2022 In Paleobotany

Template:ToC Right This paleobotany list records new fossil plant taxa that were to be described during the year 2022, as well as notes other significant paleobotany discoveries and events which occurred during 2022.

Algae

Charophytes

Name	Novelty	Status	Authors	Age	Location	Notes
Lamprothamnium elongatum^[1]	Sp. nov	In press	Feist & Floquet	Late Cretaceous	Spain	A charophyte.
Lamprothamnium ovoideum^[1]	Sp. nov	In press	Feist & Floquet	Late Cretaceous	Spain	A charophyte.
Pseudoharrisichara sedanoensis^[1]	Sp. nov	In press	Feist & Floquet	Late Cretaceous	Spain	A charophyte.

Charophyte research

A study on the Paleocene charophyte flora from the South Gobi area in the Junggar Basin (China) and on the Paleogene fossil record of charophytes is published by Cao et al. (2022), who interpret their findings as evidence of the dispersal of charophyte lineages from Asia to Europe in the middle to late Eocene, possibly facilitated by waterbirds.^[2]

Chlorophytes

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Ardeiporella^[3]	Gen. et comb. nov	Valid	Grgasović	Middle Triassic		Bosnia and Herzegovina	A green alga belonging to the group Dasycladales. Genus includes "Oligoporella" karrerioidea Pia (1935).
Earltonella^[4]	Gen. et sp. nov		LoDuca in LoDuca et al.	Silurian (Llandovery)	Earlton Formation	Canada ( Ontario)	A green alga belonging to the group Bryopsidales. Genus includes new species E. fredricksi.
Milanovicella? canadillana^[5]	Sp. nov	In press	Torromé & Schlagintweit	Late Cretaceous (Santonian–Campanian)		Spain	A green alga belonging to the group Dasycladales.
Neophysoporella^[3]	Gen. et comb. nov	Valid	Grgasović	Late Triassic, possibly also Middle Triassic		France	A green alga belonging to the group Dasycladales. Genus includes "Diplopora" lotharingica Benecke (1898), "Physoporella" jomdaensis Flügel & Mu (1982) and "Physoporella" zamparelliae Parente & Climaco (1999).
Protocodium^[6]	Gen. et sp. nov		Chai, Aria & Hua	Ediacaran	Dengying Formation	China	A green alga belonging to the family Codiaceae. Genus includes new species P. sinense.
Succodium luciae^[7]	Sp. nov	Valid	Vachard & Krainer	Permian-Triassic transition		Italy	A green alga belonging to the group Dasycladales.

Lycopodiopsida

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Lepacyclotes ordosensis^[8]	Sp. nov	Valid	Deng in Deng et al.	Middle Triassic (Ladinian)	Tongchuan Formation	China	A member of the family Isoetaceae.
Lycopodicaulis^[9]	Gen. et sp. nov	Valid	Herrera et al.	Early Cretaceous (late Barremian–early Aptian)	Huolinhe Formation	China	A member of the family Lycopodiaceae. Genus includes new species L. oellgaardii.
Multapicifolium^[10]	Gen. et sp. nov	Valid	Edwards, Li & Berry	Early Devonian		China	A member of Protolepidodendrales of uncertain phylogenetic placement. Genus includes new species M. sinense.
Nothostigma sepeensis^[11]	Sp. nov		Spiekermann, Jasper, Guerra-Sommer & Uhl in Spiekermann et al.	Permian (Cisuralian)		Brazil	A member of Lycopodiopsida of uncertain affinities.
Omprelostrobus^[12]	Gen. et sp. nov		Liu et al.	Devonian (Famennian)	Wutong Formation	China	A member of Isoetales of uncertain affinities. Genus includes new species O. gigas.
Pleuromeia obovata^[13]	Sp. nov		Deng in Deng et al.	Middle Triassic (Ladinian)	Tongchuan Formation	China	A lycopsid.
Porongodendron^[14]	Gen. et sp. nov	Valid	Prestianni et al.	Carboniferous (Mississippian)		Argentina	An isoetalean lycopsid. Genus includes new species P. minitensis.
Selaginella alata^[15]	Sp. nov		Li & Wang in Li et al.	Cretaceous (Albian-Cenomanian)	Burmese amber	Myanmar	A species of Selaginella.
Selaginella cretacea^[16]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A species of Selaginella.

Lycopsid research

Description of new fossil material of Guangdedendron micrum, providing new information on the morphology of this plant, is published by Gao et al. (2022).^[17]
Xu, Liu & Wang (2022) describe new fossil material of Sublepidodendron grabaui from the Devonian (Famennian) Wutong Formation (China), providing new information on the morphology of the female reproductive organs of this plant.^[18]

Marchantiophyta

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Radula patrickmuelleri^[19]	Sp. nov	Valid	Feldberg, Schäfer-Verwimp & Renner in Feldberg et al.	Cretaceous (Albian-Cenomanian)	Burmese amber	Myanmar	A liverwort, a species of Radula.
Radula tanaiensis^[19]	Sp. nov	Valid	Feldberg, Schäfer-Verwimp & Renner in Feldberg et al.	Cretaceous (Albian-Cenomanian)	Burmese amber	Myanmar	A liverwort, a species of Radula.
Ricciopsis asturicus^[20]	Sp. nov	Valid	Santos et al.	Late Jurassic (Kimmeridgian)	Lastres Formation	Spain	A liverwort belonging to the family Ricciaceae.
Ricciopsis cortaderitaensis^[21]	Gen. et sp. nov	Valid	Savoretti et al.	Middle Triassic		Argentina	A liverwort.
Ricciellites^[21]	Sp. nov	Valid	Savoretti et al.	Middle Triassic		Argentina	A liverwort. Genus includes new species R. unsaltoensis.

Marchantiophyta research

New specimens of Radula heinrichsii, providing new information on the morphology of this liverwort, are described from the Cretaceous Burmese amber by Wang et al. (2022).^[22]

Ferns and fern allies

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Acrostichopteris alcainensis^[23]	Sp. nov	Valid	Skog & Sender	Early Cretaceous (Albian)	Escucha Formation	Spain	A member of the family Hymenophyllaceae.
Coniopteris antarctica^[24]	Sp. nov	In press	Trevisan et al.	Late Cretaceous		Antarctica
Diodonopteris virgulata^[25]	Sp. nov	In press	Zhou et al.	Early Permian		China	A botryopteid fern.
Discosoropteris^[26]	Gen. et 2 sp. nov	In press	Pšenička et al.	Carboniferous (Pennsylvanian)	Kladno Formation	Czech Republic	A leptosporangiate fern. Genus includes new species D. chlupatum and D. zlatkokvacekii.
Dryopterites beishanensis^[27]	Sp. nov	In press	Ren & Sun in Ren et al.	Early Cretaceous	Chijinbao Formation	China	A fern Announced online in 2022 officially published in 2023
Gleichenia nagalingumiae^[28]	Sp. nov		Cantrill et al.	Miocene		Australia	A species of Gleichenia.
Hymenophyllites angustus^[15]	Sp. nov		Li & Wang in Li et al.	Cretaceous (Albian-Cenomanian)	Burmese amber	Myanmar	A member of the family Hymenophyllaceae.
Hymenophyllites kachinensis^[15]	Sp. nov		Li & Wang in Li et al.	Cretaceous (Albian-Cenomanian)	Burmese amber	Myanmar	A member of the family Hymenophyllaceae.
Hymenophyllites setosus^[15]	Sp. nov		Li & Wang in Li et al.	Cretaceous (Albian-Cenomanian)	Burmese amber	Myanmar	A member of the family Hymenophyllaceae.
Microlepia burmasia^[29]	Sp. nov		Long, Wang & Shi in Long et al.	Cretaceous	Burmese amber	Myanmar	A species of Microlepia.
Mikasapteris^[30]	Gen. et sp. nov	Valid	Nishida et al.	Late Cretaceous	Yezo Group	Japan	A probable stem polypod leptosporangiate fern. Genus includes new species M. rothwellii.
Paralophosoria^[31]	Gen. et sp. nov	Valid	Morales-Toledo, Mendoza-Ruiz & Cevallos-Ferriz	Middle Jurassic	Otlaltepec Formation	Mexico	A member of the family Dicksoniaceae. Genus includes new species P. jurassica.
Phyllotheca douroensis^[32]	Sp. nov		Barbosa et al.	Carboniferous (Gzhelian)	Douro Carboniferous Basin	Portugal	A member of Equisetales.
Scolecopteris zhoui^[33]	Sp. nov	In press	Zhang et al.	Early Permian	Taiyuan Formation	China	A member of Marattiales belonging to the family Psaroniaceae.
Wolfeniana^[34]	Nom. nov	Valid	Deshmukh	Devonian	Hampshire Group	United States ( West Virginia)	A member of Stauropteridales; a replacement name for Gillespiea Erwin & Rothwell (1989).

