, Baqri, Benammi, and Chaimanee, 2005
Phileosimias brahuiorum Marivaux, Antoine, Baqri, Benammi, and Chaimanee, 2005
Phileosimias kamali Marivaux, Antoine, Baqri, Benammi, and Chaimanee, 2005
Phenacopithecus krishtalkai Beard and Wang, 2004
Phenacopithecus xueshii Beard and Wang, 2004
Pondaungia cotteri Pilgrim, 1927
Amphipithecus mogaungensis Colbert, 1937
Krabia minuta Chaimanee et al. , 2013
Siamopithecus eocaenus Chaimanee et al. , 1997
Proteopithecus sylviae Simons, 1989
Serapia eocaena Simons, 1992
Arsinoea kallimos Simons, 1992
Apidium bowni Simons, 1995
Apidium moustafai Simons, 1962
Apidium phiomense Osborn, 1908
Parapithecus fraasi Schlosser, 1910
Parapithecus grangeri Simons, 1974
Qatrania fleaglei Simons & Kay, 1988
Qatrania wingi Simons & kay, 1983
Biretia piveteaui Bonis et al. , 1988
Biretia fayumensis Seiffert et al. , 2005
Biretia megalopsis Seiffert et al. , 2005
Platyrrhini , incertae sedis
Branisella Hoffstetter, 1969
Branisella boliviana Hoffstetter, 1969
Atelidae Gray, 1825
Pitheciinae Mivart, 1865
Tribus: Callicebini
Xenothrix Williams & Koopman, 1952
Xenothrix mcgregori Williams & Koopman, 1952
Antillothrix MacPhee et al. , 1995
Antillothrix bernensis Rímoli, 1977
Paralouatta Rivero & Arredondo, 1991
Paralouatta varonai Rivero & Arredondo, 1991
Paralouatta marianae
Tribus: Pitheciini
Soriacebus Fleagle et al. , 1987
Soriacebus adrianae Fleagle, 1990
Soriacebus ameghinorum Fleagle et al. , 1987
Proteropithecia Kay et al. , 1999
Proteropithecia neuquenensis Kay et al. , 1998
Cebupithecia Stirton & Savage, 1951
Cebupithecia sarmientoi Stirton & Savage, 1951
Nuciruptor Meldrum & Kay, 1997
Nuciruptor rubricae Meldrum & Kay, 1997
Tribus: Homunculini
Homunculus Ameghino, 1891
Homunculus patagonicus Ameghino, 1891
Carlocebus Fleagle, 1990
Carlocebus carmenensis Fleagle, 1990
Carlocebus intermedius Fleagle, 1990
Atelinae Gray, 1825
Tribus: Alouattini
Stirtonia Hershkovitz, 1970
Stirtonia tatacoensis Stirton, 1951
Stirtonia victoriae Kay et al. , 1987
Tribus: Atelini
Caipora Cartelle & Hartwig, 1996
Caipora bambuiorum Cartelle & Hartwig, 1996
Atelinae , incertae sedis
Cebidae Bonaparte, 1831
Cebinae Bonaparte, 1831
Tribus: Saimiriini
Neosaimiri Stirton, 1951
Neosaimiri fieldsi Stirton, 1951
Laventiana Rosenberger et al. , 1991
Laventiana annectens Rosenberger et al. , 1991
Dolichocebus Kraglievich, 1951
Dolichocebus gaimanensis Kraglievich, 1951
Cebinae , incertae sedis
Chilecebus Flynn & al, 1995
Chilecebus carrascoensis Flynn & al, 1995
Killikaike Tejedor et al., 2006
Killikaike blakei Tejedor et al., 2006
Aotinae Elliot, 1913
Aotus Illiger, 1811
Aotus dindensis Setoguchi & Rosenberger, 1987
Aotinae , incertae sedis
Tremacebus Hershkovitz, 1974
Tremacebus harringtoni Rusconi, 1933
Callitrichinae Thomas, 1903
Tribus: Callimiconi
Mohanamico Luchterhand et al. , 1986
Mohanamico hershkovitzi Luchterhand et al. , 1986
Callitrichinae , incertae sedis
Patasola Kay & Meldrum, 1997
Patasola magdalenae Kay & Meldrum, 1997
Lagonimico Kay, 1994
Lagonimico conclutatus Kay, 1994
Micodon Setoguchi & Rosenberger, 1985
Micodon kiotensis Setoguchi & Rosenberger, 1985
Catarrhini, incertae sedis [ edit ]
Limnopithecus evansi MacInnes, 1943
Limnopithecus legetet Hopwood, 1933
Kalepithecus songhorensis Andrews, 1978
Kalepithecus kogolensis Pickford et al. , 2017
Kamoyapithecus hamiltoni Madden, 1980
Kogolepithecus morotoensis Pickford et al. , 2003
Catopithecus browni Simons, 1989
Oligopithecus rogeri Gheerbrant et al. , 1995
Oligopithecus savagei Simons, 1962
Talahpithecus parvus Jaeger et al., 2010
