Sac spiders Temporal range:
| |
---|---|
Clubiona trivialis | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Chelicerata |
Class: | Arachnida |
Order: | Araneae |
Infraorder: | Araneomorphae |
Family: | Clubionidae Simon, 1878 |
Diversity | |
18 genera, 665 species | |
The sac spiders of the family Clubionidae[a] are nocturnal, sac-building hunting spiders with a near-worldwide distribution. Their sacs, silken retreats in which they hide during the day, are commonly spun between leafs, under bark, below rocks, or on plant terminals.[citation needed]
Although formerly a much larger catch-all taxon, in its current definition the family contains less than 700 described species across 18 genera, of which Clubiona is by far most species-rich, with 528 accepted species As of November 2024[update].[1]
The Clubionidae have a complex taxonomic history. Historically, the family was a large catch-all taxon for a variety of spiders that shared the following morphological and behavioral similarities: having eight eyes arranged in two rows; having conical anterior spinnerets that touched; and being nocturnal wandering predators that build "sacs" to retreat to during the day.[citation needed]
A large number of genera have been transferred from Clubionidae to other families, and several former subfamilies of the Clubionidae are now treated as separate families.[2] The Zoropsidae, to which genera Anachemmis, Lauricius and Liocranoides were transferred, is much more closely related to the lynx spiders of family Oxyopidae than to the remaining Clubionidae.[3]
According to 2023 cladistic research by Siddharth Kulkarni, Hannah M. Wood and Gustavo Hormiga, the remaining Clubionidae remain polyphyletic as a result of the current placement of genus Elaver.[3]
As of November 2024[update], the Clubionidae consist of over 665 species in 18 genera worldwide,[4] with by far the majority of species in genus Clubiona. The following genera are accepted by the World Spider Catalog:[5]
Additionally, the World Spider Catalog considers Carteroniella Strand, 1907 to be a nomen dubium.[6]