Fern and fern ally research

Pecopteris lativenosa is interpreted as a member of the late Paleozoic marattialean family Psaroniaceae by Li et al. (2022).^[35]

Gnetales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Bassitheca^[36]	Gen. et sp. nov	Valid	Manchester et al.	Late Jurassic	Morrison Formation	United States Utah	A gnetale. Genus includes the species B. hoodiorum.
Dichoephedra^[37]	Gen. et sp. nov	In press	Ren et al.	Early Cretaceous	Chijinbao Formation	China	A member of the family Ephedraceae. Genus includes new species D. beishanensis.

Bennettitales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Dictyozamites barnardi^[38]	Sp. nov	Valid	Saadatnejad	Late Triassic (Rhaetian)	Kalariz Formation	Iran	A member of Bennettitales.
Dictyozamites fakhri^[38]	Sp. nov	Valid	Saadatnejad	Late Triassic (Rhaetian)	Kalariz Formation	Iran	A member of Bennettitales.
Kimuriella^[39]	Gen. et sp. nov	Valid	Pott & Takimoto	Late Jurassic (Oxfordian)	Tochikubo Formation	Japan	A member of Bennettitales. Genus includes new species K. densifolia.
Zamites pateri^[40]	Sp. nov	Valid	Kvaček	Late Cretaceous (Cenomanian)	Peruc–Korycany Formation	Czech Republic

Ginkgophytes

Name	Novelty	Status	Authors	Age	Type locality	Location
Glossophyllum lanceolatum^[41]	Sp. nov		Sun & Deng in Sun et al.	Late Triassic		China
Glossophyllum panii^[41]	Sp. nov		Sun & Deng in Sun et al.	Late Triassic		China
Pseudotorellia baganuriana^[42]	Sp. nov	In press	Nosova & Kostina	Early Cretaceous (Aptian–Albian)	Khuren Dukh Formation	Mongolia
Pseudotorellia zhoui^[43]	Sp. nov	In press	Dong et al.	Middle-Late Jurassic	Daohugou Beds	China
Umaltolepis zhoui^[43]	Sp. nov	In press	Dong et al.	Middle-Late Jurassic	Daohugou Beds	China

Ginkgophyte research

Revision of Ginkgo abaniensis, based on data from leaves from the Jurassic Mura Formation (Russia ), is published by Frolov & Mashchuk (2022), who emend the diagnosis of this species, and transfer Ginkgo abaniensis, Ginkgo glinkiensis and Ginkgo capillata to the genus Ginkgoites.^[44]

Conifers

Araucariaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Agathoxylon argentinum^[45]	Sp. nov	Valid	Bodnar et al.	Late Triassic	Ischigualasto Formation	Argentina

Cheirolepidiaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Brachyoxylon yanqingense^[46]	Sp. nov		Cheng et al.	Late Jurassic	Tuchengzi Formation	China	A probable member of the family Cheirolepidiaceae.
Frenelopsis antunesii^[47]	Sp. nov	In press	Mendes & Kvaček	Early Cretaceous (Aptian–Albian)	Figueira da Foz Formation	Portugal	A member of the family Cheirolepidiaceae.
Pseudofrenelopsis zlatkoi^[48]	Sp. nov		Kvaček & Mendes	Early Cretaceous (Aptian-Albian)	Figueira da Foz Formation	Portugal

Cupressaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Cupressinanthus klebsii^[49]	Sp. nov	Valid	Sadowski, Schmidt & Kunzmann	Eocene		Europe (Baltic Sea region)	Cupressaceous pollen cone.
Patagotaxodia^[50]	Gen. et sp. nov	Valid	Andruchow-Colombo et al.	Late Cretaceous (Maastrichtian)	Lefipán Formation	Argentina	A member of the family Cupressaceae. Genus includes new species P. lefipanensis.

Pinaceae

Name	Novelty	Authors	Age	Type locality	Location	Notes
Keteleerioxylon changchunense^[51]	Sp. nov	Shi, Sun, Meng & Yu in Shi et al.	Early Cretaceous (Albian)	Yingcheng Formation	China	A Keteleeria-like wood morphogenus.
Nothotsuga mulaensis^[52]	Sp. nov	Li & Dong in Dong et al.	Miocene	Changtai Formation	China	A species of Nothotsuga.
Pinus prehwangshanensis^[53]	Sp. nov	Bazhenova, Wu & Jin in Bazhenova et al.	Late Pleistocene	Maoming Basin	China	A pine.
Pinus shengxianica^[54]	Sp. nov	Li, Hu & Xiao in Li et al.	Miocene	Shengxian Formation	China	A pine.

Podocarpaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Phyllocladoxylon antarcticum^[55]	Sp. nov	Announced	Pujana et al.	Oligocene	San José Formation	Chile	A podocarpaceous wood morphospecies Announced in 2022 Officially published in 2023
Podocarpoxylon resinosum^[55]	Sp. nov	Announced	Pujana et al.	Oligocene	San José Formation	Chile	A podocarpaceous wood morphospecies Announced in 2022 Officially published in 2023
Podocarpus mexicanoxylon^[56]	Sp. nov		Castañeda-Posadas	Miocene		Mexico	A species of Podocarpus.

Sciadopityaceae

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Zhangoxylon^[57]	Gen. et sp. nov	In press	Jiang et al.	Middle to Late Jurassic (Callovian to Kimmeridgian)		China	A member of the family Sciadopityaceae. Genus includes new species Z. yanliaoense.

Voltziales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Hexicladia^[58]	Gen. et sp. nov	Announced	Wang et al.	Permian (Cisuralian)	Shanxi Formation	China	A voltzialean conifer. The type species is H. yongchangensis. Announced in 2022 Officially published in 2023

Other conifers

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Aciphyllum^[59]	Gen. et sp. nov		Barbacka & Górecki in Barbacka et al.	Early Jurassic (Hettangian)	Zagaje Formation	Poland	A needle leaf similar to the leaves of Pinus. Genus includes new species A. triangulatum.
Ductoagathoxylon tsaaganensis^[60]	Sp. nov	In press	Cai, Zhang & Feng in Cai et al.	Late Permian		Mongolia
Sidashia^[61]	Gen. et sp. nov	In press	Forte, Kustatscher & Van Konijnenburg-van Cittert in Forte et al.	Middle Triassic (Anisian)		Italy	Genus includes new species S. tridentata.
Ullrichia^[62]	Gen. et comb. nov	Valid	Kerp et al.	Permian		Germany	The type species is "Lebachia" laxifolia (1939); genus also includes "L." intermedia (1939) and "L." mucronata (1939).

Conifer research

Bodnar et al. (2022) reassess the anatomy and systematics of the permineralized conifer-like woods from the Triassic strata from Argentina , confirm the assignment of the logs related to the families Cupressaceae and Cheirolepidiaceae, as well as three taxa related to Araucariaceae (Agathoxylon cozzoi, Agathoxylon protoaraucana and Agathoxylon argentinum), and argue that the fossil woods previously assigned to the families Podocarpaceae and Taxaceae do not have enough preserved characters to support such assignment.^[63]
A study on the pattern of conifer turnover across the Cretaceous-Paleogene boundary in the Raton and Denver basins (Colorado, United States ) is published by Berry (2022).^[64]
Mantzouka, Akkemik & Güngör (2022) describe fossil woods of Cupressinoxylon matromnense from the middle Miocene Eşelek volcanic deposits (Gökçeada, Turkey), preserved with feeding damage produced by members of the agromyzid genus Protophytobia, and supporting the existence of an eastern Mediterranean Miocene Climatic Optimum hotspot which additionally included Greek islands of Lemnos and Lesbos.^[65]

Flowering plants

Chloranthales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Canrightia foveolata^[66]	Sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation	Portugal
Proencistemon^[66]	Gen. et sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation	Portugal	Genus includes new species P. portugallicus.