Moeripithecus markgrafi Schlosser, 1910
Propliopithecus ankeli Simons et al. , 1987
Propliopithecus chirobates Simons, 1965
Propliopithecus haeckeli Schlosser, 1910
Aegyptopithecus zeuxis Simons, 1965
Pliopithecidae Zapfe, 1960
Lomorupithecus Rossie and MacLatchy, 2006
Dionysopithecinae
Dionysopithecus Li, 1978
Dionysopithecus orientalis Suteethorn et al. , 1990
Dionysopithecus shuangouensis Li, 1978
Platodontopithecus Li, 1978
Platodontopithecus jianghuaiensis Li, 1978
Pliopitheciinae
Epipliopithecus Zapfe & Hurzeler, 1957
Pliopithecus Gervais, 1849
Pliopithecus antiquus Gervais, 1849
Pliopithecus piveteaui Hürzeler, 1954
Pliopithecus platyodon Bidermann, 1863
Pliopithecus zhanxiangi Harrison et al. , 1991
Egarapithecus Moyà-Solà et al. , 2001
Egarapithecus narcisoi Moyà-Solà et al. , 2001
Crouzeliinae Ginsburg & Mein, 1980
Plesiopliopithecus Zapfe, 1961
Plesiopliopithecus auscitanensis Bergounioux & Crouzel, 1965
Plesiopliopithecus lockeri Zapfe, 1961
Plesiopliopithecus priensis Welcomme et al. , 1991
Plesiopliopithecus rhodanica Ginsburg & Mein, 1980
Anapithecus Kretzoi , 1975
Laccopithecus Wu & Pan, 1984
Laccopithecus robustus Wu & Pan, 1984
Paidopithex rhenanus Pohlig, 1895
Dendropithecus macinnesi Le Gros Clark & Leakey, 1950
Dendropithecus ugandensis Pickford et al. , 2010
Micropithecus clarki Fleagle & Simons, 1978
Micropithecus leakeyorum Harrison, 1989
Simiolus andrewsi Harrison, 2010
Simiolus cheptumoae Pickford & Kunimatsu, 2005
Simiolus enjiessi Leakey & Leakey, 1987
Simiolus minutus Rossie & Hill, 2018 [ 26] [ 27]
Proconsul
Nyanzapithecus alesi Nengo, Tafforeau, Gilbert, Fleagle, Miller, Feibel, Fox, Feinberg, Pugh, Berruyer, Mana, Engle, Spoor, 2017
Nyanzapithecus harrisoni Kunimatsu, 1997
Nyanzapithecus pickfordi Harrison, 1986
Nyanzapithecus vancouveringorum Andrews, 1974
Homo habilis
Pliobatidae Alba et al. , 2015
Afropithecidae Begun, 2002
Hominidae Gray, 1825
Kenyapithecinae Leakey, 1962
Ponginae Elliot, 1913
Sivapithecini
Lufengpithecini
Homininae Gray, 1825
Dryopithecini Gregory & Hellman, 1939
Gorillini
Hominini
Graecopithecus von Koenigswald, 1972
Sahelanthropus Brunet et al. , 2002
Orrorin Senut et al. , 2001
Ardipithecus White et al. , 1995
Australopithecus Dart, 1925 - paraphyletic in respect to Paranthropus and Homo
Paranthropus Broom, 1938
Homo Linnaeus , 1758
Homo gautengensis Curnoe, 2010
Homo rudolfensis Alexeev, 1986
Homo habilis Leakey et al. , 1964
Homo luzonensis Détroit et al. , 2019
Homo erectus Dubois, 1892
Homo floresiensis P. Brown et al. , 2004
Homo ergaster Groves & Mazak, 1975
Homo antecessor Bermúdez de Castro et al. , 1997
Homo heidelbergensis Schoetensack, 1908
Homo cepranensis Mallegni et al. , 2003
Homo neanderthalensis King, 1864
Homo rhodesiensis Woodward, 1921
Homo naledi Berger et al., 2015
Kenyanthropus Leakey et al. , 2001
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By class By subclass or infraclass By order
Afrosoricids (golden moles, otter shrews, and tenrecs)
Artiodactyls (even-toed ungulates)
Carnivorans (cats, dogs, seals, etc.)
Chiropterans (bats)
Cingulates (armadillos)
Dasyuromorphs (quolls, dunnarts, and other Australian carnivores)
Didelphimorphs (opossums)
Diprotodonts (kangaroos, possums, and wombats)
Eulipotyphlans (hedgehogs, moles, and shrews)
Lagomorphs (hares, rabbits, and pikas)
Macroscelids (elephant shrews)
Peramelemorphs (bandicoots and bilbies)
Perissodactyls (odd-toed ungulates)
Pholidotans (pangolins)
Pilosans (anteaters and sloths)
Primates
Rodents
Scandentians (treeshrews)
By suborder, infraorder, or family
By species