Magnoliids

Laurales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Araliaephyllum silvapinedae^[67]	Sp. nov	In press	Rubalcava-Knoth & Cevallos-Ferriz	Cretaceous (Albian–Cenomanian)	La Cintura Formation	Mexico
Argapaloxylon salvadorensis^[68]	Sp. nov	Valid	Vasquez-Loranca & Cevallos-Ferriz	Miocene		El Salvador	A member of the family Lauraceae.
Catula^[69]^[70]	Gen. et sp. nov	Valid	Maccracken et al.	Late Cretaceous (Campanian)	Kaiparowits Formation	United States ( Utah)	A member of the family Lauraceae. Genus includes new species C. gettyi.
Cryptocaryoxylon irregularis^[71]	Sp. nov	Valid	Akkemik, Iamandei & Çelik	Early Miocene	Hançili Formation	Turkey	Fossil wood of a member of the family Lauraceae.
Laurinoxylon scalariforme^[68]	Sp. nov	Valid	Vasquez-Loranca & Cevallos-Ferriz	Miocene		El Salvador	A member of the family Lauraceae.
Mezilaurinoxylon americana^[68]	Sp. nov	Valid	Vasquez-Loranca & Cevallos-Ferriz	Miocene		El Salvador	A member of the family Lauraceae.
Mezilaurinoxylon draconis^[68]	Sp. nov	Valid	Vasquez-Loranca & Cevallos-Ferriz	Miocene		El Salvador	A member of the family Lauraceae.
Mezilaurinoxylon miocenica^[68]	Sp. nov	Valid	Vasquez-Loranca & Cevallos-Ferriz	Miocene		El Salvador	A member of the family Lauraceae.

Magnoliales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Magnolia allasoniae^[72]	Sp. nov	Valid	Martinetto in Niccolini et al.	Miocene (Messinian)	Piedmont Basin	Italy	A species of Magnolia. First named in 1995 but did not meet ICBN requirements; subsequently validated in 2022.

Piperales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Aristospermum^[73]	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Early Cretaceous (Aptian–Albian)		Portugal United States ( Virginia)	A member of the family Aristolochiaceae. Genus includes new species A. huberi.
Siratospermum^[73]	Gen. et sp. nov	Valid	Friis, Crane & Pedersen	Late Cretaceous (Cenomanian)		United States ( Maryland)	A member of the family Aristolochiaceae. Genus includes new species S. mauldinense.

Monocots

Lilioid monocots

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Pandanus estellae^[74]	Sp. nov	Valid	Rozefelds et al.	Oligocene		Australia	A species of Pandanus.

Commelinid monocots

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Sabalites ghughuaensis^[75]	Sp. nov	In press	Kumar, Hazra & Khan in Kumar et al.	Late Cretaceous-Paleocene (Maastrichtian-Danian)	Deccan Intertrappean Beds	India	A member of the family Arecaceae belonging to the subfamily Coryphoideae.
Sabalites umariaensis^[75]	Sp. nov	In press	Kumar, Hazra & Khan in Kumar et al.	Late Cretaceous-Paleocene (Maastrichtian-Danian)	Deccan Intertrappean Beds	India	A member of the family Arecaceae belonging to the subfamily Coryphoideae.

Monocot research

Leaf fossils of costapalmate-palms belonging to the genus Sabalites are described from the ?Santonian–Campanian Belly River Group, Campanian Foremost Formation (Alberta, Canada ) and Maastrichtian Frenchman Formation (Saskatchewan, Canada) by Greenwood, Conran & West (2022), who interpret the studied fossils as constraining climate reconstructions for the Late Cretaceous high mid-latitudes of North America (c. 55° N) to exclude significant freezing episodes; the authors also transfer the Late Cretaceous species "Geonomites" imperialis to the genus Phoenicites, and reassess Sabalites carolinensis as more likely to be Campanian than Coniacian–Santonian in age.^[76]
A study on the impact of the absence of megaherbivores in the aftermath of the Cretaceous–Paleogene extinction event on the evolution of palms is published by Onstein, Kissling & Linder (2022).^[77]
A study on the evolutionary history of palms belonging to the group Mauritiinae, as inferred from a phylogenetic analysis incorporating fossil data, is published by Bacon et al. (2022).^[78]

Basal eudicots

Proteales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Distefananthus^[79]	Gen. et sp. nov		Huegele & Wang	Early Cretaceous (Albian)	Dakota Formation	United States ( Kansas)	A platanaceous inflorescence. Genus includes new species D. hoisingtonensis.
Langeranthus^[80]	Gen et sp nov	Valid	Huegele & Manchester	Eocene Ypresian	Klondike Mountain Formation	USA Washington	A platanaceous flowering head. The type species is L. dillhoffiorum
Meliosma eosinica^[81]	Sp. nov		Moiseeva, Kodrul & Jin in Moiseeva et al.	Late Eocene	Huangniuling Formation	China	A species of Meliosma.
Nelumbo delinghaensis^[82]	Sp. nov		Luo & Jia in Luo et al.	Miocene	Upper Youshashan Formation	China	A species of Nelumbo.
Nelumbo fujianensis^[83]	Sp. nov	In press	Dong et al.	Miocene	Fotan Group	China	A species of Nelumbo.
Platimeliphyllum durhamensis^[84]	Comb nov	in press	(Wolfe)	Late Eocene	Puget Group "Upper Fultonian" Loc 9832	USA Washington	A platanaceous leaf. Moved from "Fothergilla" durhamensis (1968).
Platimeliphyllum fushunensis^[84]	Comb nov	in press	(Chen)	Eocene	Fushun Formation	China	A platanaceous leaf. Moved from "Betula" fushunensis.
Sapindopsis chinensis^[85]	Sp. nov		Golovneva et al.	Early Cretaceous (Albian)	Dalazi Formation	China	A member of the family Platanaceae.
Sapindopsis orientalis^[85]	Sp. nov		Golovneva et al.	Early Cretaceous (Albian)	Frentsevka Formation	Russia (Template:Country data Primorsky Krai)	A member of the family Platanaceae.

Protealean research

Redescription of the Okanagan Highlands genus Langeria with description of associated stipules and reproductive structures plus formal reassignment of the genus to Platanaceae by Huegele & Manchester is published.^[80]

Ranunculales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Berberis auriolensis^[86]	Sp. nov	Valid	Denk & Sami in Denk et al.	Pleistocene (Calabrian)		Italy	A species of Berberis.
Mahonia mangbangensis^[87]	Sp. nov		Tang et al.	Pliocene	Mangbang Formation	China	A species of Mahonia.
Palaeosinomenium hengduanensis^[88]	Sp. nov		Wu & Zhou in Wu et al.	Eocene	Shuanghe Formation	China	A member of the family Menispermaceae.

Superasterids

Aquifoliales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Ilex antiquorum^[89]	Nom. nov	Valid	Doweld	Late Cretaceous (Maastrichtian)		Germany	A holly; a replacement name for Ilex antiqua Knobloch & Mai (1986).
Ilex myricina^[89]	Nom. nov	Valid	Doweld	Miocene (Messinian)		Italy	A holly; a replacement name for Ilex myricoides Massalongo (1858).

Caryophyllales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Podopterus mijangosae^[90]	Sp. nov	In press	Estrada-Ruiz	Miocene	Mexican amber	Mexico	A species of Podopterus.

Cornales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Blackwelloxylon^[91]	Nom. nov	Valid	Deshmukh	Pleistocene		United States ( Mississippi)	A member of the family Cornaceae; a replacement name for Cornoxylon Blackwell (1982).
Exbeckettia^[92]	Gen. et comb. nov	Valid	Manchester & Collinson	Early Eocene	London Clay	United Kingdom	A mastixioid fruit; a new genus for Beckettia mastixioides Reid & Chandler (1933).

Dipsacales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Sambucus heqingensis^[93]	Sp. nov	In press	Huang & Zhou in Huang et al.	Late Pliocene	Heqing Basin	China	A species of Sambucus.

Ericales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Paradiospyroxylon^[94]	Gen. et sp. nov	In press	Koutecký & Sakala in Koutecký, Sakala & Chytrý	Oligocene	Ústí Formation	Czech Republic	A member of the family Ebenaceae. Genus includes new species P. kvacekii.

Icacinales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Palaeophytocrene chicoensis^[95]	Sp. nov		Atkinson	Late Cretaceous (Campanian)	Chico Formation	United States ( California)	A member of the family Icacinaceae.

Metteniusales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Calatola verae^[96]	Sp. nov		Estrada-Ruiz et al.	Miocene	Mexican amber	Mexico	A species of Calatola.

Superrosids

Cucurbitales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Libasperma^[97]	Gen. et sp. nov	In press	Huegele & Manchester	Paleocene	Fort Union Formation	United States ( Montana)	A member of the family Cucurbitaceae. Genus includes new species L. potamoglossensis.

Fabales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Albizia yenbaiensis^[98]	Sp. nov	Announced	Nguyen, Su & J. Huang in Nguyen et al.	Miocene	Yen Bai Basin	Vietnam	An Albizia species. Announced in 2022 Officially published January 2023
Anadenantheroxylon kurupaum^[99]	Sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A member of the family Fabaceae.
Cedrelinga paleocatenaeformis^[99]	Sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A species of Cedrelinga.
Cercioxylon mediterraneum^[71]	Sp. nov	Valid	Akkemik, Iamandei & Çelik	Early Miocene	Hançili Formation	Turkey	Fossil wood of a member of the family Fabaceae.
Chloroleucoxylon^[99]	Gen. et sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A member of the family Fabaceae. Genus includes new species C. yukeriense.
Enterolobiumoxylon vassalloae^[99]	Sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A member of the family Fabaceae.
Leguminocarpum meghalayensis^[100]	Sp. nov	Announced	Bhatia, Srivastava & Mehrotra	Late Paleocene	Tura Formation	India	A fabaceous seed pod morphospecies. Announced in 2022 Officially published in 2023
Microlobiusxylon parafoetidus^[99]	Sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A member of the family Fabaceae.
Paleobowdichia^[101]	Gen. et comb. nov	Valid	Herendeen et al.	Latest Paleocene to late early Eocene	Lamar River Formation	United States ( Colorado Wyoming)	A member of Papilionoideae; a new genus for "Acacia" lamarensis Knowlton (1899).
Parapiptadenioxylon^[99]	Gen. et sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A Fabaceae genus. The type species is P. pararigida.
Parvileguminophyllum damalgiriensis^[100]	Sp. nov	Announced	Bhatia, Srivastava & Mehrotra	Late Paleocene	Tura Formation	India	A fabaceous leaf morphospecies. Announced in 2022 Officially published in 2023
Podocarpium tibeticum^[102]	Sp. nov	In press	Li, Huang & Su in Li et al.	Late Eocene	Lunpola Basin	China	A member of the family Fabaceae.
Pseudopiptadenioxylon^[99]	Gen. et sp. nov	Valid	Ramos et al.	Late Pleistocene	El Palmar Formation	Argentina	A member of the family Fabaceae. Genus includes new species P. uniseriatum.
Tobya^[101]	Gen. et comb. nov	Valid	Herendeen et al.	Eocene	Cockfield Formation	United States ( Kentucky Tennessee)	A member of Papilionoideae; a new genus for "Diplotropis" claibornensis Herendeen & Dilcher (1990).

Fabalean research

New fossil material of members of the genus Bauhinia is described from the Eocene of the Puyang Basin (China) by Jia et al. (2022), who interpret their findings as the earliest reliable fossil records of Bauhinia in Asia.^[103]
Moya et al. (2022) study the affinities of fossil legumes Entrerrioxylon victoriensis, Gossweilerodendroxylon palmariensis, Paraoxystigma concordiensis and Cylicodiscuxylon paragabunensis from the Cenozoic Paraná, Arroyo Feliciano and El Palmar formations (Argentina) with extant West African legumes, and discuss the possible migration routes by which these plants may have arrived in South America from Africa.^[104]

Fagales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Castanopsis zhoui^[105]	Sp. nov	In press	Wang et al.	Miocene	Fotan Group	China	A species of Castanopsis.
Comptonia hirsuta^[106]	Sp. nov		Xiao & Ji in Ji et al.	Miocene	Hannuoba Formation	China	A species of Comptonia.
Myricoxylon doganyurtensis^[71]	Sp. nov	Valid	Akkemik, Iamandei & Çelik	Early Miocene		Turkey	Fossil wood of a member of the family Myricaceae.
Nothofagoxylon ruei^[55]	Sp. nov	Announced	Pujana et al.	Oligocene	San José Formation	Chile	A nothofagaceous wood morphospecies Announced in 2022 Officially published in 2023
Pterocarya magnifructa^[107]	Sp. nov	Valid	Stults, Tiffney & Axsmith	Pliocene		United States ( Alabama)	A species of Pterocarya.
Quercus nanningensis^[108]	Sp. nov	In press	Liu & Jin in Liu et al.	Late Oligocene	Yongning Formation	China	An oak.
Quercus paleodisciformis^[108]	Sp. nov	In press	Liu & Jin in Liu et al.	Late Oligocene	Yongning Formation	China	An oak.
Quercus paleohui^[108]	Sp. nov	In press	Liu & Jin in Liu et al.	Late Oligocene	Yongning Formation	China	An oak.
Quercus yongningensis^[108]	Sp. nov	In press	Liu & Jin in Liu et al.	Late Oligocene	Yongning Formation	China	An oak.

Malpighiales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Belenocarpa^[109]	Gen. et comb. nov	Valid	Hamersma et al.	Early Oligocene		Peru	A member of the family Euphorbiaceae; a new genus for "Jatropha" tertiara Berry.
Elatine odgaardii^[110]	Sp. nov	Valid	Bennike in Bennike et al.	Probably early Pleistocene		Greenland	A species of Elatine. Announced in 2022; the final article version was published in 2023.
Mammeoxylon beylikduezuense^[111]	Sp. nov	In press	Akkemik et al.	Late Oligocene-Early Miocene	İstanbul Formation	Turkey
Parinari hilliana^[112]	Sp. nov	Valid	Grote in Grote, Duangkrayom & Jintasakul	Late Miocene	Tha Chang beds	Thailand	A species of Parinari.
Parinari khoratensis^[112]	Sp. nov	Valid	Grote in Grote, Duangkrayom & Jintasakul	Late Miocene	Tha Chang beds	Thailand	A species of Parinari.
Plukenetia minima^[113]	Sp. nov	In press	Poinar	Miocene	Dominican amber	Dominican Republic	A species of Plukenetia.

Malvales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Malvacipolloides deccanensis^[114]	Sp. nov		Manchester et al.	Late Cretaceous-Paleocene (Maastrichtian–Danian)	Deccan Intertrappean Beds	India	A member of the family Malvaceae.
Malvacipolloides intertrappea^[114]	Sp. nov		Manchester et al.	Late Cretaceous-Paleocene (Maastrichtian–Danian)	Deccan Intertrappean Beds	India	A member of the family Malvaceae.
Thespesia neopopulnea^[115]	Sp. nov	Valid	Hazra, Mahato & Khan in Hazra et al.	Pliocene	Rajdanda Formation	India	A species of Thespesia.

Malvalean research

A study on the evolutionary history of Dipterocarpaceae, as indicated by biogeography of pollen fossils from Africa and India, molecular data and fossil amber records, is published by Bansal et al. (2022).^[116]

Myrtales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Hemitrapa zhangpuensis^[117]	Sp. nov		Dong et al.	Miocene		China	A member of the family Lythraceae belonging to the subfamily Trapoideae.
Myrtineoxylon hoffmannae^[55]	Sp. nov	Announced	Pujana et al.	Oligocene	San José Formation	Chile	A myrtaceous wood morphospecies. Announced in 2022 Officially published in 2023
Trapa natanifolia^[118]	Sp. nov		Han & Jia in Han et al.	Late Eocene	Bailuyuan Formation	China	A water caltrop.
Trapa qaidamensis^[119]	Sp. nov		Cai et al.	Miocene	Shangyoushashan Formation	China	A water caltrop.

Oxalidales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Ceratopetalum suciensis^[120]	Sp nov	In press	Tang, Smith, & Atkinson	Late Cretaceous Campanian	Cedar District Formation	United States ( Washington)	A Cunoniaceous species.
Connaroxylon^[121]	Gen. et sp. nov	Valid	Baas et al.	Cretaceous Maastrichtian-earliest Paleocene	Deccan Intertrappean Beds	India	A probable Connaraceous wood morphotaxon. The type species is C. dimorphum. First named in 2017 but failed ICBN requirements;^[122] subsequently validated in 2022.^[121]
Cunoniocarpa^[123]	Gen. et sp. nov	Valid	Matel et al.	Early Eocene	Huitrera Formation	Argentina	A member of Cunoniaceae. The type species is C. stylosa.
Racemofructus^[123]	Gen. et sp. nov	Valid	Matel et al.	Early Eocene	Huitrera Formation	Argentina	A member of Cunoniaceae. The type species is R. fasciculatus.
Weinmannioxylon trichospermoides^[55]	Sp. nov	Announced	Pujana et al.	Oligocene	San José Formation	Chile	A cunoniaceous wood morphospecies. Announced in 2022 Officially published in 2023

Oxalidalean research

Tand, Smith, and Atkinson describe the first North American instance of the previously Paleo-Antarctic Rainforest Lineage Cunoniaceae fruits from Sucia Island. Previously considered solely a Gondwanan family, the new species indicate a complex geographic history for the group.^[120]

Rosales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Eophylica^[124]	Gen. et sp. nov	Valid	Shi et al.	Cretaceous	Burmese amber	Myanmar	A rhamnaceous floral morphotaxon. The type species is E. priscastellata.
Ficoxylon fusiforme^[125]	Sp. nov	Valid	El-Noamani	Late Cretaceous	Taref Formation	Egypt	A member of the family Moraceae.
Ficus fujianensis^[126]	Sp. nov	In press	Dong et al.	Miocene		China	A species of Ficus.
Ficus zhangpuensis^[126]	Sp. nov	In press	Dong et al.	Miocene		China	A species of Ficus.
Phylica piloburmensis^[124]	Sp. nov	Valid	Shi et al.	Cretaceous	Burmese amber	Myanmar	A species of Phylica.
Ventilago pliocenica^[127]	Sp. nov		Hazra et al.	Pliocene		India	A species of Ventilago.
Ventilago siwalika^[127]	Sp. nov		Hazra et al.	Miocene		India	A species of Ventilago.

Sapindales

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Barkleya^[128]	Gen. et comb. nov	Valid	Manchester & Judd	Eocene Ypresian	Green River Formation	United States Colorado	An anacardiaceous samara. Type species Anacardites schinoloxus (1929). Possibly the fruits of Rhus nigricans.
Canarium haominiae^[129]	Sp. nov	In press	Yin et al.	Miocene		China	A species of Canarium.
Canarium maomingense^[130]	Sp. nov		Xiang & Jin in Xiang et al.	Late Pleistocene	Maoming Basin	China	A species of Canarium.
Choerospondias mioaxillaris^[131]	Sp. nov		Xiao & Wu in Xiao et al.	Miocene	Shengxian Formation	China	A species of Choerospondias.
Choerospondias tiantaiensis^[131]	Sp. nov		Xiao & Wu in Xiao et al.	Miocene	Shengxian Formation	China	A species of Choerospondias.
Grimmipollis^[132]	Gen. et sp. nov		Huang, Morley & Hoorn in Huang et al.	Eocene	Yaw Formation	Myanmar	A member of the family Sapindaceae. Genus includes new species G. burmanica.
Koelreuteria kvacekii^[133]	Sp. nov	Valid	Chen, Del Rio & Su in Chen et al.	Eocene	Niubao Formation	China	A species of Koelreuteria.
Loxopteroides^[134]	Gen. et sp. nov	In press	Manchester & Judd	Eocene	Ione Formation	United States ( California Oregon)	A member of the family Anacardiaceae. Genus includes new species L. weeksae.
Vaudoisia^[135]	Gen. et comb. nov	Valid	Strullu-Derrien et al.	Eocene		France	A fruit of likely sapindalean affinity; a new genus for "Juglandicarya" gruetii Vaudois-Miéja (1976).

Other Eudicots

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Zlatkovia^[136]	Gen. et sp. nov	In press	Rothwell & Stockey	Late Cretaceous	St. Mary River Formation	Canada ( Alberta)	An aquatic eudicot. The type species is Z. crenulata.

Other angiosperms

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Archaebuda^[137]	Gen. et sp. nov		Chen & Wang	Early Cretaceous (Barremian-Aptian)	Yixian Formation	China	A flower bud of an early angiosperm. Genus includes new species A. lingyuanensis.
Ascarinophyllum^[138]	Gen. et sp. nov	Announced 2022	Čepičková & Kvaček	Late Cretaceous (Cenomanian)	Peruc–Korycany Formation	Czech Republic	A Basal angiosperm leaf morphogenus Similar to Mesodescolea plicata and Chloranthaceae. The type species is A. pecinovense. Officially published in 2023
Covidifructus^[139]	Gen. et sp. nov	Valid	Heřmanová et al.	Late Cretaceous (late Turonian-Santonian)	Klikov Formation	Czech Republic	An angiosperm fruit of uncertain affinities, with similarities to the family Dilleniaceae. The type species is C. multicarpellatus.
Elasmostemon^[66]	Gen. et sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation	Portugal	A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species E. paisii.
Endressistemon^[66]	Gen. et sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation	Portugal	A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species E. cateficensis.
Fairlingtonia microgyna^[140]	Sp. nov		Du et al.	Early Cretaceous	Zhonggou Formation	China	A herbaceous eudicot.
Florigerminis^[141]	Gen. et sp. nov	In press	Cui et al.	Middle-Late Jurassic	Jiulongshan Formation	China	A possible flower bud. The type species is F. jurassica. First announced online 2021, Final article published in 2022.
Gansupeltata^[142]	Gen. et sp. nov	Valid	Wu et al.	Early Cretaceous (Aptian)	Chijinpu Formation	China	An early flowering plant. Genus includes new species G. beishanensis.
Herbifolia^[143]	Gen. et sp. nov	In press	Frolov & Enushchenko	Middle Jurassic (Aalenian)	Irkutsk Coal Basin	Russia	An angiosperm with leaf epidermal structure most similar to those of modern Asparagales and Liliales. Genus includes new species H. antiqua.
Honeytheca^[97]	Gen. et sp. nov	In press	Huegele & Manchester	Paleocene	Fort Union Formation	United States ( Montana)	A flowering plant of uncertain affinities. Genus includes new species H. bighornensis.
Ibericarpus^[66]	Gen. et sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation	Portugal	A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species I. cuneiformis.
Lingyuananthus^[144]	Gen. et sp. nov		Wang	Early Cretaceous (Barremian–Aptian)	Yixian Formation	China	An early angiosperm. Genus includes new species L. inexpectus.
Santaniella^[145]	Gen. et 2 sp. nov		Gobo et al.	Early Cretaceous (Barremian-Aptian)	Crato Formation	Brazil	Originally described as a member or a relative of the family Ranunculaceae, but subsequently considered to be a mesangiosperm of uncertain affinities, possibly a magnoliid.^[146] Genus includes new species S. lobata and S. acuta.
Todziaphyllum saportanum^[138]	Comb. nov	Announced	(Velenovský) Čepičková & Kvaček	Late Cretaceous (Cenomanian)	Peruc–Korycany Formation	Czech Republic	A Basal angiosperm leaf morphogenus A new combination for Banksites saportanus Officially published in 2023
Tolmania^[147]	Gen. et sp. nov	Valid	Edmonds, Stockey & Rothwell	Late Cretaceous (Maastrichtian)	St. Mary River Formation	Canada ( Alberta)	An aquatic dicot. Genus includes new species T. aquatica.
Valvidistemon^[66]	Gen. et sp. nov	Valid	Friis et al.	Early Cretaceous (Aptian-Albian)	Almargem Formation	Portugal	A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species V. globiferus.

General angiosperm research

Surangea mohgaoensis, originally interpreted as fern megaspores, is reinterpreted as angiosperm fruits by Ramteke et al. (2022).^[148]
Zhang et al. (2022) describe rich assemblages of spiny plant fossils from the Eocene (Bartonian) Niubao Formation (Tibet, China), preserving seven different spine morphologies, and interpret this finding as evidence of the presence of a diversity of spiny plants in Eocene central Tibet, as well as evidence of a rapid diversification of spiny plants in Eurasia around that time.^[149]
A preliminary report on a new fossil angiosperm flora of the Lesvos Petrified Forest at Akrocheiras east of Sigri on Lesbos, Greece is given by Kafetzidou et al. Preliminary taxa identifications are given and commentary on the climactic implications are made.^[150]
A study aiming to determine the relationship between past atmospheric CO₂ and temperature fluctuations and the shifts in diversification rates of Poaceae and Asteraceae is published by Palazzesi et al. (2022).^[151]

Other plants

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Aegianthus irkutensis^[152]	Sp. nov	In press	Nosova & Tekleva	Middle Jurassic	Prisayan Formation	Russia	Pollen cone with pollen of ginkgoalean or gnetophytalean affinity.
Aysenoxylon^[55]	Gen et sp nov	Announced	Pujana et al.	Oligocene	San José Formation	Chile	A wood morphospecies of uncertain affinity. The type species is A. patorarensis. Announced in 2022 Officially published in 2023
Bryokhutuliinia ignatovii^[153]	Sp. nov	Valid	Frolov, Kazanovsky & Enushchenko	Early Jurassic (Toarcian)	Middle Subformation of Prisayan Formation	Russia (Template:Country data Irkutsk Oblast)	A member of Bryopsida of uncertain affinities.
Combina^[154]	Gen. et sp. nov		Santos & Wang	Middle Triassic (Anisian)	Calcena Formation	Spain	A cone-like reproductive organ of a seed plant. Genus includes new species C. triassica.
Dioonitocarpidium rossicum^[155]	Sp. nov	Valid	Gomankov	Permian		Russia	A member of Cycadales.
Europoxylon garapensis^[156]	Sp. nov	In press	Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A gymnosperm.
Jarudia^[157]	Gen. et sp. nov		Shi et al.	Early Cretaceous	Huolinhe Formation	China	A seed-bearing structure of a corystosperm seed fern. Genus includes new species J. zhoui.
Komlopteris distinctiva^[59]	Sp. nov		Barbacka in Barbacka et al.	Early Jurassic (Hettangian)	Zagaje Formation	Poland	Cuticle of a seed fern.
Lesleya ceriacoi^[158]	Sp. nov	In press	Correia et al.	Carboniferous (Gzhelian)	Douro Carboniferous Basin	Portugal	An early gymnosperm.
Palaeodichelyma kiritchkovae^[153]	Sp. nov	Valid	Frolov, Kazanovsky & Enushchenko	Early Jurassic (Pliensbachian)	Lower Subformation of Prisayan Formation	Russia (Template:Country data Irkutsk Oblast)	A member of Bryopsida of uncertain affinities.
Paragigantopteris^[159]	Gen. et sp. nov	In press	Ma et al.	Permian (Wuchiapingian)	Lungtan Formation	China	A gigantopterid. Genus includes new species P. qingloongensis.
Pauthecophyton hezhangensis^[160]	Sp. nov		Wang et al.	Devonian (Pragian-Emsian)	Danlin Formation	China	A euphyllophyte of uncertain affinities.
Polycanaloxylon^[156]	Gen. et sp. nov	In press	Conceição et al.	Permian (Cisuralian)	Pedra de Fogo Formation	Brazil	A gymnosperm. Genus includes new species P. merlottii.
Psilophyton diakanthon^[161]	Sp. nov		Colston, Landaw & Tomescu	Devonian (Emsian)	Battery Point Formation	Canada ( Quebec)	A member of the group Trimerophytopsida.
Renbernia^[162]	Gen. et sp. nov	In press	Friis, Crane & Pedersen	Early Cretaceous (Albian)	Potomac Group	United States ( Virginia)	A seed plant similar to Brenneria potomacensis. Genus includes new species R. zhoui.
Rhyniotaenium^[163]	Gen. et sp. nov	In press	Krings	Early Devonian	Rhynie chert	United Kingdom	An alga, probably a green alga belonging to the family Mesotaeniaceae. Genus includes new species R. velatum.
Sinoglossa^[164]	Gen. et sp. nov	Valid	Zhang et al.	Middle Triassic	Linjia Formation	China	A member of Glossopteridales. The type species is S. sunii.
Taimyria^[165]	Gen. et sp. nov	Valid	Naugolnykh & Mogutcheva	Early Triassic (Induan)	Keshinskian/Keshinskaya Formation	Russia (Template:Country data Krasnoyarsk Krai)	A member of Peltaspermales belonging to the family Angaropeltaceae. Genus includes new species T. triassica.
Taungurungia^[166]	Gen. et sp. nov	Valid	McSweeney, Shimeta & Buckeridge	Devonian (Pragian–Emsian)	Norton Gully Sandstone Formation	Australia	A plant of uncertain affinities, similar to members of Zosterophyllopsida. Genus includes new species T. garrattii.
Teyoua^[167]	Gen. et sp. nov	In press	Huang, Liu & Xue	Devonian (probably Pragian)	Mangshan Group	China	A polysporangiate land plant. Genus includes new species T. antrorsa.
Traskia^[168]	Gen. et sp. nov	Valid	Rothwell et al.	Jurassic		Canada ( British Columbia)	A stem-cycad. Genus includes new species T. maahlae.
Vetiplanaxis obtusus^[169]	Sp. nov	In press	Li et al.	Cretaceous	Burmese amber	Myanmar	A moss belonging to the group Hypnodendrales.
Wilhowia^[170]	Gen. et sp. nov	Valid	Gensel	Devonian (Emsian)	Battery Point Formation	Canada ( Quebec)	A basal euphyllophyte. Genus includes new species W. phocarum.
Xadzigacalix^[171]	Gen. et sp. nov	Valid	Klymiuk, Rothwell & Stockey	Early Cretaceous (Valanginian)		Canada ( British Columbia)	A gymnosperm of uncertain phylogenetic placement, possibly having affinities with gnetophytes or angiosperms. Genus includes new species X. quatsinoensis.
Xinhangia^[172]	Gen. et sp. nov		Yang & Wang	Devonian (Famennian)	Wutong Formation	China	A fern-like plant of uncertain affinities. Genus includes new species X. spina.

Other plant research

A study on the xylem development in Leptocentroxyla, and on its implications for the knowledge of the evolution of pith, is published by Tomescu & McQueen (2022).^[173]
Decombeix et al. (2022) report evidence of tylosis formation in permineralized wood of Dameria hueberi from the Tournaisian of Australia .^[174]
The first comprehensive crown reconstruction of Medullosa stellata var. typica, based on data from a specimen from the Chemnitz petrified forest (Germany ), is presented by Luthardt et al. (2022).^[175]
Fossil material of Rhabdotaenia is reported from the Permian Umm Irna Formation (Jordan) by Blomenkemper et al. (2022), representing the northernmost occurrence of this Gondwanan leaf type reported to date.^[176]

Palynology

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Anapiculatisporites radiatus^[177]	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation	Australia	A trilete spore.
Apricasporites^[177]	Gen. et sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation	Australia	A trilete spore. Genus includes new species A. cancellosus.
Camptotriletes inaequabilis^[177]	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation	Australia	A trilete spore.
Camptotriletes suggrandis^[177]	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation	Australia	A trilete spore.
Convolutispora inreligata^[177]	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation	Australia	A trilete spore.
Endosporites circumsaeptus^[177]	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation	Australia	A trilete spore.
Foveosporites magnus^[177]	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation	Australia	A trilete spore.
Granulatisporites commutabilis^[177]	Sp. nov	Valid	Playford	Carboniferous (Mississippian)	Lyall Formation	Australia	A trilete spore.
Paxillitriletes permicus^[178]	Sp. nov	In press	Sui, McLoughlin & Feng in Sui et al.	Permian (Wuchiapingian–Changhsingian)	Xuanwei Formation	China	A lycophyte megaspore.
Sergipea multipapillata^[179]	Sp. nov		Hu et al.	Early Cretaceous	Bongor Basin	Chad	A gymnosperm pollen.
Volkheimerites^[180]	Gen. et sp. nov	In press	Narváez et al.	Paleocene (Danian)	Salamanca Formation	Argentina	Pollen of a flowering plant. The type species is V. labyrinthus.
Yezopollis^[181]	Gen. et sp. nov	In press	Legrand, Yamada & Nishida	Late Cretaceous (Cenomanian–Turonian)	Mikasa Formation	Japan	A Normapolles-type flowering plant pollen. Genus includes new species Y. mikasaensis.

Research

Review of the studies on the origin of the land flora is published by Bowman (2022).^[182]
A study on the evolution of body plans of members of Viridiplantae, based on a review of the fossil record, molecular data and developmental biology, is published by Niklas & Tiffney (2022).^[183]
A study on the biodiversity of land plants at the equator during their first major diversification in the Late Silurian–Early Devonian is published by Wellman et al. (2022).^[184]
A study on the evolution of heterospory during the Devonian is published by Leslie & Bonacorsi (2022).^[185]
Seven coniferous nurse logs that have been colonized by conifer and equisetalean roots are reported from four Permian intervals in the Ordos Basin (China ) by Feng et al. (2022), indicating that conifer tree stems probably functioned as hosts to both conspecific and interspecific seedlings in the Cathaysian Flora.^[186]
A study on the impact of the Intertropical Convergence Zone in the emerging South Atlantic region on Aptian plant communities from eight Brazilian sedimentary basins is published by Carvalho et al. (2022), who report evidence of an overall predominance of xerophytic plants, attesting to more dry conditions, and of a humidification trend towards the end of the late Aptian resulting in the predominance of hydrophytes, hygrophytes, tropical lowland flora and upland flora, indicative of prevalence of lowland and montane rainforests.^[187]
A study on the distribution and relative abundances of major plant groups from the Albian Gates Formation (Alberta, Canada ) is published by Kalyniuk et al. (2022).^[188]
A study on the relationship between whole-genome duplication, seed traits and the selectivity of the survival of plants during the Cretaceous–Paleogene extinction event is published by Berry & Jaganathan (2022).^[189]
New Oligocene flora is described from the Dong Ho Formation (Vietnam) by Huang et al. (2022), who interpret the studied fossils as evidence of long-term environmental, floristic and vegetational stability in this region since the Paleogene.^[190]
Gentis et al. (2022) describe fossil wood specimens from the Miocene Natma Formation (Myanmar), representing an assemblage dominated by members of the families Fabaceae and Dipterocarpaceae, interpreted as coming from different types of low altitude forest ecosystems (tropical wet evergreen, tropical dry and deciduous, and tropical littoral), and interpreted as indicative of a monsoonal climate with an alternance of a dry season and a wet season.^[191]
Abundant compression floras dominated by angiosperm leaves are described from two sites of probable Pliocene age in Brunei by Wilf et al. (2022), who interpret these floras as evidence of dipterocarp-dominated lowland rainforests in the Malay Archipelago before the Pleistocene.^[192]
A study on the impact of the extinct Neotropical megafauna on the variability in plant functional traits and biome geography in Central and South America is published by Dantas & Pausas (2022).^[193]
A study on plant material from rock overhangs from mid-late Holocene sites along the Kawarau-Cromwell-Roxburgh Gorges in Central Otago (New Zealand), much of which was likely transported as roosting material or consumed by moa birds, and on its implications for the knowledge of the mid-late Holocene regional vegetation of Central Otago and the knowledge of vegetation changes since mid-late Holocene, is published by Pole (2022).^[194]
A study on the role of hydraulic failure in the evolution of early vascular plants is published by Bouda et al. (2022), suggesting that drought selection played a key role in the diversification of vascular arrangements beginning with the Devonian explosion.^[195]

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↑ ^38.0 ^38.1 Saadatnejad, J. (2022). "Two new species of Dictyozamites (Bennettitales) from the Rhaetic Kalariz Formation, North of Iran". Revista Brasileira de Paleontologia 25 (2): 135–143. doi:10.4072/rbp.2022.2.03.
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↑ Kvaček, J. (2022). "New species of Zamites from the Cenomanian of the Bohemian Cretaceous Basin". Fossil Imprint 78 (2): 425–431. doi:10.37520/fi.2022.017. http://fi.nm.cz/en/clanek/new-species-of-zamites-from-the-cenomanian-of-the-bohemian-cretaceous-basin-2/.
↑ ^41.0 ^41.1 Sun, Y.; Deng, S.; Lu, Y.; Fan, R.; Ma, X.; Lü, D. (2022). "Emendation of the Triassic plant species Glossophyllum shensiense (Ginkgoales) with a review of the genus Glossophyllum Kräusel". Review of Palaeobotany and Palynology 301: 104657. doi:10.1016/j.revpalbo.2022.104657. Bibcode: 2022RPaPa.30104657S.
↑ Nosova, N.; Kostina, E. (2022). "New findings of the female reproductive structures of Umaltolepis Krassilov and associated leaves of Pseudotorellia Florin in the Lower Cretaceous of Mongolia". Review of Palaeobotany and Palynology 304: Article 104696. doi:10.1016/j.revpalbo.2022.104696. Bibcode: 2022RPaPa.30404696N.
↑ ^43.0 ^43.1 Dong, C.; Shi, G.; Zhang, X.; Wang, Z.; Wang, Y. (2022). "Middle-Late Jurassic fossils from Northeast China confirm the affiliation of Umaltolepis seed-bearing structure and Pseudotorellia leaves". Review of Palaeobotany and Palynology 306: Article 104763. doi:10.1016/j.revpalbo.2022.104763. Bibcode: 2022RPaPa.30604763D.
↑ Frolov, A. O.; Mashchuk, I. M. (2022). "New Discoveries and New Combinations of the Fossil-genus Ginkgoites Seward (Ginkgoales) from the Lower and Middle Jurassic of East Siberia (Russia)". Phytotaxa 567 (1): 49–60. doi:10.11646/phytotaxa.567.1.4.
↑ Bodnar, J.; Sagasti, A. J.; Correa, G. A.; Miranda, V.; Medina, F. (2022). "Araucariaceous fossil woods from the Upper Triassic Ischigualasto Formation (San Juan Province, Argentina): paleofloristic and paleoclimatic implications". Journal of Paleontology 96 (6): 1354–1378. doi:10.1017/jpa.2022.45. Bibcode: 2022JPal...96.1354B.
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↑ Shi, X.; Sun, Y.; Meng, F.; Yu, J.; Lan, Z. (2022). "Early Cretaceous Keteleerioxylon Wood in the Songliao Basin, Northeast China, and Its Geographic and Environmental Implications". Biology 11 (11): 1624. doi:10.3390/biology11111624. PMID 36358325.
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↑ Li, X.-C.; Hu, Y.; Zhang, X.; Xiao, L.; Liang, L.-N.; Zhang, R.-Z.; Qiao, L. (2022). "A novel seed cone of Pinus from the Miocene of coastal Southeast China indicates kinship with Southeast Asian pines". Plant Diversity. doi:10.1016/j.pld.2022.12.002.
↑ ^55.0 ^55.1 ^55.2 ^55.3 ^55.4 ^55.5 Pujana, R. R.; Bostelmann, J. E.; Ugalde, R. A.; Riquelme, M. P.; Torres, T. (2022). "Fossil woods from the Pato Raro Heights, Patagonia National Park, Aysén, Chile: A new paleobotanical assemblage at the Oligocene climate transition". Review of Palaeobotany and Palynology 309: 104814. doi:10.1016/j.revpalbo.2022.104814.
↑ Castañeda, C. (2022). "Podocarpus (Podocarpaceae) wood from Miocene rocks in Panotla, Tlaxcala, Mexico". Journal of South American Earth Sciences 121: 104118. doi:10.1016/j.jsames.2022.104118.
↑ Jiang, Z.; Tian, N.; Wang, Y.; Li, Y.; Zheng, S.; Xie, A.; Zhu, Y. (2022). "A new structurally preserved fossil umbrella pine from the Jurassic of East Asia". Geological Journal 57 (9): 3521–3537. doi:10.1002/gj.4467.
↑ Wang, X.; Yang, Y.; Hua, Y.; Sun, B.; Miao, Y. (2022). "Hexicladia, a new genus of the Cisuralian conifer from Hexi Corridor, China". Review of Palaeobotany and Palynology 308: 104789. doi:10.1016/j.revpalbo.2022.104789.
↑ ^59.0 ^59.1 Barbacka, M.; Górecki, A.; Pacyna, G.; Pieńkowski, G.; Philippe, M.; Bóka, K.; Ziaja, J.; Jarzynka, A. et al. (2022). "Early Jurassic coprolites: insights into palaeobotany and the feeding behaviour of dinosaurs". Papers in Palaeontology 8 (2): e1425. doi:10.1002/spp2.1425.
↑ Cai, Y.; Zhang, H.; Feng, Z.; Gou, X.; Byambajav, U.; Zhang, Y.; Yuan, D.; Qie, W. et al. (2022). "A new conifer stem, Ductoagathoxylon tsaaganensis, from the Upper Permian of the South Gobi Basin, Mongolia and its palaeoclimatic and palaeoecological implications". Review of Palaeobotany and Palynology 304: Article 104719. doi:10.1016/j.revpalbo.2022.104719. Bibcode: 2022RPaPa.30404719C.
↑ Forte, G.; Kustatscher, E.; Nowak, H.; Van Konijnenburg-van Cittert, J. H. A. (2022). "Conifer Cone and Dwarf Shoot Diversity in the Anisian (Middle Triassic) of Kühwiesenkopf/Monte Prà della Vacca (Dolomites, Northeastern Italy)". International Journal of Plant Sciences 183 (9): 729–767. doi:10.1086/722036.
↑ Kerp, H.; Bödige, H.; Bomfleur, B.; Schneider, J. W. (2022). "First records of the conifers Majonica and Ortiseia from the German Zechstein (upper Permian) of east Thuringia and west Saxony, Germany". Botany Letters 169 (4): 423–441. doi:10.1080/23818107.2022.2122555.
↑ Bodnar, J.; Cuesta, V.; Escapa, I. H.; Nunes, G. C. (2022). "Exploring the first appearance of the main derived conifer families of Gondwana: evidence provided by the Triassic woods from Argentina". Ameghiniana 60 (1): 18–47. doi:10.5710/AMGH.16.11.2022.3520.
↑ Berry, K. (2022). "Conifer turnover across the K/Pg boundary in Colorado, U.S.A., parallels South American patterns: New and emerging perspectives". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 303 (1): 11–28. doi:10.1127/njgpa/2022/1035.
↑ Mantzouka, D.; Akkemik, Ü.; Güngör, Y. (2022). "Miocene Cupressinoxylon from Gökçeada (Imbros), Turkey with Protophytobia cambium mining and the study of ecological signals of wood anatomy". PeerJ 10: e14212. doi:10.7717/peerj.14212. PMID 36530400.
↑ ^66.0 ^66.1 ^66.2 ^66.3 ^66.4 ^66.5 Friis, E. M.; Crane, P. R.; Pedersen, K. R.; Mendes, M. M.; Kvaček, J. (2022). "The Early Cretaceous mesofossil flora of Catefica, Portugal: angiosperms". Fossil Imprint 78 (2): 341–424. doi:10.37520/fi.2022.016. http://fi.nm.cz/en/clanek/the-early-cretaceous-mesofossil-flora-of-catefica-portugal-angiosperms-2/.
↑ Rubalcava-Knoth, M. A.; Cevallos-Ferriz, S. R. S. (2022). "Lauraceous palmately lobed leaf from the middle Cretaceous Cintura Formation (Albian–Cenomanian), Sonora, Mexico: identification based on two comparative models". Cretaceous Research 140: 105355. doi:10.1016/j.cretres.2022.105355. Bibcode: 2022CrRes.14005355R.
↑ ^68.0 ^68.1 ^68.2 ^68.3 ^68.4 Vasquez-Loranca, A. R.; Cevallos-Ferriz, S. R. S. (2022). "A diverse assemblage of Miocene Lauraceae in Chalatenango, El Salvador". IAWA Journal 43 (4): 479–507. doi:10.1163/22941932-bja10096.
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[CRcharophytes-1] 1.0 ^1.1 ^1.2 Feist, M.; Floquet, M. (2022). "Charophytes from the Upper Cretaceous Castilian marine ramp and continental basins (central northern Spain): fossil assemblages and depositional environments". Cretaceous Research 140: Article 105325. doi:10.1016/j.cretres.2022.105325. Bibcode: 2022CrRes.14005325F.

[2] Cao, W.; Li, S.; Li, Q.; Stidham, T. A.; Wan, X.; Ni, X. (2022). "Asian Paleocene charophyte records demonstrate Eocene dispersals from Asia to Europe". Journal of Paleontology 96 (3): 706–714. doi:10.1017/jpa.2021.118. Bibcode: 2022JPal...96..706C.

[Ardeiporella-3] 3.0 ^3.1 Grgasović, T. (2022). "Taxonomy of the fossil calcareous algae: Revision of genera Physoporella Steinmann and Oligoporella Pia (Dasycladales)". Carnets Geol. 22 (7): 171–310. doi:10.2110/carnets.2022.2207. http://paleopolis.rediris.es/cg/22/07/index.html.

[4] LoDuca, S. T.; Meacher, M.; Pepper, P.; Brett, K.; Isotalo, P. A. (2022). "Earltonella fredricksi n. gen n. sp. and Thalassocystis striata (Chlorophyta, Bryopsidales) from the Silurian (Llandoverian) of the Timiskaming outlier, Ontario, Canada". Journal of Paleontology 97 (2): 516–532. doi:10.1017/jpa.2022.86.

[5] Torromé, D.; Schlagintweit, F. (2022). "Milanovicella? canadillana sp. nov., an Upper Cretaceous supposedly calcitic Dasycladale (green algae) from the middle Santonian–lower Campanian of northeastern Spain". Cretaceous Research 141: Article 105365. doi:10.1016/j.cretres.2022.105365.

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[7] Vachard, D.; Krainer, K. (2022). "Calcareous algae and foraminifers across the Permian-Triassic boundary interval (uppermost Bellerophon Formation and basal Werfen Formation) in the Dolomites (South Tyrol – Trentino, Italy)". Palaeontographica Abteilung A 324 (1–6): 1–173. doi:10.1127/pala/2022/0128. Bibcode: 2022PalAA.324....1V.

[8] Deng, S.; Lu, Y.; Fan, R.; Luo, Z.; Ma, X.; Lyu, D.; Sun, Y. (2022). "Lycopsid Lepacyclotes Emmons from the Middle Triassic of the Ordos Basin, North China and reviews of the genus". Review of Palaeobotany and Palynology 308: 104660. doi:10.1016/j.revpalbo.2022.104660.

[9] Herrera, F.; Testo, W. L.; Field, A. R.; Clark, E. G.; Herendeen, P. S.; Crane, P. R.; Shi, G. (2022). "A permineralized Early Cretaceous lycopsid from China and the evolution of crown clubmosses". New Phytologist 233 (5): 2310–2322. doi:10.1111/nph.17874. PMID 34981832.

[10] Edwards, D.; Li, C.-S.; Berry, C. M. (2022). "Lower Devonian lycophytes from Sichuan and the paleogeographic context of coeval plant assemblages from South China". International Journal of Plant Sciences 183 (6): 413–431. doi:10.1086/720387.

[11] Spiekermann, R.; Jasper, A.; Pozzebon-Silva, Â.; Carniere, J. S.; Benício, J. R. W.; Guerra-Sommer, M.; Uhl, D. (2022). "Small but not trivial: Nothostigma sepeensis sp. nov., a lycopsid from the Cisuralian (early Permian) of the Paraná basin, Brazil". Journal of South American Earth Sciences 122: 104188. doi:10.1016/j.jsames.2022.104188.

[12] Liu, L.; Wang, D.-M.; Zhou, Y.; Qin, M.; Ferguson, D. K.; Meng, M.-C. (2022). "A Late Devonian tree lycopsid with large strobili and isotomous roots". Communications Biology 5 (1): 966. doi:10.1038/s42003-022-03934-4. PMID 36109665.

[13] Deng, S.; Lu, Y.; Fan, R.; Ma, X.; Lyu, D.; Luo, Z.; Sun, Y. (2022). "A new species of Pleuromeia (Lycopsid) from the upper Middle Triassic of Northern China and discussion on the spatiotemporal distribution and evolution of the genus". Geobios 75: 1–15. doi:10.1016/j.geobios.2022.10.001. Bibcode: 2022Geobi..75....1D.

[14] Prestianni, C.; Rustán, J. J.; Balseiro, D.; Vaccari, N. E. (2022). "Porongodendron minitensis gen. nov. sp. nov. a new lycopsid from the Mississippian of Argentina with adaptations to tundra-like conditions". Botany Letters 169 (4): 527–539. doi:10.1080/23818107.2022.2101515. https://orbi.uliege.be/bitstream/2268/302490/1/Porongodendron minitensis gen nov sp nov a new lycopsid from the Mississippian of Argentina with adaptations to tundra like conditions.pdf.

[Biology1111Hymenophyllaceae-15] 15.0 ^15.1 ^15.2 ^15.3 Li, Y.; Wang, Y.-D.; Nosova, N.; Lu, N.; Xu, Y.-Y. (2022). "Filmy Ferns (Hymenophyllaceae) and Associated Spike-Mosses (Selaginellaceae) from the Mid-Cretaceous Kachin Amber, Myanmar". Biology 11 (11): 1629. doi:10.3390/biology11111629. PMID 36358330.

[16] Li, Y.; Li, Y.-D.; Wang, Y.-D.; Schneider, H.; Shi, G.-L. (2022). "Re-appraisal of lacewing mimicry of liverworts from the mid-Cretaceous Kachin amber, Myanmar with a description of Selaginella cretacea sp. nov. (Selaginellales, Selaginellaceae)". Cretaceous Research 133: Article 105143. doi:10.1016/j.cretres.2022.105143. Bibcode: 2022CrRes.13305143L.

[17] Gao, X.; Liu, L.; Qin, M.; Zhou, Y.; Mao, L.; Wang, D.-M. (2022). "Re-study of Guangdedendron micrum from the Late Devonian Xinhang forest". BMC Ecology and Evolution 22 (1): Article number 69. doi:10.1186/s12862-022-02021-w. PMID 35606742.

[18] Xu, P.; Liu, L.; Wang, D.-M. (2022). "Reinvestigation of the Late Devonian Lycopsid Sublepidodendron grabaui from Anhui Province, South China". Biology 11 (10): 1544. doi:10.3390/biology11101544. PMID 36290447.

[FR251Feldbergetal-19] 19.0 ^19.1 Feldberg, K.; Schäfer-Verwimp, A.; Li, Y.; Renner, M. A. M. (2022). "Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae)". Fossil Record 25 (1): 213–230. doi:10.3897/fr.25.82362.

[20] Santos, A. A.; Sender, L. M.; Piñuela, L.; García-Ramos, J. C.; Diez, J. B. (2022). "First evidence of Ricciaceae in the Jurassic of the Iberian Peninsula (Asturias, NW Spain): Ricciopsis asturicus sp. nov". Botany Letters 169 (4): 557–567. doi:10.1080/23818107.2022.2124452.

[Ricciocarpolites-21] 21.0 ^21.1 Savoretti, A.; Bodnar, J.; Coturel, E. P.; Beltrán, B. (2022). "Fossil bryophytes from the Middle Triassic Sorocayense Group, San Juan Province, central-western Argentina". Ameghiniana 59 (3): 179–200. doi:10.5710/AMGH.26.02.2022.3469. https://www.ameghiniana.org.ar/index.php/ameghiniana/article/view/3469.

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[23] Skog, J. E.; Sender, L. M. (2022). "New information and family relationship (Hymenophyllaceae) for the fossil fern genus Acrostichopteris Fontaine and a new species from the Lower Cretaceous (Albian) of Spain". American Journal of Botany 109 (9): 1443–1455. doi:10.1002/ajb2.16050. PMID 36045579.

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